�@�@ •ª—Þƒgƒbƒv
Family Cichlidae�@ƒJƒ�ƒXƒYƒ�‰È

---

Sub-Family Geophaginae HASEMAN 1911�@ƒQƒIƒtƒ@ƒOƒXˆŸ‰È
�@�V‘å—¤�B



Genus Acarichthys^♂ EIGENMANN 1912�@ƒAƒJƒŠƒNƒeƒBƒX‘®
�@�@�@�¦�© [Acarichthyini KULLANDER 1998]�B

�Ÿ
�@Acarichthys heckelii (Müuller & Troschel 1848)
�@�@�@syn.Acara heckelii Müuller & Troschel 1848
�@�@�@�@�@Acara subocularis Cope 1878
�@�@�@�@�@Acarichthys heckeli Eigenmann 1912�@�¦ƒXƒyƒ‹ƒ~ƒX�B
�@�@�@�@�@Geophagus thayeri Steindachner 1875
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@"ƒXƒŒƒbƒhƒtƒBƒ“ƒAƒJƒ‰"�B

---

Genus Apistogramma^♀ REGAN 1913�@ƒAƒsƒXƒgƒOƒ‰ƒ€ƒ}‘®
�@�@�@�¦�‚Genus Heterogramma^♀ GUENÉE 1854 [Noctuidae, Lepidoptera, Insecta]�B
�@�@�@�¦��Genus Heterogramma^♀ REGAN 1906�B
�@�@�@�¦�{Genus Pintoichthys^♂ FOWLER 1954 (Type species : Biotodoma trifasciatum Eigenmann & Kennedy 1903)�B

�Ÿ
�@Apistogramma acrensis


�Ÿ
�@Apistogramma agassizii (Steindachner 1875)
�@�@�@syn.Apistogramma parvum Ahl 1931
�@�@�@�@�@Geophagus agassizii Steindachner 1875�@�¦��Geophagus (Mesops) agassizii Steindachner 1875�B
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<agassizii group>�B


�Ÿ
�@Apistogramma aguarico Römer & Hahn 2013

�Ÿ
�@Apistogramma alacrina

�Ÿ
�@Apistogramma allpahuayo Römer, Beninde, Duponchelle, Vela-Díaz, Ortega, Hahn, Soares, Cachay, Davila, Cornejo & Renno 2012

�Ÿ
�@Apistogramma amoena (Cope 1872)
�@�@�@syn.Geophagus amoenus Cope 1872
�@^*1871-1872�wOn the Fishes of the Ambyiacu River 1872�FEdward Drinker Cope�x�B
�@"ƒvƒŒƒ[ƒ“ƒg"�B

�Ÿ
�@Apistogramma angayuara

�Ÿ
�@Apistogramma arua
�@�@�@syn.Apistogramma sp. 'Highfin Yellow'
�@<cacatuoides group I>�B

�Ÿ
�@Apistogramma atahuala
�@�@�@syn.Apistogramma sp. 'Sunset'
�@<cacatuoides group I>�B

�Ÿ
�@Apistogramma baenschi


�Ÿ
�@Apistogramma barlowi

�Ÿ
�@Apistogramma bitaeniata (Pellégrin 1936)
�@�@�@syn.Apistogramma klausewitzi Meinken 1962
�@�@�@�@�@Apistogramma kleei Meinken 1964
�@�@�@�@�@Apistogramma pertensis var.bitaeniata Pellégrin 1936
�@�@�@�@�@Apistogramma sweglesi Meinken 1961
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<agassizii group>�B


�Ÿ
�@Apistogramma borellii (Regan 1906)
�@�@�@syn.Apistogramma aequipinnis Ahl 1938
�@�@�@�@�@Apistogramma reitzigi Mitsch 1938
�@�@�@�@�@Apistogramma ritensis (Haseman 1911)
�@�@�@�@�@Apistogramma rondoni (Miranda-Ribeiro 1918)
�@�@�@�@�@Heterogramma borellii Regan 1906
�@�@�@�@�@Heterogramma ritensis Haseman 1911
�@�@�@�@�@Heterogramma rondoni Miranda-Ribeiro 1918
�@<borellii group>�B


�Ÿ
�@Apistogramma brevis
�@<agassizii group>�B


�Ÿ
�@Apistogramma cacatuoides Hoedeman 1951
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B

�@<cacatuoides group I>�B

�Ÿ
�@Apistogramma caetei
�@<regani group>�B


�Ÿ
�@Apistogramma caudomaculata Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma cinilabra Römer, Duponchelle, Diaz, Davilla, Sirvas, Catchay & Renno 2011

�Ÿ
�@Apistogramma commbrae (Regan 1906)
�@�@�@syn.Apistogramma corumbae (Eigenmann & Ward ex Eigenmann, McAtee & Ward 1907)
�@�@�@�@�@Heterogramma commbrae Regan 1906
�@�@�@�@�@Heterogramma corumbae Eigenmann & Ward ex Eigenmann, McAtee & Ward 1907
�@"ƒRƒ‹ƒ“ƒo"�B<commbrae group>�B


