Conspectus cobitidum - Raffles Museum of Biodiversity Research
Conspectus cobitidum - Raffles Museum of Biodiversity Research
Conspectus cobitidum - Raffles Museum of Biodiversity Research
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THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012 Supplement No. 26: 1–199<br />
Date <strong>of</strong> Publication: 28 Dec.2012<br />
© National University <strong>of</strong> Singapore and the author<br />
CONSPECTUS COBITIDUM*:<br />
AN INVENTORY OF THE LOACHES OF THE WORLD<br />
(TELEOSTEI: CYPRINIFORMES: COBITOIDEI)<br />
Maurice Kottelat<br />
Case postale 57, CH-2952 Cornol, Switzerland (address for correspondence), and<br />
<strong>Raffles</strong> <strong>Museum</strong> <strong>of</strong> <strong>Biodiversity</strong> <strong>Research</strong>, Department <strong>of</strong> Biological Sciences, National University <strong>of</strong> Singapore,<br />
6 Science Drive 2, Singapore 117546, Republic <strong>of</strong> Singapore.<br />
E-mail: mkottelat@dplanet.ch<br />
CONTENTS<br />
Abstract ...................................................................................................................... 1<br />
Introduction ............................................................................................................... 3<br />
Name-bearing types ................................................................................................... 6<br />
Ending and spelling <strong>of</strong> species names ....................................................................... 8<br />
Abbreviations used .................................................................................................. 12<br />
Family Gyrinocheilidae ........................................................................................... 14<br />
Family Botiidae ....................................................................................................... 14<br />
Family Vaillantellidae .............................................................................................. 21<br />
Family Cobitidae ..................................................................................................... 21<br />
Family Ellopostomatidae......................................................................................... 43<br />
Family Barbuccidae................................................................................................. 43<br />
Family Balitoridae ................................................................................................... 44<br />
Family Gastromyzontidae ....................................................................................... 57<br />
Family Serpenticobitidae......................................................................................... 73<br />
Family Nemacheilidae ............................................................................................. 74<br />
Appendix: New genus-group and family-group names ........................................ 137<br />
Acknowledgements ............................................................................................... 141<br />
Bibliography .......................................................................................................... 142<br />
Index ...................................................................................................................... 175<br />
ABSTRACT. — The taxonomy and nomenclature <strong>of</strong> the fishes <strong>of</strong> the superfamily Cobitoidea<br />
(suborder Cobitoidei minus Catostomidae) are reviewed. Original descriptions <strong>of</strong> all 1499<br />
recorded species-group names and 185 genus-group names have been checked for correct spelling,<br />
types and bibliographic references. The bibliography includes about 1010 titles. 1043 valid<br />
species in 111 valid genera are recognised. Synonymies are given, based on published information<br />
as well as unpublished observations. Endings consistent with the three possible Latin grammatical<br />
* Cobitidum is formed on the common name cobites or cobitis, meaning loaches in general, and not on the formal name <strong>of</strong><br />
the suborder Cobitoidei.<br />
1
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
genders <strong>of</strong> genus-group names are indicated for all species-group names; basic tools are provided<br />
to establish the correct endings in most simple cases. The main nomenclatural acts are:<br />
– new family-group names: Serpenticobitidae, Barbuccidae;<br />
– new genera: Ambastaia (type species: Botia nigrolineata), Theriodes (type species:<br />
Acanthophthalmus sandakanensis), Speonectes (type species: Sundoreonectes tiomanensis);<br />
– lectotype designation: Cobitis stephanidisi (sensu Economidis, 1992);<br />
– declaration as nomen protectum: Cobitis biwae;<br />
– declaration as nomen oblitum: Cobitis schlegeli;<br />
– first reviser action on precedence <strong>of</strong> simultaneous synonyms: Parabotia kimluani over P.<br />
vancuongi, Cobitis taenia turcica over Cobitinula anatoliae, Sewellia medius over S. grandis,<br />
Parasewellia monolobata over P. polylobata, Cobitis turio over C. bilturio, Triplophysa<br />
bashanensis over T. longchiensis, Yunnanilus macrositanus over Y. forkicaudalis;<br />
– first reviser action on correct spelling <strong>of</strong>: Parabotia vancuongi, Cobitis fahireae, Hemimyzon<br />
songamensis, Sewellia analis, Parasewellia polylobata, Vanmanenia mon<strong>of</strong>asciodorsalata,<br />
V. trifaseudorsala, Oreonectes microphthalmus, Paracobitis posterodorsalus, Yunnanilus<br />
macrositanus, Y. forkicaudalis, Y. spanisbripes.<br />
2
Loaches, fishes <strong>of</strong> the suborder Cobitoidei, are small benthic<br />
fishes known throughout Eurasia, with two named species<br />
in Africa (Morocco and Ethiopia). Most <strong>of</strong> them inhabit rivers<br />
and streams in hilly areas but they are also known from<br />
streams in rainforest, in the black waters <strong>of</strong> peat swamp forests,<br />
or in oases (in the Taklamakan Desert). Many <strong>of</strong> them<br />
are specialised cave fishes, others inhabit rapids and waterfalls.<br />
The fishes reported from the highest altitude in Asia<br />
are loaches (Triplophysa stolickai at 5200 masl, in hot springs<br />
near Lungmu Co [lake], Ngari County in western Tibet<br />
[34°37'N 80°28'E]; Zhu, 1981: 1063); those reported from<br />
the lowest altitude are also loaches (Triplophysa zhaoi in<br />
swamps <strong>of</strong> the Lükqün oasis, about 50 m below sea level in<br />
the Turpan depression, Xinjiang, China).<br />
The suborder Cobitoidei derives its name from Cobitis, a<br />
genus established by Linnaeus (1758). This name has an old<br />
history. It first appears as κωβτις in Aristotle's Περ ζων<br />
στοραζ for small fishes that bury in the bottom (Aristotle,<br />
ca. 343 BC; first printed edition in 1476, Historia animalium,<br />
Inquiries on animals, book VI, chapter XIV). Cobitis<br />
means 'like the gudgeon' (κωβιζ). The word was first transcribed<br />
in the Latin alphabet by Rondelet (1555) who used<br />
it for three small fishes identifiable as today's Cobitis taenia,<br />
Barbatula barbatula, and, apparently, Alburnus alburnus.<br />
The name was then used by classical authors like Gesner,<br />
Aldrovandi, Jonston and Willughby. Cobitis is the second<br />
genus in Artedi's system (1738), who also added Misgurnus<br />
fossilis. As Artedi's system was adopted by Linnaeus (1758),<br />
Cobitis entered formal zoological nomenclature.<br />
Work on this checklist began around 1980 when I became<br />
interested in the taxonomy and ecology <strong>of</strong> Southeast Asian<br />
loaches. On 10 November 2012 the superfamily includes<br />
1043 valid species, out <strong>of</strong> 1499 nominal species (valid species<br />
plus all their synonyms, emendations, nomina nuda and<br />
some incorrect subsequent spelling occasionally used). They<br />
are presently placed in 111 genera (out <strong>of</strong> 185 nominal genera).<br />
I have examined the original description <strong>of</strong> all. About<br />
1400 publications have been checked and about 1010 relevant<br />
ones are listed.<br />
By loaches I understand here the suborder Cobitoidei. Some<br />
authors have considered the family Catostomidae as the sister-group<br />
<strong>of</strong> all other Cobitoidei (e.g. Šlechtová et al., 2007).<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
INTRODUCTION<br />
3<br />
Μα χελιδν αρ ο ποιε<br />
[One nucleotide does not a species make *]<br />
Aristotle (384–322 BC)<br />
[*apocryphal; usually translated as<br />
"one swallow doest not a summer make"]<br />
Other authors have found that Catostomidae are the sistergroup<br />
<strong>of</strong> all other Cypriniformes (e.g. Mayden et al., 2009;<br />
Chen et al., 2009 [labelled as part <strong>of</strong> Cobitoidea in their tree,<br />
but clearly sister-group to all other Cypriniformes]); part <strong>of</strong><br />
these same authors had also found Catostomidae as sistergroup<br />
<strong>of</strong> all other Cobitoidei (Tang et al., 2010) or as sistergroup<br />
<strong>of</strong> Gyrinocheilidae, together being the sister-group to<br />
Cyprinidae (He et al., 2008). In a morphological study, Conway<br />
(2011) found that Catostomidae are the sister-group <strong>of</strong><br />
Gyrinocheilidae, these together are sister-group <strong>of</strong> the remaining<br />
Cobitoidei, which is sister-group <strong>of</strong> Cyprinidae +<br />
Psilorhynchidae.<br />
In view <strong>of</strong> these conflicting conclusions (and awaiting the<br />
next ones ....), I follow Šlechtová et al. (2007) and exclude<br />
Catostomidae from Cobitoidei. Among the above analyses,<br />
theirs is based on the largest number <strong>of</strong> genera and species<br />
<strong>of</strong> Cobitoidei.<br />
Content. — First, the following checklist is far from an ultimate<br />
inventory <strong>of</strong> the loaches <strong>of</strong> the world: it merely presents<br />
what is known today. Many discoveries are still ahead<br />
<strong>of</strong> us and a great amount <strong>of</strong> work remains to be done before<br />
we reach an acceptable level <strong>of</strong> knowledge. I am aware <strong>of</strong><br />
the existence <strong>of</strong> about 100 unnamed species which, hopefully,<br />
should be described in the next few years, from throughout<br />
the range <strong>of</strong> the superfamily.<br />
This list deals with taxonomy and nomenclature. The two<br />
topics are intimately related but should not be confused.<br />
Taxonomy is about species (the animals) and studies their<br />
distinctness, their morphology, their phylogeny and ultimately<br />
organises them in a system, under scientific or philosophical<br />
concepts that may vary among individual scientists. Nomenclature<br />
is only about the names <strong>of</strong> animals; it is governed by<br />
a set <strong>of</strong> rules (the International Code <strong>of</strong> Zoological Nomenclature;<br />
ICZN, 1999; hereunder Code) intended to be applied<br />
universally. Nomenclature is not concerned by scientific<br />
conclusions and hypotheses, or philosophical concepts,<br />
and leaves total taxonomic freedom to individual scientists;<br />
it is concerned only with the names applied to an animal.<br />
In using the checklist, attention should be paid so as to not<br />
confuse taxonomic and nomenclatural discussions. Taxonomy<br />
is a matter <strong>of</strong> hypotheses and open to discussion, while
nomenclature is a more objective issue, almost administrative,<br />
automatically decided by application <strong>of</strong> the Code.<br />
The checklist aims to include all the known species and their<br />
correct names. Synonymies are usually based on published<br />
information, but in some cases I used personal communication<br />
from colleagues, my own interpretation <strong>of</strong> the literature,<br />
or unpublished personal observations.<br />
The list includes references to the original descriptions <strong>of</strong><br />
all species and their synonyms, <strong>of</strong> all the genera and their<br />
synonyms, and all relevant nomenclatural acts. All the cited<br />
references have been checked in the original publications.<br />
The details that I could not check are marked in bold face. A<br />
few references have been checked by trusted colleagues.<br />
Names based on fossil taxa and which have never been used<br />
for Recent taxa are not included in this checklist.<br />
Taxonomy and systematics have two main goals. One is primarily<br />
<strong>of</strong> academic interest: the study <strong>of</strong> the diversity <strong>of</strong> living<br />
organisms and their phylogenetic relationships. The<br />
other is <strong>of</strong> immediate practical interest: inventories, surveys,<br />
documentation <strong>of</strong> biodiversity, and compilation <strong>of</strong> identification<br />
tools. For the proper management <strong>of</strong> natural resources,<br />
we need information on numbers <strong>of</strong> species and their identification<br />
now, not sometime in the distant future. If definitive<br />
conclusions are not possible with the available data, then<br />
tentative decisions are needed. As for other components <strong>of</strong><br />
environmental management strategies, the precautionary<br />
approach should be the rule. In the present context, in case<br />
<strong>of</strong> doubt on the distinctness <strong>of</strong> two species, the precautionary<br />
approach would be to retain them as distinct awaiting<br />
(possible) further research.<br />
This precautionary approach, however, has its limits. The<br />
development <strong>of</strong> molecular techniques has led some to recognise<br />
as 'species' populations distinguished only by very<br />
few nucleotides; complex statistics have been used to justify<br />
the recognition <strong>of</strong> 'species' otherwise not distinguishable by<br />
human eyes. These species I have not given the benefit <strong>of</strong><br />
the doubt. The now fashionable discussions about 'cryptic'<br />
species do not change the situation very much: Aristotle (384–<br />
322 BC) already commented "one nucleotide does not a species<br />
make", be it ever so cryptic [apocryphal translation <strong>of</strong><br />
Μα χελιδν αρ ο ποιε, usually translated in English as<br />
"one swallow doest not a summer make"]. Further, in fishes,<br />
the 'cryptic' species discovered by molecular techniques that<br />
I have been told about have been cryptic not because taxonomists<br />
could not distinguish them, but because no trained<br />
taxonomist ever had an opportunity to examine them.<br />
Another problem with the precautionary approach in Southeast<br />
Asian fishes is shown by the huge number <strong>of</strong> 'new' species<br />
that have been described from Vietnam in recent years.<br />
The description <strong>of</strong> many is in a quality that makes it impossible<br />
to even guess whether or not they might be valid. Their<br />
identity and possible distinctness will remain in limbo as<br />
long as they are not competently re-described.<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
4<br />
Limitations. — One <strong>of</strong> the limitations <strong>of</strong> the checklist is that<br />
I started work on it around 1980. In 30 years, its scope, coverage<br />
and format unavoidably evolved and this caused slight<br />
internal inconsistencies in formatting. It has been updated<br />
continuously so that the technical content is not affected by<br />
this 'formatting evolution'. Further, between 1980 and today,<br />
three different editions <strong>of</strong> the International Code <strong>of</strong><br />
Zoological Nomenclature (ICZN, 1964, 1985, 1999) have<br />