�Ÿ
�@Apistogramma cruzi Kullander 1986
�@�@�@syn.Apistogramma sp.'Parallelstreifen'
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<regani group>�B

�Ÿ
�@Apistogramma diplotaenia
�@�@�@syn.Apistogramma sp.'Doppelband'
�@<diplotaenia group>�B

�Ÿ
�@Apistogramma eleutheria Varella & Britzke 2016

�Ÿ
�@Apistogramma elizabethae
�@<agassizii group>�B


�Ÿ
�@Apistogramma eremnopygis

�Ÿ
�@Apistogramma erythrura Staeck & Schindler 2008


�Ÿ
�@Apistogramma eunota Kullander 1981
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<regani group>�B

�Ÿ
�@Apistogramma feconat Römer, Soares, Dávila, Duponchelle, Renno & Hahn 2015

�Ÿ
�@Apistogramma flabellicauda Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma flavipedunculata Varella & Britzke 2016

�Ÿ
�@Apistogramma geisleri
�@<regani group>�B

�Ÿ
�@Apistogramma gephyra
�@<agassizii group>�BIUCN Least Concern ver.3.1�B

�Ÿ
�@Apistogramma gibbiceps
�@�@�@syn.Apistogramma roraimae
�@<gibbiceps group>�B

�Ÿ
�@Apistogramma gossei
�@<regani group>�B

�Ÿ
�@Apistogramma guttata
�@<macmasteri group>�B

�Ÿ
�@Apistogramma helkeri Schindler & Staeck 2013

�Ÿ
�@Apistogramma hippolytae
�@<steindachneri group>�B

�Ÿ
�@Apistogramma hoignei
�@<macmasteri group>�B

�Ÿ
�@Apistogramma hongsloi
�@�@�@syn.Apistogramma sp.'Rostrich'
�@<macmasteri group>�B


�Ÿ
�@Apistogramma huascar

�Ÿ
�@Apistogramma inconspicua
�@�@�@syn.Apistogramma sp.'Rostbrauner'
�@<commbrae group>�B

�Ÿ
�@Apistogramma iniridae
�@<pertensis group>�B


�Ÿ
�@Apistogramma inornata

�Ÿ
�@Apistogramma intermedia Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma juruensis Kullander 1986
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<cacatuoides group I>�B

�Ÿ
�@Apistogramma kullanderi Varella & Pérez 2014

�Ÿ
�@Apistogramma lineata Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma linkei
�@<commbrae group>�B

�Ÿ
�@Apistogramma luelingiKullander 1976
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<cacatuoides group I>�B

�Ÿ
�@Apistogramma macmasteri
�@<macmasteri group>�B


�Ÿ
�@Apistogramma martini

�Ÿ
�@Apistogramma megaptera Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma megastoma Römer, Römer, Estivals, Díaz, Duponchelle, Dávila, Hahn & Renno 2017
�@�@�@syn.Apistogramma sp.'Jutai' Römer & Römer 2015
�@^*2017�wDescription of a New Maternal Larvophilic Mouth-Brooding Cichlid Species, Apistogramma megastoma sp.n. [Teleostei, Perciformes, Geophaginae], from Loreto, Peru�FUwe Römer, Clara Isis Römer, Guillain Estivals, Antonia Vela Díaz, Fabrice Duponchelle, Carmen Rosa Garcia Dávila, Ingo Hahn & Jean-Francois Renno�x�B
�@Œû�o“à›z‰»�B

�Ÿ
�@Apistogramma meinkeni
�@<pertensis group>�B

�Ÿ
�@Apistogramma mendezi
�@�@�@syn.Apistogramma sp.'Langsstreifen'
�@�@�@�@�@Apistogramma sp.'LuƒÀkopf'
�@<agassizii group>�B

�Ÿ
�@Apistogramma minima Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma moae Kullander 1980
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<regani group>�B

�Ÿ
�@Apistogramma nijsseni Kullander 1979
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@<cacatuoides group II>�B

�Ÿ
�@Apistogramma norberti
�@"ƒrƒbƒOƒ}ƒEƒX"�B<cacatuoides group II>
�Ÿ
�@Apistogramma nororientalis Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma ortegai Britzke, de Oliveira & Kullander 2014


�Ÿ
�@Apistogramma ortmanni (Eigenmann 1912)
�@�@�@syn.Heterogramma ortmanni Eigenmann 1912
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@<regani group>�B

�Ÿ
�@Apistogramma panduro
�@�@�@syn.Apistogramma coriati
�@�@�@�@�@Apistogramma sp.'pandurini'
�@<cacatuoides group I>�B


�Ÿ
�@Apistogramma pantalonis


�Ÿ
�@Apistogramma paucisquamis�@<agassizii group>�B

�Ÿ
�@Apistogramma paulmuelleri Römer, Beninde, Duponchelle, Dávila, Díaz & Renno 2013

�Ÿ
�@Apistogramma payaminonis
�@<cacatuoides group II>�B

�Ÿ
�@Apistogramma pedunculata Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma personata
�@<gibbiceps group>�B


�Ÿ
�@Apistogramma pertensis (Haseman 1911)
�@�@�@syn.Heterogramma taeniata var.pertensis Haseman 1911
�@"ƒAƒ}ƒ]ƒ“ƒhƒ��[ƒt"�B<pertensis group>�B


�Ÿ
�@Apistogramma piaroa Mesa-Salazar & Lasso 2012

�Ÿ
�@Apistogramma piauiensis
�@�@�@syn.Apistogramma sp.'Wild linkei'
�@<regani group>�B

�Ÿ
�@Apistogramma playayacu Römer, Beninde & Hahn 2011

�Ÿ
�@Apistogramma pleurotaenia (Regan 1909)
�@�@�@syn.Heterogramma pleurotaenia Regan 1909
�@<regani group>�B

�Ÿ
�@Apistogramma psammophila Staeck & Schindler 2019
�@^*2019�wApistogramma psammophila, a New Geophagine Dwarf Cichlid [Teleostei, Cichlidae] from the Rio Atabapo Drainage in Colombia and Venezuela�FWolfgang Staeck & Ingo Schindler�x�B

�Ÿ
�@Apistogramma pulchra
�@�@<agassizii group>�B

�Ÿ
�@Apistogramma regani
�@<regani group>�B

�Ÿ
�@Apistogramma resticulosa
�@<regani group>�B

�Ÿ
�@Apistogramma rositae

�Ÿ
�@Apistogramma rubrolineata


�Ÿ
�@Apistogramma rupununi
�@"ƒgƒD�[ƒXƒ|ƒbƒg"�@<steindachneri group>�B

�Ÿ
�@Apistogramma salpinction

�Ÿ
�@Apistogramma similis

�Ÿ
�@Apistogramma sororcula Staeck & Schindler 2016

�Ÿ
�@Apistogramma staecki
�@<cacatuoides group I>�B


�Ÿ
�@Apistogramma steindachneri (Regan 1908)
�@�@�@syn.Apistogramma ornatipinnis Ahl 1936
�@�@�@�@�@Apistogramma wickleri Meinken 1960
�@�@�@�@�@Heterogramma steindachneri Regan 1908
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@<steindachneri group>�B

�Ÿ
�@Apistogramma taeniata (Güunther 1862)
�@�@�@syn.Mesops taeniatus Güunther 1862
�@<regani group>�B

�Ÿ
�@Apistogramma trifasciata (Eigenmann & Kennedy 1903)
�@�@�@syn.Apistogramma maciliensis (Haseman 1911)
�@�@�@�@�@Apistogramma sp. 'Rio Mamore'
�@�@�@�@�@Apistogramma trifasciata haraldschultzi Meinken 1960
�@�@�@�@�@Heterogramma trifasciata var.maciliensis Haseman 1911
�@�@�@�@�@Apistogramma trifasciata maciliensis (Haseman 1911)
�@�@�@�@�@Biotodoma trifasciatum Eigenmann & Kennedy 1903
�@�@�@�@�@Heterogramma trifasciata maciliensis (Haseman 1911)
�@�@�@�@�@Pintoichthys trifasciatus (Eigenmann & Kennedy 1903)
�@<trifasciata group>�B


�Ÿ
�@Apistogramma tucurui

�Ÿ
�@Apistogramma uaupesi
�@�@�@syn.Apistogramma sp.'Blutkehl'
�@�@�@�@�@Apistogramma sp.'Rotkeil'
�@<pertensis group>�B


�Ÿ
�@Apistogramma urteagai Kullander 1986
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@IUCN Least Concern ver.3.1�B
�@<regani group>

�Ÿ
�@Apistogramma velifera

�Ÿ
�@Apistogramma viejita
�@<macmasteri group>�B


�Ÿ
�@Apistogramma wapisana

�Ÿ
�@Apistogramma wolli Römer, Soares, Dávila, Duponchelle, Renno & Hahn 2015

�Ÿ
�@Apistogramma sp. 'Maulbruter'
�@��Apistogramma amoena (Cope 1872)�H

---

Genus Apistogrammoides^♂ MEINKEN 1965�@ƒAƒsƒXƒgƒOƒ‰ƒ€ƒ‚ƒCƒfƒX‘®

�Ÿ
�@Apistogrammoides pucallpaensis Meinken 1965
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B

---

Genus Biotodoma^⚲ EIGENMANN & KENNEDY 1903�@ƒrƒIƒgƒhƒ}‘®
�@�@�@�¦�‚Genus Mesops^♂ AUDINET-SERVILLE 1831 [Orthoptera, Insecta]�B
�@�@�@�¦�‚Genus Mesops^♂ BILLBERG 1820 [Acrididae, Coleoptera, Insecta]�B
�@�@�@�¦��Genus Mesops^♂ GÜNTHER 1862�B

�Ÿ
�@Biotodoma cupido (Heckel 1840)
�@�@�@syn.Geophagus cupido Heckel 1840
�@�@�@�@�@Mesops cupido (Heckel 1840)
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@"ƒOƒŠ�[ƒ“ƒXƒgƒŠ�[ƒNƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Biotodoma wavrini
�@"ƒIƒŠƒmƒRƒA�[ƒXƒC�[ƒ^�["�B