been in use, which differ slightly. I have tried to update all<br />
entries affected by the changes in the Code but I may have<br />
missed details.<br />
Another limitation is that the core target <strong>of</strong> my work and my<br />
own experience is Southeast Asian freshwater fishes. My<br />
treatment <strong>of</strong> these taxa is probably quite complete. But I am<br />
likely to have missed some literature on the species occurring<br />
outside this area. When I encountered nomenclatural<br />
problems concerning Southeast Asian species I had no hesitation<br />
in introducing changes to clear the problems. When it<br />
came to the same situation with taxa from other areas, I tried<br />
to solve the routine problems but decided not to address the<br />
more complex ones. However, I discuss these cases and their<br />
possible solutions where pertinent.<br />
Families vs. subfamilies. — The family-level organisation<br />
is based on the phylogeny established by Šlechtová et al.<br />
(2007) and Bohlen & Šlechtová (2009), and on my personal<br />
considerations (see below and Appendix). The naming and<br />
ranking <strong>of</strong> supraspecific taxa (genera, tribes, families, etc.)<br />
is partly a matter <strong>of</strong> individual taste, that is, some may prefer<br />
to have, for example, a family Balitoridae divided into subfamilies<br />
A, B, C, or a superfamily Balitoroidea divided into<br />
families A, B, C, or a subfamily Balitorinae divided into tribes<br />
A, B, C. What is important is not the rank, but that the organisation<br />
reflects phylogeny.<br />
The recognition <strong>of</strong> sublineages within a lineage (e.g. subfamilies<br />
within a family) implies that the relationships <strong>of</strong> the<br />
lineage have been resolved and that it is demonstrated that<br />
the lineage is monophyletic. If it is not demonstrated, then<br />
there is no justification to recognise sublineages within this<br />
lineage and they are better either ignored or each recognised<br />
as a separate lineage.<br />
Various published phylogenies (Šlechtová et al., 2007;<br />
Bohlen & Šlechtová, 2009; Chen et al., 2009; Mayden et al.,<br />
2009) are congruent in recognising that Ellopostoma, Barbucca,<br />
Serpenticobitis, Balitoridae and Nemacheilidae form<br />
a monophyletic lineage, but they show different relationships<br />
within this lineage, or between some <strong>of</strong> the branches. In such<br />
a situation I consider that it is not justified to recognise some<br />
lineages at family rank and others at subfamily rank; this is<br />
why I treat all at the same rank. Because <strong>of</strong> their very distinctive<br />
morphology and because they are not positioned<br />
within any <strong>of</strong> the sublineages, Barbucca and Serpenticobitis<br />
are treated at the same rank and two family-group names<br />
(Barbuccidae, Serpenticobitidae) are established for them.<br />
I use only the family rank; in my opinion intermediate ranks<br />
such as subfamily, subtribe, etc., are rarely justified. But
others may disagree and might prefer to recognise a family<br />
Balitoridae with subfamilies Ellopostomatinae, Nemacheilinae,<br />
Balitorinae, Barbuccinae, Serpenticobitinae and Gastromyzontinae;<br />
this makes no difference.<br />
Note that a family-group name retains its original author and<br />
date even if used at different ranks. For example Nemacheilinae<br />
Regan, 1911 retains Regan, 1911 as author even if treated<br />
as family Nemacheilidae or tribe Nemacheilini.<br />
Listed names. — I list all the names I found, including infrasubspecific<br />
names, which are given in their original form.<br />
The headings <strong>of</strong> all generic and specific accounts have the<br />
correct spelling <strong>of</strong> all valid names. In the synonymies, however,<br />
all names are given with their original combination (except<br />
that interpolated subgeneric names are omitted) and with<br />
their original spelling, including misspellings and diacritic<br />
marks [ü, è, ñ, etc.]. Capitalised letters and diacritic marks<br />
are not permitted by the Code (arts. 27, 28, Glossary) in<br />
species- and genus-group names and must be corrected. Incorrect<br />
original spellings are used only in the synonymies<br />
but they have been corrected in all other circumstances, especially<br />
in the discussions under Taxonomic notes and Nomenclatural<br />
notes.<br />
Genera and species. — Genera are listed in alphabetic sequence<br />
within families. Species are listed in alphabetic sequence<br />
within genera.<br />
Entries for genera include the valid name <strong>of</strong> the genus (in<br />
bold, as a heading), the name <strong>of</strong> the genus with the spelling<br />
in the original description, the author, the year <strong>of</strong> publication,<br />
and the number <strong>of</strong> the page with principal information.<br />
This is followed by information on possible subgeneric status<br />
in the original description, type species, mode <strong>of</strong> designation,<br />
information on possible nomenclatural acts associated<br />
with the name, and grammatical gender. This information<br />
is provided for all names considered to be synonyms, in chronological<br />
sequence.<br />
Entries for species include the valid name <strong>of</strong> the species (in<br />
bold, as a heading), the name <strong>of</strong> the species with the spelling<br />
in the original description, the author, the year <strong>of</strong> publication,<br />
the number <strong>of</strong> the page on which the actual description<br />
starts (or where the elements necessary to make the name<br />
available occur) and the number <strong>of</strong> the main illustration<br />
(ignoring those showing maps, anatomical details, portrait<br />
<strong>of</strong> collector, etc.). This is followed by a block in parentheses<br />
with information on type locality, primary types and possible<br />
nomenclatural acts associated with the name. If the name<br />
is based largely or totally on references to the older literature,<br />
this information is listed first in the block in parentheses.<br />
The block in parentheses also includes information on<br />
the spelling <strong>of</strong> the end <strong>of</strong> the name associated with the three<br />
grammatical genders <strong>of</strong> genera (see explanation below). This<br />
information is provided for all names considered to be synonyms,<br />
in chronological sequence.<br />
Transliteration <strong>of</strong> non-Latin alphabets. — Author names,<br />
place names, and journal names in non-Latin alphabets, and<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
5<br />
in languages using other notations, have been transcribed;<br />
titles <strong>of</strong> books and papers have been translated. When a transliteration<br />
is used in the original work (e.g. in the text, in an<br />
abstract, in a table <strong>of</strong> contents), the same spelling is used<br />
here. There are some inconsistencies as it happens that transliterations<br />
or translations used in abstracts or tables <strong>of</strong> contents<br />
may be different from the actual title <strong>of</strong> a paper. Frequently,<br />
transliteration systems have changed with time and<br />
no standardisation has been attempted here. Older bibliographies<br />
or indexes may have used earlier transliteration systems<br />
and I consider that a standardised use could actually<br />
complicate bibliographic search, especially for those not<br />
familiar with these languages.<br />
Unfortunately, some accents and diacritic marks may have<br />
disappeared as standard western European keyboards and<br />
s<strong>of</strong>twares do not support them. This especially applies in the<br />
case <strong>of</strong> the Vietnamese alphabet.<br />
Type localities. — The type locality is the locality at which<br />