---

Genus Biotoecus^♂ EIGENMANN & KENNEDY 1903�@ƒrƒIƒgƒGƒNƒX‘®
�@�@�@�¦�‚Genus Saraca^♀ WALKER 1865 [Noctuidae, Lepidoptera, Insecta]�B
�@�@�@�¦��Genus Saraca^♀ STEINDACHNER 1875�B

�Ÿ
�@Biotoecus dicentrarchus

�Ÿ
�@Biotoecus opercularis (Steindachner 1875)
�@�@�@syn.Saraca opercularis Steindachner 1875
�@"ƒOƒŠ�[ƒ“ƒhƒ��[ƒtƒVƒNƒŠƒbƒh"�B

---

Genus Crenicara^♀ STEINDACHNER 1875�@ƒNƒŒƒjƒJƒ‰‘®
�@�@�@�¦��Genus Crenacara^♀ REGAN 1905�B
�@�@�@�¦�© [Crenicaratini KULLANDER 1998]�B
�@�@�@�¦�© [Crenicarini HOEDEMAN 1947]�B

�Ÿ
�@Crenicara latruncularia
�@"ƒwƒ‰ƒNƒŒƒXƒVƒNƒŠƒbƒh"�B

�Ÿ
�@Crenicara punctulata (Günther 1863)
�@�@�@syn.Acara punctulata Günther 1863
�@�@�@�@�@Aequidens hercules Allen 1942
�@�@�@�@�@Aequidens madeirae Fowler 1913
�@�@�@�@�@Crenacara elegans (Steindachner 1875)
�@�@�@�@�@Crenicara elegans Steindachner 1875
�@�@�@�@�@�@�@�¦–ÍŽ®Ží�A’A‚µ�æ‚É‹L�Ú‚³‚ꂽ Acara punctulata Günther 1863 ‚É“��‡�B
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@"ƒ`ƒFƒbƒJ�[ƒ{�[ƒh(ŠCŠO)"�B

---

Genus Dicrossus^♂ STEINDACHNER 1875�@ƒfƒBƒNƒ�ƒbƒXƒX‘®

�Ÿ
�@Dicrossus filamentosus
�@�@�@syn.Crenicara filamentosa
�@"ƒ`ƒFƒbƒJ�[ƒ{�[ƒh"�B


�Ÿ
�@Dicrossus foirni Römer, Hahn & Vergara 2010


�Ÿ
�@Dicrossus gladicaudus

�Ÿ
�@Dicrossus maculatus Steindachner 1875
�@�@�@syn.Crenicara maculata (Steindachner 1875)
�@�@�@�@�@Crenicara praetoriusi Weise 1935 non Ahl 1936


�Ÿ
�@Dicrossus praetoriusi (Ahl 1936)
�@�@�@syn.Crenicara praetoriusi Ahl 1936 non Weise 1935
�@��Dicrossus maculatus Steindachner 1875�H

�Ÿ
�@Dicrossus warzeli Römer, Hahn & Vergara 2010

---

Genus Geophagus^♂ HECKEL 1840�@ƒQƒIƒtƒ@ƒOƒX‘®
�@�@�@�¦�{Genus Macracara^♀ WOODWARD 1939 (Type species : Macracara prisca Woodward 1939)�B

�Ÿ
�@Geophagus abalios



�Ÿ
�@Geophagus altifrons Heckel 1840


�Ÿ
�@Geophagus argyrostictus


�Ÿ
�@Geophagus brachybranchus
�@IUCN Least Concern ver.3.1�B



�Ÿ
�@Geophagus brasiliensis (Quoy & Gaimard ex de Freycinet 1824)
�@�@�@syn.Acara gymnopoma Gunther 1862
�@�@�@�@�@Acara minuta Hensel 1870
�@�@�@�@�@Chromis brasiliensis Quoy & Gaimard ex de Freycinet 1824 non Kner 1865
�@�@�@�@�@Chromis unimaculatus (de Castelnau 1855)
�@�@�@�@�@Chromys unimaculatus de Castelnau 1855
�@�@�@�@�@Geophagus brasiliensis Kner 1865 non Quoy & Gaimard ex de Freycinet 1824
�@^*1824�wZoology by Jean René Constant Quoy & Joseph Paul Gaimard in Voyage Autour du Monde, Entrepris par Ordre du Roi, sur les Corvettes de S.M. l'Uranie et la Physicienne, Pendant les Années 1817, 1818, 1819 et 1820 3�FLouis Claude Desaulses de Freycinet�x�B
�@^*1855�wAnimaux Nouveaux ou Rares Recueillis Pendant l'Expedition dans les Parties Centrales de L'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para 2, Poissons�FFrançois-Louis la Porte/comte de Castelnau�x�B
�@^*2020�wUn-Raveling the Systematics and Evolution of the Geophagus brasiliensis [Cichliformes, Cichlidae] Species Complex�FLeandro Araujo Argoloa, Hernán López-Fernández, Henrique Batalha-Filho & Paulo Roberto Antunes de Mello Affonsod�x�B
�@"ƒp�[ƒ‹ƒVƒNƒŠƒbƒh"�B