the holotype, lectotype or neotype was collected. Although<br />
mentioned in the Code, the type locality has no nomenclatural<br />
role. Simply, it is a convenient wording, it is shorter to<br />
say 'type locality' than 'the locality at which the primary namebearing<br />
type was collected', or to give the locality data in<br />
full.<br />
In cases where there is no primary type but a series <strong>of</strong> syntypes<br />
from different localities, the type locality is the sum <strong>of</strong><br />
all the localities <strong>of</strong> the syntypes, and all their localities are<br />
listed (separated by a slash [ / ] where clarity requires it).<br />
Localities are usually given with the original spelling; this<br />
sometimes results in different spellings being used for the<br />
same locality under different headings; I have tried to introduce<br />
some consistency, but only in cases where I was certain<br />
that the different spellings were really referring to the same<br />
place, or when the different spellings were used for the locality<br />
<strong>of</strong> the very same specimen, or referring to the very same<br />
bibliographic source.<br />
Alternative spellings, modern equivalents, present names <strong>of</strong><br />
political entities, transliterations and coordinates (if any) have<br />
been added in square brackets, but this has not been systematically<br />
attempted. Most text in square brackets is not from<br />
original authors.<br />
Locality descriptions have been translated into English when<br />
possible and/or justified. In some cases, words meaning river,<br />
lake, etc. are part <strong>of</strong> the name in the original language<br />
and they have not been deleted in order to avoid ambiguities<br />
when using local maps (but the word river, lake, etc. has<br />
been added). Local names have been used, except for a few<br />
well known rivers and lakes with a common English name<br />
used in international literature (e.g. Mekong, Irrawaddy,<br />
Salween, Ganges, Red River). For most localities, when feasible<br />
I have tried to add information on present political entities<br />
(country, province, state, etc.) and river drainages as<br />
an aid to the reader. For larger topographic features which<br />
have several different names, a single one has been consistently<br />
used; this especially applies to those extending in dif-
ferent countries (e.g. Mekong River); I usually retained the<br />
name easiest to find for readers not familiar with local toponymy<br />
or, when it exists, the English name used in international<br />
literature (e.g. Salween River and not Nu Jiang,<br />
Salawin, Thanlwin, Salouen, or fGyl mo rNGul chu [a Tibetan<br />
transcription]; Irrawaddy and not Ayeyarwady; Red River<br />
and not Song Hong, Yuan Jiang or Fleuve Rouge).<br />
Infrasubspecific names and nomina nuda having no nomenclatural<br />
status, they do not have type specimens and therefore<br />
do not have a type locality and I thus list only a 'locality',<br />
when justified. Localities are usually not indicated for<br />
infrasubspecific names based on aberrant specimens; they<br />
are given only if the name has been created for a particular<br />
geographical area.<br />
When a neotype has been designated, the type locality is the<br />
locality <strong>of</strong> the neotype. The original type locality (the locality<br />
<strong>of</strong> the primary type mentioned in the original description),<br />
if different, is usually listed in square brackets in order<br />
not to lose the information.<br />
When a lectotype has been designated, the type locality is<br />
the locality <strong>of</strong> the lectotype. The original type locality [the<br />
sum <strong>of</strong> the localities <strong>of</strong> all the syntypes mentioned in the<br />
original description], is not listed since it is no longer relevant.<br />
Available name. — An available name is a name that satisfies<br />
the criteria <strong>of</strong> the Code and may be used for a valid<br />
species. An available name is not automatically a valid name.<br />
Valid name. — A valid name is the correct name applied to<br />
a species. To be valid, a name must first be available. But an<br />
available name is not automatically valid; junior synonyms<br />
are available names but invalid. A 'valid name' should not be<br />
confused with a 'valid species'.<br />
Nomen oblitum, nomen protectum. — The earliest <strong>of</strong> two<br />
synonyms always has priority. In some circumstances defined<br />
by Code art. 23.9. it is possible to ignore an old name<br />
and instead use a younger synonym. The older name is then<br />
called nomen oblitum (forgotten name) and the youger one<br />
is called nomen protectum (protected name). To be declared<br />
nomen oblitum, a name must have been unused after 1899<br />
and the nomen protectum must have been used at least 25<br />
times in the last 50 years.<br />
Incertae sedis, genera inquirenda, species inquirendae,<br />
nomina dubiae. — Incertae sedis are valid family, genera<br />
and species <strong>of</strong> uncertain taxonomic position. Species incertae<br />
sedis are listed at the beginning <strong>of</strong> the family they belong<br />
to. Sometimes they are placed in the genus in which my experience<br />
or the experience <strong>of</strong> knowledgeable colleagues suggests<br />
they belong. Alternatively, for species placed in genera<br />
where there is a suspicion they do not belong, the generic<br />
name is placed in single quotation marks (e.g. 'Genus'<br />
species), sometimes with a comment under Taxonomic notes.<br />
The fate <strong>of</strong> a species incertae sedis is to be placed in a genus.<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
6<br />
A species inquirenda (plural: species inquirendae) is a species<br />
<strong>of</strong> doubtful identity. Often they can be placed in a genus<br />
but the description and the known material do not allow a<br />
decision as to whether or not it is valid. These names are<br />
listed immediately under the heading <strong>of</strong> the genus to which<br />
they belong. Species inquirendae that cannot be placed in<br />
any genus are listed immediately under the heading <strong>of</strong> the<br />
family to which they belong. A species inquirenda may have<br />
great similarity to a valid species but its identity may remain<br />
open to doubt; these are listed in the synonymy <strong>of</strong> that species,<br />
and are indicated by a question mark in front <strong>of</strong> the<br />
name. Some species inquirendae are poorly described but<br />
are nevertheless tentatively accepted as possibly valid, for<br />
example because an illustration in the original description<br />
suggests they may be valid; awaiting confirmation or a usable<br />
description, they are listed as 'normal' species but with<br />
a question mark. It is noteworthy that a substantial number<br />
<strong>of</strong> the taxa described from Vietnam in the last 15 years falls<br />
into the category species inquirendae. The fate <strong>of</strong> a species<br />
incertae sedis is that future studies will allow to decide in<br />
which genus or family to place them.<br />
A species inquirenda should not be confused with a nomen<br />
dubium. A nomen dubium (plural: nomina dubia) is a name<br />
<strong>of</strong> doubtful application that is impossible to link with a known<br />
species, or may apply to several species, etc. Typically, a<br />
nomen dubium would have been described in the 18 th or 19 th<br />
century, with a few laconic sentences including no diagnostic<br />
characters usable today, may be based on a painting or on<br />
artificially prepared specimens (examples include species<br />
described from Chinese paintings in the 19 th century; these<br />
paintings usually were not based on a given specimen but<br />
<strong>of</strong>ten were an artistic or idealised view <strong>of</strong> the species, copied<br />
from earlier classical paintings, or sometimes simply imaginary).<br />
The fate <strong>of</strong> a nomen dubium is not to remain so, but to<br />
become either a valid name or a synonym after either taxonomic<br />
examination or appropriate nomenclatural decisions.<br />
A genus inquirendum (plural: genera inquirenda) is a generic<br />
name that can be placed in a family but whose description<br />
and associated species (usually species inquirendae and nomina<br />
dubia) do not allow a decision as to whether or not it is<br />
valid.<br />
Nominal species. — Any <strong>of</strong> the available names created for<br />
a species, irrespective <strong>of</strong> its validity. If a valid species has x<br />
synonyms, the valid name and the x synonyms are x+1 nominal<br />
species.<br />
NAME-BEARING TYPES<br />
It is not the place here to detail all basic principles <strong>of</strong> nomenclature<br />
(for details, see Kottelat, 1997; Kottelat & Freyh<strong>of</strong>,<br />
2007). It is however necessary to clarify a few concepts<br />
that are <strong>of</strong>ten misunderstood.<br />
Type specimens. — Each species-group name has a type.<br />
The type <strong>of</strong> a species name is the specimen on which the<br />
name is based; the phrase name-bearing type is more appro-
priate but, in order to simplify texts, is not usually used. The<br />
type specimen is the type <strong>of</strong> a name, not <strong>of</strong> a species. It is<br />
therefore erroneous to understand the type as a 'model' representation<br />
<strong>of</strong> a species or a specimen to which all specimens<br />
must be identical in order to be called the same species.<br />
The type concept is exclusively a nomenclatural standard<br />
and totally independent <strong>of</strong> any taxonomic judgements<br />
or philosophical theory. The type is only used to objectively<br />
relate to which species the name must be applied. After its<br />
original description, if a species X is redescribed, re-defined<br />
or re-diagnosed and it is found that the type specimen <strong>of</strong><br />
name Y belongs to that species, then this fixes Y as the name<br />
to be applied to species X. If type specimens <strong>of</strong> more than<br />
one name are found to belong to that same species X, this<br />
makes these names synonyms.<br />
Only primary types (name bearing types) are listed here.<br />
Primary types belong to the categories holotype, lectotype,<br />
neotype and syntypes. Other type categories recognised by<br />
the Code are paratypes and paralectotypes but have no nomenclatural<br />
status. Other 'type' categories (e.g. allotypes,<br />
topotypes, paratopotypes, paraneotypes) are not recognised<br />
by the Code, should not be used and are ignored here. Among<br />
them, the word allotype is sometime used to designate one<br />
<strong>of</strong> the paratypes <strong>of</strong> a sex different from that <strong>of</strong> the holotype;<br />
the word topotype is used as a shortened way to say "specimens<br />
collected at the locality where the primary type was<br />
collected".<br />
The holotype is the specimen that has been explicitly designated<br />
so (or by a similar wording) in the original description,<br />
or if there is clear evidence that the author based the<br />
nominal species on a single specimen. There is only one<br />
holotype per nominal species. In all cases where there is clear<br />
evidence that the author based the species name on more<br />
than one specimen (including literature records) but did not<br />
designate a holotype, then all these specimens are syntypes.<br />
If it is not possible to determine from the original description<br />
if a name is based on one or several specimens, I am<br />
usually listing 'syntypes' or 'holotype ?').<br />
If the nominal species is based on a specimen explicitly<br />
designated as holotype and a number <strong>of</strong> additional specimens<br />
are also explicitly designated as types, these are<br />
paratypes; allotypes are thus paratypes. The sum <strong>of</strong> the holotype<br />
and paratypes or the sum <strong>of</strong> the syntypes is called<br />
type series.<br />
In cases where there is no holotype but only a series <strong>of</strong> syntypes,<br />
one <strong>of</strong> the syntypes may be selected as lectotype; it<br />
then has the same value as the holotype. The remaining syntypes<br />
then become paralectotypes and lose their status as<br />
primary types. Lectotypes are designated when it is demonstrated<br />
or suspected that the type series includes more than<br />
one species; it allows the name to be definitively fixed to the<br />
nominal species to which the lectotype belongs. Incidentally,<br />
the designation <strong>of</strong> a lectotype also restricts the type locality<br />
to the locality <strong>of</strong> the lectotype, no longer the sum <strong>of</strong> the<br />
localities <strong>of</strong> the syntypes. Paratypes and paralectotypes are<br />
not listed hereunder as they are not name-bearing types.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
7<br />
If none <strong>of</strong> the specimens <strong>of</strong> the original type series remains,<br />
or if the holotype or lectotype no longer exist (they have not<br />
been preserved, are lost, or destroyed) and if the name cannot<br />
be unambiguously linked to a valid species, then (and<br />
only then) a specimen can be designated as neotype that will<br />
have the same function as the holotype. All designations <strong>of</strong><br />
neotypes that do not fully satisfy these conditions are invalid<br />
and must be ignored. Incidentally, the designation <strong>of</strong> a<br />
neotype also fixes the type locality to the locality <strong>of</strong> the neotype.<br />
A number <strong>of</strong> neotype designations are invalid because the<br />
need for a neotype is not stated or demonstrated. This requirement<br />
did not exist in the 1985 Code (art. 75(b)) and an<br />
implicit justification was enough. The requirement became<br />
explicit in the 1999 Code, with an added clause (art. 75.3.1)<br />
requiring "a statement that [the neotype] is designated with<br />
the express purpose <strong>of</strong> clarifying the taxonomic status or the<br />
type locality <strong>of</strong> a nominal taxon".<br />
After the original description, it may be necessary for later<br />
authors to re-examine the primary type <strong>of</strong> a nominal species<br />
in order to decide to which taxonomic species it applies, for<br />
example in cases when several similar species are later discovered<br />
and the original description does not mention the<br />
characters now decisive to determine to which <strong>of</strong> these species<br />
the name must be applied. It is, however, not a necessity<br />
to examine a primary type if the original description provides<br />
all the information needed for identification. In fact,<br />
types may be fragile specimens, and they should not be handled<br />
if not justified, and persons without experience should<br />
not be permitted to handle them. Primary types must be deposited<br />
in museums or other responsible institutions and with<br />
staff able to conserve them and make them accessible to later<br />
researchers. Even if there is political pressure in some<br />
countries to consider that types are a national property, types<br />
do not belong to a country but to science and must be accessible<br />
to competent scientists irrespective <strong>of</strong> their nationality.<br />
Neotypes must be deposited in a museum (Code art. 75.3.7).<br />
A number <strong>of</strong> species described by earlier authors do not have<br />
known types or they have been lost since the original description.<br />
This does not affect the availability <strong>of</strong> a name. For<br />
example, a specimen described in the field and later eaten<br />