Chromys unimaculatus=Chromis brasiliensis

br> �Ÿ
�@Geophagus brokopondo


�Ÿ
�@Geophagus camopiensis
�@"ƒIƒCƒ„ƒ|ƒbƒNƒA�[ƒXƒC�[ƒ^�["�B



�Ÿ
�@Geophagus crassilabris Steindachner 1876�@�¦��Geophagus (Satanoperca) crassilabris Steindachner 1876�B
�@�@�@syn.Satanoperca crassilabris (Steindachner 1876) Jordan & Evermann 1896
�@^*1876�wIchthyologische Beiträge 5�FFranz Steindachner�x�B
�@^*1896�wA Check-List of the Fishes and Fish-Like Vertebrates of North and Middle America 1895�FDavid Starr Jordan & Barton Warren Evermann�x�B
�@^*1898�wThe Fishes of North and Middle America 2, a Descriptive Catalogue of the Species of Fish-Like Vertebrates Found in the Waters of North America, North of the Isthmus of Panama�FDavid Starr Jordan & Barton Warren Evermann�x�B
�@"ƒpƒiƒ}ƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Geophagus crocatus Hauser & López-Fernández 2013
�@^*2013�wGeophagus crocatus, a New Species of Geophagine Cichlid from the Berbice River, Guyana, South America [Teleostei, Cichlidae]�FFrances E.Hauser & Hernan López-Fernández�x�B

�Ÿ
�@Geophagus diamantinensis Mattos, Costa & Santos 2015

�Ÿ
�@Geophagus dicrozoster


�Ÿ
�@Geophagus gottwaldi

�Ÿ
�@Geophagus grammepareius

�Ÿ
�@Geophagus harreri
�@"ƒ}ƒ�ƒjƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Geophagus iporangensis
�@�@�@syn.Geophagus brasiliensis iporangensis


�Ÿ
�@Geophagus itapicuruensis
�@�@�@syn.Geophagus brasiliensis itapicuruensis

�Ÿ
�@Geophagus megasemus

�Ÿ
�@Geophagus mirabilis Deprá, Kullander, Pavanelli & da Graça 2014


�Ÿ
�@Geophagus multiocellus Mattos & da Costa 2018
�@^*2018�wThree New Species of the Geophagus brasiliensis Species Group from the Northeast Brazil [Cichlidae, Geophagini]�FJosé L.O.Mattos & Wilson José Eduardo Moreira da Costa�x�B

�Ÿ
�@Geophagus neambi Lucinda, de Lucena & Assis 2010


�Ÿ
�@Geophagus olfersi Gosse 1976
�@��Geophagus brasiliensis (Quoy & Gaimard ex de Freycinet 1824)�H

�Ÿ
�@Geophagus parnaibae


�Ÿ
�@Geophagus pellegrini
�@"ƒCƒGƒ��[ƒnƒ“ƒvƒA�[ƒXƒC�[ƒ^�["�B


�ŸƒQƒIƒtƒ@ƒOƒX�EƒvƒŠƒXƒNƒX (�â–Å)
�@Geophagus priscus (Woodward 1939)
�@�@�@syn.Macracara prisca Woodward 1939
�@^*1939�wTertiary Fossil Fishes from the Maranhão, Brazil�FArthur Smith Woodward�x�B


�Ÿ
�@Geophagus proximus (de Castelnau 1855)
�@�@�@syn.Chromys proximus de Castelnau 1855
�@^*1855�wAnimaux Nouveaux ou Rares Recueillis Pendant l'Expedition dans les Parties Centrales de L'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para 2, Poissons�FFrançois-Louis la Porte/comte de Castelnau�x�B
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B


�Ÿ
�@Geophagus pyrineusi Deprá, Ohara & Silva 2022
�@^*2022�wGeophagus pyrineusi, a New Species from the Rio Teles Pires, Rio Tapajós Basin, Brazil [Cichliformes, Cichlidae, Geophagini]�FGabriel de Carvalho Deprá, Willian Massaharu Ohara & Hugmar Pains Silva�x�B

�Ÿ
�@Geophagus pyrocephalus Chuctaya, Nitschke, Andrade & Wingert 2022
�@^*2022�wA New Species of Geophagus [Teleostei, Cichlidae], Naming a Cichlid Species Widely Known in the Aquarium Hobby as Geophagus sp.Tapajos Red-Head�FJunior Chuctaya, Pedro Nitschke, Marcelo Costa Andrade & Juliana Mariani Wingert�x�B

�Ÿ
�@Geophagus rufomarginatus Mattos & da Costa 2018
�@^*2018�wThree New Species of the Geophagus brasiliensis Species Group from the Northeast Brazil [Cichlidae, Geophagini]�FJosé L.O.Mattos & Wilson José Eduardo Moreira da Costa�x�B

�Ÿ
�@Geophagus santosi Mattos & da Costa 2018
�@^*2018�wThree New Species of the Geophagus brasiliensis Species Group from the Northeast Brazil [Cichlidae, Geophagini]�FJosé L.O.Mattos & Wilson José Eduardo Moreira da Costa�x�B

�Ÿ
�@Geophagus steindachneri
�@�@�@syn.Geophagus hondae
�@�@�@�@�@Geophagus magdalenus
�@"ƒŒƒbƒhƒnƒ“ƒvƒA�[ƒXƒC�[ƒ^�["�B