by a hungry author remains the type specimen. Or a specimen<br />
used as model for a figure remains the type specimen,<br />
even if it has not been preserved.<br />
When known, institutions in which primary types are deposited<br />
are listed, together with register numbers and, when<br />
known, the number <strong>of</strong> specimens in square brackets (example:<br />
"AAA 1234 [2], AAA 1233 [1]"). When the primary<br />
types were deposited in a collection but cannot presently be<br />
located, the institution is listed as they may still be present<br />
(misidentified, misplaced, uncatalogued), or as a starting<br />
point for further search. The source for the catalogue number<br />
is given when it is not the original description; besides,<br />
many catalogue numbers listed in the original descriptions<br />
have also been checked in published catalogues or in the<br />
institutions themselves. When there is a series <strong>of</strong> syntypes, I
listed those I could trace in the literature, but made no effort<br />
to trace the whole series; this would have been tedious, many<br />
<strong>of</strong> them having possibly been later used for exchanges between<br />
institutions, etc. NT indicates that there is no (or apparently<br />
no) preserved type material, LU that there was apparently<br />
preserved type material but that its whereabouts are<br />
not known. A question mark in front <strong>of</strong> the abbreviation <strong>of</strong><br />
an institution indicates that the type(s) is possibly there or<br />
that the type status <strong>of</strong> the specimen is not certain.<br />
Institutional abbreviations used in the text are listed below.<br />
For institutions for which no abbreviations have been used<br />
in literature, the abbreviations follow current use by workers<br />
at these institutions (where possible) or Leviton et al.<br />
(1985; Leviton & Gibbs, 1988) or Eschmeyer & Fricke<br />
(2010). I did not automatically follow Leviton et al. and Eschmeyer<br />
& Fricke as a standard because for non-US collections<br />
the abbreviations they list are <strong>of</strong>ten not those actually<br />
used by the institutions themselves. In case the abbreviations<br />
used in these lists differ from those used by workers at<br />
these institutions, I retain the second one (as long as they<br />
make sense and do not represent personal, bureaucratic or<br />
chauvinistic idiosyncrasy). I use the word abbreviation, not<br />
acronym. The word acronym means an abbreviation that can<br />
be pronounced as a word, which is not the case here.<br />
Type species. — Each genus-group name has a type species.<br />
The type species <strong>of</strong> a genus name is the species whose name<br />
determines the validity <strong>of</strong> a genus. If several species are<br />
placed in genus X, with type species Xx and this genus is<br />
later divided into two genera, the genus which include species<br />
Xx will continue to be genus X while the other genus<br />
will have another name. If two genera have the same type<br />
species, they are objective synonyms.<br />
Type genus. — Each family-group name has a type genus.<br />
The type genus is the genus whose name has been used to<br />
form the name <strong>of</strong> the family. For example Cobitis is the name<br />
<strong>of</strong> the genus used to form the family group names Cobitidae,<br />
Cobitoidei, Cobitinae, etc. For nomenclature purposes these<br />
three words are a single name. Whatever the rank within the<br />
family-group, a given family-group name retains the same<br />
author and date. In this example, Cobitidae Swainson, 1838,<br />
Cobitoidei Swainson, 1838, etc.<br />
ENDING AND SPELLING OF SPECIES NAMES<br />
The following discussions refers only to scientific names,<br />
which are written in a language supposed to be Latin. When<br />
the system <strong>of</strong> binominal nomenclature was created, the genus<br />
was the important entity and the species was <strong>of</strong> secondary<br />
significance. As a result, the names <strong>of</strong> the species were<br />
made <strong>of</strong> a noun (the genus name) and a qualifying word (the<br />
species name), in most cases an adjective or a noun in the<br />
genitive. As genus names are nouns, they have a grammatical<br />
gender (masculine, feminine or neuter). In Latin, French,<br />
German and most western languages the ending <strong>of</strong> adjectives<br />
varies to agree with the gender <strong>of</strong> the noun (English is<br />
a notable exception in which adjectives are not variable).<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
8<br />
When new research shows that a species must be transferred<br />
from one genus to another, if the two genera have names<br />
with different grammatical genders, it may happen that the<br />
ending <strong>of</strong> the name <strong>of</strong> the species must be changed to agree<br />
in gender with the name <strong>of</strong> the genus (Code art. 31.2).<br />
This may seem complicated, but actually it does not require<br />
to learn Latin, but only to follow a handful <strong>of</strong> trivial rules:<br />
a) check the gender <strong>of</strong> the name <strong>of</strong> the genus in the original<br />
publication or on a reliable list;<br />
b) if the species name is a noun, its spelling remains unchanged;<br />
c) if the species name is an adjective, the name must agree in<br />
gender with the genus.<br />
Besides, answers to most questions can be found in dictionaries<br />
and, in case <strong>of</strong> doubt, it does not take a great effort to<br />
ask knowledgeable colleagues. In the last resort, if nobody<br />
knows or if it seems too time consuming, one should treat<br />
the name as a noun in apposition and retain the original spelling<br />
or follow common practice.<br />
Hereunder, the grammatical gender <strong>of</strong> all genus names is<br />
listed. For all species names, the list includes the three possible<br />
endings (for the three genders), so that it is instantly<br />
possible to check the correct ending when moving a species<br />
from one genus to another. The names marked 'indeclinable'<br />
never change; for those which change, the ending in masculine,<br />
femine and neuter is indicated. For example, for maculatus,<br />
"-us, -a, -um" means that if the name is placed in a<br />
genus with a masculine name, the species must be spelled<br />
maculat-us, if placed in a genus with a feminine name it<br />
must be spelled maculat-a and if placed in a genus with a<br />
neuter name it must be spelled maculat-um.<br />
Out <strong>of</strong> about 1391 available names, 439 are adjectives ending<br />
in -us, 317 adjectives ending in -is, 7 other declinable<br />
adjectives, 306 nouns in genitive, 154 compound nouns, 143<br />
nouns in apposition, 14 indeclinable adjectives, and 11<br />
indeclinable participles.<br />
Common situations. — The basic principles above and the<br />
information in the list below are enough to determine the<br />
correct ending <strong>of</strong> names in any combination. For those interested<br />
in more details, this chapter presents examples <strong>of</strong> the<br />
main types <strong>of</strong> name constructions, how to distinguish which<br />
existing names are declinable and which are not, and the<br />
status <strong>of</strong> corrections <strong>of</strong> the original spelling. Note that this is<br />
for existing names and not for creating new names. These<br />
common cases cover the vast majority <strong>of</strong> existing names; a<br />
few rare cases make exceptions and are not discussed.<br />
1. Names based on personal names<br />
1.1. Names in the genitive case are indeclinable. They<br />
mean "<strong>of</strong>".<br />
1.1.1. If the name is based on a modern name, the<br />
genitive ending is -i for men (e.g. bleekeri, <strong>of</strong><br />
Bleeker); -ae for woman (e.g. pfeifferae, <strong>of</strong> Mrs.<br />