�Ÿ
�@Geophagus surinamensis (Bloch 1791) Göldi 1900
�@�@�@syn.Cichla surinamensis (Bloch 1791) Bloch post Schneider 1801
�@�@�@�@�@Sparus surinamensis Bloch 1791
�@�@�@�@�@Sparus trimaculatus Shaw 1809
�@^*1801�wSystema Ichthyologiae Iconibus cx Illustratum�FMarcus Elieser Bloch after Johann Gottlob Schneider�x�B
�@^*1817�wLe Règne Animal Distribué d'Après son Organisation Pour Servir de Base à l'Histoire Naturelle des Animaux et d'Introduction à l'Anatomie Comparée 2, les Reptiles, les Poissons, les Mollusques et les Annélides�FGeorges Léopold Chrétien Frédéric Dagobert Cuvier�x�B
�@^*1900�wDie Fischwelt des Amazonas-Gebietes 2�FÉmil August Göldi�x�B
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B


�Ÿ
�@Geophagus sveni Lucinda, de Lucena & Assis 2010


�Ÿ
�@Geophagus taeniopareius


�Ÿ
�@Geophagus tuberosus Gosse 1976
�@��Geophagus brasiliensis (Quoy & Gaimard ex de Freycinet 1824)�H


�Ÿ
�@Geophagus unipunctatus (de Castelnau 1855)
�@�@�@syn.Chromis obscurus (de Castelnau 1855)
�@�@�@�@�@Chromis unipunctatus (de Castelnau 1855)
�@�@�@�@�@Chromys obscurus de Castelnau 1855
�@�@�@�@�@Chromys unipunctatus de Castelnau 1855
�@�@�@�@�@Geophagus obscurus (de Castelnau 1855)
�@^*1855�wAnimaux Nouveaux ou Rares Recueillis Pendant l'Expedition dans les Parties Centrales de L'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para 2, Poissons�FFrançois-Louis la Porte/comte de Castelnau�x�B
�@ƒJƒXƒeƒ‹ƒm�[‚Ì�˜—ñ‚Í Chromys 1.punctata�H�A2.unipunctata�A3.unimaculata�A4.obscura�B

Chromys obscurus=Chromys unipunctatus

�Ÿ
�@Geophagus winemilleri

---

Genus Guianacara^♀ KULLANDER & NIJSSEN 1989�@ƒOƒCƒAƒiƒJƒ‰‘®
�@�@�@�¦�{Genus Guianacara^♀ KULLANDER & NIJSSEN 1989 Sub-Genus Oelemaria^♀ KULLANDER & NIJSSEN 1989 (Type species : Guianacara oelemariensis Kullander & Nijssen 1989)�B
�@�@�@�¦�{Genus Oelemaria^♀ (KULLANDER & NIJSSEN 1989) (Type species : Guianacara oelemariensis Kullander & Nijssen 1989)�B

�Ÿ
�@Guianacara cuyunii

�Ÿ
�@Guianacara dacrya Arbour & López-Fernández 2011


�Ÿ
�@Guianacara geayi (Pellégrin 1902)
�@�@�@syn.Acara geayi Pellégrin 1902
�@�@�@�@�@Acarichthys geayi (Pellégrin 1902)
�@�@�@�@�@Aequidens geayi (Pellégrin 1902) Eigenmann 1912
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B


�Ÿ
�@Guianacara oelemariensis Kullander & Nijssen 1989
�@�@�@syn.Oelemaria oelemariensis (Kullander & Nijssen 1989)
�@IUCN Least Concern ver.3.1�B

�Ÿ
�@Guianacara owroewefi Kullander & Nijssen 1989


�Ÿ
�@Guianacara sphenozona


�Ÿ
�@Guianacara stergiosi

---

Genus Gymnogeophagus^♂ DE MIRANDA-RIBEIRO 1918�@ƒMƒ€ƒmƒQƒIƒtƒ@ƒOƒX‘®

�Ÿ
�@Gymnogeophagus australis
�@�@�@syn.Geophagus australe
�@"ƒEƒ‹ƒOƒAƒCƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Gymnogeophagus balzanii (Perugia 1891)
�@�@�@syn.Geophagus balzanii Perugia 1891
�@�@�@�@�@Geophagus duodecimspinosus Boulenger 1895
�@�@�@�@�@Gymnogeophagus cyanopterus de Miranda-Ribeiro 1918
�@�@�@�@�@�@�@�¦–ÍŽ®Ží�A’A‚µ�æ‚É‹L�Ú‚³‚ꂽ Geophagus balzanii Perugia 1891 ‚É“��‡�B
�@"ƒAƒ‹ƒ[ƒ“ƒ`ƒ“ƒnƒ“ƒvƒwƒbƒh"�B

�Ÿ
�@Gymnogeophagus caaguazuensis


�Ÿ
�@Gymnogeophagus che

�Ÿ
�@Gymnogeophagus constellatus Malabarba, Malabarba & Reis 2015

�ŸƒMƒ€ƒmƒQƒIƒtƒ@ƒOƒX�EƒGƒIƒPƒjƒNƒX (�â–Å)
�@Gymnogeophagus eocenicus Malabarba, Malabarba, & del Papa 2010
�@^*2010�wGymnogeophagus eocenicus n.sp. [Perciformes, Cichlidae], an Eocene Cichlid from the Lumbrera Formation in Argentina�FMaria Cláudia de Souza Lima Malabarba, Luiz Roberto Malabarba, & Cecilia del Papa�x�B

�Ÿ
�@Gymnogeophagus gymnogenys
�@�@�@syn.Geophagus camurus
�@�@�@�@�@Geophagus pygmaeus
�@"ƒXƒ€�[ƒXƒ`�[ƒNƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Gymnogeophagus jaryi Alonso, Terán, Aguilera, Ríčan, Casciotta, Serra, Almirón, Benítez, García & Mirande 2019
�@^*2019�wDescription of a New Species of the Neotropical Cichlid Genus Gymnogeophagus Miranda Ribeiro 1918 [Teleostei, Cichliformes] from the Middle Paraná Basin, Misiones, Argentina�FFelipe Alonso, Guillermo Enrique Terán, Gastón Aguilera, Oldřich Ríčan, Jorge Rafael Casciotta, Wilson Sebastián Serra, Adriana Edith Almirón, Mauricio Fábian Benítez, Ignacio García & Juan Marcos Mirande�x�B

�Ÿ
�@Gymnogeophagus labiatus (Hensel 1870)
�@�@�@syn.Geophagus bucephalus Hensel 1870
�@�@�@�@�@Geophagus labiatus Hensel 1870
�@�@�@�@�@Geophagus scymnophilus Hensel 1870
�@^*2021�wHydrography Rather than Lip Morphology Better Explains the Evolutionary Relationship Between Gymnogeophagus labiatus and G.lacustris in Southern Brazil [Cichlidae, Geophagini]�FPedro Ivo Campani de Castro Figueiredo, Luiz Roberto Malabarba & Nelson Jurandi Rosa Fagundes�x�B
�@"ƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Gymnogeophagus lacustris Reis & Malabarba 1988
�@^*2021�wHydrography Rather than Lip Morphology Better Explains the Evolutionary Relationship Between Gymnogeophagus labiatus and G.lacustris in Southern Brazil [Cichlidae, Geophagini]�FPedro Ivo Campani de Castro Figueiredo, Luiz Roberto Malabarba & Nelson Jurandi Rosa Fagundes�x�B

�Ÿ
�@Gymnogeophagus lipokarenos Malabarba, Malabarba & Reis 2015
�@�@�@syn.Gymnogeophagus sp.'E' Wimberger, Reis & Thornton 1998


�Ÿ
�@Gymnogeophagus mekinos Malabarba, Malabarba & Reis 2015
�@�@�@syn.Gymnogeophagus aff.gymnogenys Serra, Bessonart, de Mello, Duarte, Malabarba & Loureiro 2014
�@�@�@�@�@Gymnogeophagus cf.gymnogenys-complex Clade 1 Pereyra & Garcia 2008
�@�@�@�@�@Gymnogeophagus sp.'A' Wimberger, Reis & Thornton 1998


�Ÿ
�@Gymnogeophagus meridionalis


�Ÿ
�@Gymnogeophagus missioneiro Malabarba, Malabarba & Reis 2015
�@�@�@syn.Gymnogeophagus sp.'D' Wimberger, Reis & Thornton 1998

�Ÿ
�@Gymnogeophagus peliochelynion Turcati, Serra-Alanis & Malabarba 2018
�@^*2018�wA New Mouth Brooder Species of Gymnogeophagus with Hypertrophied Lips [Cichliformes, Cichlidae]�FAndréia Turcati, Wilson Sebastián Serra-Alanis & Luiz Roberto Malabarba�x�B

�Ÿ
�@Gymnogeophagus pseudolabiatus Malabarba, Malabarba & Reis 2015
�@�@�@syn.Gymnogeophagus cf.gymnogenys-complex Clade 2 Pereyra & Garcia 2008
�@�@�@�@�@Gymnogeophagus sp.'C' Wimberger, Reis & Thornton 1998

�Ÿ
�@Gymnogeophagus rhabdotus
�@�@�@syn.Geophagus brachyurus
�@"ƒXƒgƒ‰ƒCƒvƒtƒBƒ“ƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Gymnogeophagus setequedas


�Ÿ
�@Gymnogeophagus tarobus Casciotta, Almirón, Piálek & Řičan 2017
�@^*2017�wGymnogeophagus taroba [Teleostei, Cichlidae], a New Species from the Río Iguazú Basin, Misiones, Argentina�FJorge Rafael Casciotta, Adriana Edith Almirón, Lubomir Piálek & Oldřich Řičan�x�B


�Ÿ
�@Gymnogeophagus terrapurpurus Loureiro, Zarucki, Malabarba & González-Bergonzoni 2016