Pfeiffer), -arum for several women together (e.g.<br />
mariarum, <strong>of</strong> Mary Inger and Mary Chin), -orum<br />
for several men together (e.g. grummorum, <strong>of</strong> the
others Grigorii and Vladimir Grumm-Grzhimaylo)<br />
or for men and women together (e.g, potaninorum,<br />
<strong>of</strong> Grigorii Potanin and Aleksandra Potanina,<br />
baluchiorum, <strong>of</strong> the inhabitants <strong>of</strong><br />
Balochistan).<br />
1.1.1.1. Exception: names <strong>of</strong> men ending in -a may<br />
be considered Latinized and become -ae (e.g.<br />
vinciguerrae, <strong>of</strong> Vinciguerra).<br />
1.1.1.2. If the name ends with -i, the ending is -ii<br />
(e.g. choii, <strong>of</strong> Choi; modiglianii, <strong>of</strong> Modigliani).<br />
1.1.1.3. Names may be formed by adding the endings<br />
-i, -ae, etc. to the name or to the stem <strong>of</strong><br />
the name. Example: agassizi is based on Agassiz<br />
and agassii is based on the stem agassi-<br />
[this corresponds with the pronunciation, since<br />
the final -z in Agassiz is not pronounced in<br />
French in the area <strong>of</strong> Switzerland where he was<br />
born].<br />
1.1.2. If the name is based on a Latin name, or the<br />
Latinized form <strong>of</strong> a modern name, the name is<br />
formed <strong>of</strong> the Latin root and the endings as above<br />
(<strong>of</strong>ten becomes -ii, -is, -iorum, -iarum). Examples:<br />
cuvieri is based on Cuvier's name, cuvierii<br />
is based on the Latinized form Cuvierius.<br />
1.2. Nouns in apposition are indeclinable (but the Code<br />
recommends that species-group names should not be<br />
created as personal names in apposition as this may<br />
lead to confusion with the author <strong>of</strong> the name).<br />
1.3. Adjectives formed on a personal name are declined<br />
to agree in gender with the genus name. Examples:<br />
bleekerianus, -a, -um (P. Bleeker; means Bleekerian),<br />
pavianus, -a, -um (A. Pavie).<br />
2. Names based on geographical names. Same rules as<br />
names based on personal names (see 1).<br />
2.1. If the name is in the genitive case, it is indeclinable.<br />
Examples: italiae (<strong>of</strong> Italy), cyri (<strong>of</strong> Cyrus, the Latin<br />
name <strong>of</strong> Kura River).<br />
2.2. If the name is in apposition, it is indeclinable: Examples:<br />
Nemacheilus marang (Marang loach; for<br />
Marang karst).<br />
2.3. If the name is an adjective, it is declined to agree in<br />
gender with the genus name. The most common cases<br />
are:<br />
2.3.1. names based on classical and modern Latin adjectives.<br />
Examples: hispanicus, -a, -um (Hispanic,<br />
Spanish), helveticus, -a, -um (Helvetian,<br />
Swiss), sumatranus, -a, -um (Sumatran);<br />
2.3.2. names based on modern place names with the<br />
addition <strong>of</strong> the ending -ensis, -ensis, -ense). Example:<br />
yunnanensis, -is, -e.<br />
2.3.2.1. The final letter(s) can be omitted, or a<br />
letter can be added (usually to smoothen pronunciation).<br />
Examples: borneensis instead <strong>of</strong><br />
borneoensis (Bornean), sumatrensis instead <strong>of</strong><br />
sumatraensis (Sumatran; but sumatranus<br />
would be preferable), tientainensis instead <strong>of</strong><br />
tientaiensis (<strong>of</strong> Tientai).<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
9<br />
3. Latin adjectives. Names that are Latin adjectives or participles,<br />
or end with Latin adjectives or participles, are declined<br />
to agree in gender with the name <strong>of</strong> the genus. (For<br />
nomenclature purposes, only the nominative case is relevant;<br />
Latin has five more cases, but they are irrelevant here).<br />
3.1. The most common declinable endings are:<br />
3.1.1. -us, -a, -um. Examples: maculatus, maculata,<br />
maculatum (spotted), barbatus, barbata, barbatum<br />
(bearded);<br />
3.1.2. -is, is, -e. Examples: lateralis, lateralis, laterale<br />
(lateral); meridionalis, meridionalis, meridionale<br />
(southern);<br />
3.1.3. -ger, -gra, grum. Example: niger, nigra, nigrum<br />
(black);<br />
3.1.4. -ger, -gera, -gerum. Example: filiger, filigera,<br />
filigerum (that carries a filament);<br />
3.1.5. -fer, -fera, -ferum. Example: aurifer, aurifera,<br />
auriferum (that carries or contains gold).<br />
3.2. When declined, some adjectives are identical in the<br />
three genders in the nominative case. They are labelled<br />
here as indeclinable. Examples: elegans (nice),<br />
splendens (brilliant, shining), pallens (pale). Some<br />
common endings <strong>of</strong> indeclinable adjectives are:<br />
3.2.1. adjectives ending in -oides (sometimes -ides).<br />
Examples: siluroides (silurus-like), elongatoides<br />
(like [Cobitis] elongata), misgurnoides (misgurnus-like),<br />
salmonides (salmon-like);<br />
3.2.2. adjectives ending in -color. Examples: discolor<br />
(differently coloured), unicolor (<strong>of</strong> a single<br />
colour).<br />
3.3. Participles (verbs in the conjugation tense ending in<br />
-ing in English) usually end in -ens or -ans and are<br />
indeclinable in the nominative case. Examples: alternans<br />
(alternating), ridens (smiling).<br />
3.4. If an adjective has been erroneously declined in<br />
cases other than nominative it is treated as a noun in<br />
apposition. Example: albicoloris (should have been<br />
albicolor, indeclinable).<br />
4. Latin nouns. Names that are Latin or Latinized nouns are<br />
indeclinable. Examples: Schistura cataracta (cataract), Ellopostoma<br />
mystax (mustache).<br />
5. Compound names (noun-phrases)<br />
5.1. A compound name made <strong>of</strong> two nouns is a noun and<br />
indeclinable (examples: ophiolepis, from ophius,<br />
snake, and lepis, scale, with snake scales; bellibarus,<br />
from the incorrectly Latinized English nouns belly<br />
and bar, for the six bars on the belly); names made <strong>of</strong><br />
an adjective and a noun are also nouns and indeclinable<br />
(example: alticorpus, from altus, high, and corpus,<br />
body, with high body; flavicauda, from flavus,<br />
yellow, and cauda, tail, with yellow tail).<br />
5.2. If a compound name ends with a non-Latin word, it<br />
is indeclinable. This applies to all languages. Some<br />
Greek words are used as ending <strong>of</strong> compound names.<br />
The Code makes a distinction between Greek words<br />
and Latinized Greek words. For example kephale is<br />
a Greek word (meaning head) while cephalus is Latinized<br />
Greek; oura is a Greek word (meaning tail)
while ura is Latinized Greek. If the compound name<br />
ends with a Greek word (thus non-Latin), the name is<br />
indeclinable, irrelevant <strong>of</strong> whether it is a noun or an<br />
adjective (examples: hipporhynchos, from hippos,<br />
horse, and rhynchos, snout or snouted, that has a horse<br />
muzzle; microphthalmos, from micros, small, and<br />
ophthalmos, eye, small-eyed). If the compound name<br />
ends with a Latinized Greek word, it is indeclinable<br />
if that last word is a noun, or declinable if that last<br />
word is an adjective.<br />
Many (not all) compound names ending with a<br />
Latinized Greek word are adjectives. Among the most<br />
common ones in ichthyology are -ophthalmus, -a,<br />
-um (-eyed; example: microphthalmus, small-eyed),<br />
-rhynchus, -a, -um (-snouted; example: macrorhynchus,<br />
big-snouted), -cheilus, -a, -um (-lipped; example:<br />
pachycheilus, thick-lipped); -cephalus, -a, -um<br />
(-headed; example: pachycephalus, broad-headed),<br />
-stomus, -a, -um (-mouthed; example: macrostomus,<br />
large mouthed), -urus, -a, -um (-tailed; example: stenurus;<br />
slender-tailed), -gnathus, -a, -um (-jawed; example:<br />
prognathus, prognathous). However, not all<br />
Latinized Greek words in compound names are adjectives.<br />
Some are nouns; examples: -pomus (from poma,<br />
cover, lid, opercle; the adjective is -pomatus, -a, -um),<br />