�Ÿ
�@Gymnogeophagus tiraparae

---

Genus Mazarunia^♀ KULLANDER 1990�@ƒ}ƒUƒ‹ƒjƒA‘®

�Ÿ
�@Mazarunia charadrica López-Fernández, Taphorn-Baechle & Liverpool 2012

�Ÿ
�@Mazarunia mazarunii Kullander 1990

�Ÿ
�@Mazarunia pala López-Fernández, Taphorn-Baechle & Liverpool 2012

---

Genus Mikrogeophagus^♂ MEULENGRACHT-MADSEN ex SCHIOTZ & CHRISTENSEN 1968�@ƒ~ƒNƒ�ƒQƒIƒtƒ@ƒOƒX‘®
�@�@�@�¦��Genus Microgeophagus^♂ AXELROD 1971�B
�@�@�@�¦��Genus Microgeophagus^♂ FREY 1957�B
�@�@�@�¦��Genus Papiliochromis^♂ KULLANDER 1977�B��Genus Papilochromis�B
�@�@�@�¦�{Genus Pseudoapistogramma^♀ AXELROD 1971�B–ÍŽ®Ží“™‹L�Ú‚È‚µ�B
�@�@�@�¦��Genus Pseudogeophagus^♂ HOEDEMAN 1974�B
�@ƒtƒŒƒC‚ÍŽ”ˆç�‘‚Ƀ~ƒNƒ�ƒQƒIƒtƒ@ƒOƒX‚ð’ñŽ¦‚µ‚½‚ª�AŠw–¼‚Å‚Í‚È‚­‚ ‚ç‚ä‚é‹L�q‚ª‘¶�Ý‚µ‚È‚¢�B

�Ÿ
�@Mikrogeophagus altispinosus


�Ÿ
�@Mikrogeophagus maculicaudus Staeck, Ottoni & Schindler 2022
�@^*2022�wMikrogeophagus maculicauda, a New Dwarf Cichlid [Teleostei, Cichlidae] from the Eastern Drainage of the Upper Rio Guaporá, Brazil�FWolfgang Staeck, Felipe Polivanov Ottoni & Ingo Schindler�x�B


�Ÿ
�@Mikrogeophagus ramirezi (Myers & Harry 1948)
�@�@�@syn.Apistogramma ramirezi Myers & Harry 1948
�@�@�@�@�@Microgeophagus ramirezi (Myers & Harry 1948)
�@�@�@�@�@Pseudogeophagus ramirezi (Myers & Harry 1948)
�@"ƒ‰ƒ€(+ƒhƒCƒc�EƒIƒ‰ƒ“ƒ_)"�B

---

Genus Satanoperca^♀ GÜNTHER 1862�@ƒTƒ^ƒmƒyƒ‹ƒJ‘®

�Ÿ
�@Satanoperca acuticeps
�@�@�@syn.Geophagus acuticeps
�@"ƒVƒƒ�[ƒvƒwƒbƒhƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Satanoperca curupira Ota, Kullander, Deprá, da Graça & Pavanelli 2018
�@^*2018�wSatanoperca curupira, a New Cichlid Species from the Rio Madeira Basin in Brazil [Teleostei: Cichlidae]�FRenata Rúbia Ota, Sven Oscar Kullander, Gabriel C.Deprá, Weferson Júnio da Graça & Carla Simone Pavanelli�x�B

�Ÿ
�@Satanoperca daemon (Heckel 1840)
�@�@�@syn.Geophagus daemon Heckel 1840
�@"ƒXƒŠ�[ƒXƒ|ƒbƒgƒA�[ƒXƒC�[ƒ^�["�B



�Ÿ
�@Satanoperca jurupari (Hecklel 1840)
�@�@�@syn.Geophagus jurupari Hecklel 1840
�@^*1871-1872�wOn the Fishes of the Ambyiacu River 1872�FEdward Drinker Cope�x�B
�@^*1986�wCichlid Fishes of the Amazon River Drainage of Peru�FSven Oscar Kullander�x�B
�@^*1912�wThe Fresh-Water Fishes of British Guiana, Including a Study of the Ecological Grouping of Species and the Relation of the Fauna of the Plateau to That of of the Low-Land 8, Systematic Account of the Fresh-Water Fishes of British Guiana�FCarl Henry Eigenmann�x�B
�@"ƒf�[ƒ‚ƒ“ƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Satanoperca leucosticta
�@�@�@syn.Geophagus leucostictus
�@�@�@�@�@Satanoperca macrolepis


�Ÿ
�@Satanoperca lilith


�Ÿ
�@Satanoperca mapiritensis


�Ÿ
�@Satanoperca pappaterra
�@"ƒpƒ“ƒ^ƒiƒ‹ƒA�[ƒXƒC�[ƒ^�["�B


�Ÿ
�@Satanoperca rhynchitis Kullander 2012


�Ÿ
�@Satanoperca setepele Ota, Deprá, Kullander, da Graça & Pavanelli 2021
�@^*2021�wA New Species of Satanoperca [Teleostei, Cichlidae] from the Rio Tocantins Basin, Brazil�FRenata Rúbia Ota, Gabriel de Carvalho Deprá, Sven Kullander, Weferson Júnio da Graça & Carla Simone Pavanelli�x�B

---

Genus Taeniacara^♀ MYERS 1935�@ƒ^ƒGƒjƒAƒJƒ‰‘®

�Ÿ
�@Taeniacara candidi Myers 1935

---