soma (body; the adjective is -somatus, -a, -um). Some<br />
nouns may be both; example: the Latinized adjective -<br />
pterus, -a, -um, winged, and the Latin noun ptera.<br />
5.3. A compound name made <strong>of</strong> two words <strong>of</strong> which the<br />
last word is an adjective qualifying the genus name is<br />
treated as an adjective and has to agree in gender (see<br />
3). Examples: in Xus flavicaudatus (yellow-tailed<br />
Xus), caudatus is an adjective and flavicaudatus must<br />
agree in gender, in Xus flavicauda (yellow tail Xus,<br />
Xus with yellow tail), cauda is a noun in nominative<br />
and flavicauda does not have to agree in gender with<br />
the genus name.<br />
5.4. A compound name made <strong>of</strong> two words <strong>of</strong> which the<br />
last word is an adjective qualifying the first word is a<br />
noun in apposition and indeclinable. Examples: in<br />
Nemacheilus lacusnigri, nigri (genitive <strong>of</strong> niger) qualifies<br />
lacus (lake) and lacusnigri is indeclinable; the<br />
name means loach <strong>of</strong> Black Lake and not black lakeloach.<br />
6. Names that can be a noun or an adjective in Latin<br />
6.1. If it was first used as an adjective, it is treated as an<br />
adjective and must be declined;<br />
6.2. If it was first used as a noun, it is a noun and is<br />
indeclinable;<br />
6.3. If its first use does not allow a decision as to whether<br />
it was treated as a noun or an adjective, then it is<br />
treated as a noun in apposition and is not declinable.<br />
Note that in some cases, although the spellings <strong>of</strong> the<br />
noun and <strong>of</strong> the adjective do not differ, syntax and<br />
meaning may allow to make a decision.<br />
6.4. Example 1. In Barbatula sellaefer, Barbatula is feminine,<br />
sellaefer can be a noun in apposition or an adjective<br />
in the masculine nominative; because in Barbatula<br />
sellaefer it is spelt sellaefer it is a noun in ap-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
10<br />
position and indeclinable; if it had been spelt sellaefera<br />
it would have been an adjective and declinable.<br />
6.5. Example 2. Imaginary case: in Nemacheilus criniger,<br />
Nemacheilus is masculine, criniger can be a noun<br />
in apposition or an adjective in nominative masculine<br />
and the author did not indicate which one he used;<br />
therefore it is treated as a noun in apposition.<br />
7. Non-Latin words. Names (or the final word <strong>of</strong> a compound<br />
noun) that are not Latin words are treated as nouns in<br />
apposition and are indeclinable. Examples: Botia zebra (zebra<br />
comes from an African language), Botia helodes (helodes<br />
is a Greek word meaning marshy).<br />
8. Distinction between some adjectives and nouns<br />
8.1. There is <strong>of</strong>ten confusion between a noun and adjectives<br />
based on the same noun. The most common ones<br />
are compound names ending in -atus, -ata, -atum.<br />
Example: lateristriga and lateristrigata in which<br />
striga is a noun (stripe) and strigata is an adjective<br />
(striped). A simple 'rule' to decide if the last word is<br />
an adjective or a noun (beside checking a dictionary)<br />
is that words ending in -atus, -ata, -atum are usually<br />
adjectives and declinable (flavilinea [with yellow<br />
stripe] vs. flavilineata [yellow-striped], anguillicauda<br />
[with eel-tail] vs. anguillicaudata [eel-tailed]).<br />
There are exceptions to the 'rule'.<br />
8.2. If a Latin adjective has been explicitly treated as a<br />
noun in apposition in the original description (Code<br />
art. 11.9.1.2.), then it is indeclinable.<br />
8.3. If a Latin noun has been explicitly treated as an adjective<br />
in the original description (Code art. 11.9.1.1),<br />
then it is treated as adjective and is declinable. Example:<br />
names ending in -cola (inhabitant) are nouns<br />
and indeclinable; but some names have been created<br />
as -colus with the erroneous statement that they were<br />
adjectives (e.g. Nemacheilus arenicolus) and must be<br />
retained as adjectives and declined.<br />
8.4. If a name has been treated as a Latin noun in the<br />
original description but is not Latin, then it is a noun<br />
in apposition and indeclinable.<br />
8.5. Some words are adjectives in Latin, but in ichthyology<br />
they are used as nouns. These are treated as noun<br />
in apposition. The most common ones are ventralis<br />
and pectoralis, which originally are the adjectives<br />
ventral and pectoral but in ichthyology are commonly<br />
used as substantives to mean the ventral (pelvic)<br />
and pectoral fins. In such cases, they are used as nouns<br />
in apposition (e.g. in longiventralis, longipectoralis);<br />
but in case they had been used as adjectives in the<br />
original descriptions, they would <strong>of</strong> course be treated<br />
as adjectives. In Cobitis pectoralis, pectoralis was<br />
obviously treated as a noun in apposition, otherwise<br />
the name would not make sense. The words analis<br />
(anal fin), dorsalis (dorsal fin) and caudalis (caudal<br />
fin) do not exist in Latin and are treated as nouns in<br />
apposition.<br />
9. Correction <strong>of</strong> spelling <strong>of</strong> names<br />
9.1. The spelling used in the first description making a
name available is called the original spelling. The<br />
original spelling is the correct spelling and cannot be<br />
changed. There are only few exceptions.<br />
9.1.1. If in a later work the spelling is changed unintentionally,<br />
it is called an incorrect subsequent<br />
spelling. If it is changed intentionally, then it is an<br />
emendation. An emendation is justified if it is<br />
made to follow requirements <strong>of</strong> the Code; all other<br />
emendations are unjustified.<br />
9.1.2. Incorrect spellings are not available names (they<br />
do not exist in nomenclature). Unjustified emendations<br />
are available names, although <strong>of</strong>ten invalid.<br />
9.2. The original spelling can be corrected if there is evidence<br />
<strong>of</strong> an inadvertent error. An inadvertent error is<br />
an incorrect spelling not intended by the author. The<br />
Code mentions lapsus calami, copyist's error and<br />
printer's error as examples <strong>of</strong> inadvertent error. This<br />
is sometimes interpreted as a list <strong>of</strong> accepted cases.<br />
The Code lists these as examples, which means that<br />
the list is not exhaustive. Additional cases would include,<br />
for example, damaged font (a case is described<br />
by Bogutskaya et al., 2005), errors introduced by a<br />
translator (see under Triplophysini) or, with modern<br />
technology, a file-conversion problem. An error by<br />
the printer in deciphering a hand-written manuscript<br />
is an inadvertent error, but it is debatable whether an<br />
error by the author in deciphering hand-written information<br />
(his own notes, a place name on a label, or<br />
the signer <strong>of</strong> a letter) is an inadvertent error.<br />
Speech recognition s<strong>of</strong>twares and the commands<br />
'search and replace' in word processors are promising<br />
fields for inadvertent errors.<br />
9.2.1. Latinization errors cannot be corrected. For example,<br />
nujiangensa should have been nujiangensis<br />
but cannot be corrected; the erroneous use <strong>of</strong><br />
ovis (sheep) or ovalis (ovation) instead <strong>of</strong> ovum<br />
(egg, oval) or ovatus (oval) for an animal with an<br />
oval shape cannot be corrected.<br />
9.2.2. Special letters must be replaced (e.g. German<br />
ü, ä, ö become ue, ae, oe) or modified (e.g. Spanish<br />
ñ becomes n, non-German ö becomes o, etc.),<br />
spaces and hyphens are suppressed, capitalised<br />
initials become lower case, numbers must be<br />
spelled out, etc.<br />
9.3. Erroneous spellings <strong>of</strong> names based on personal<br />
names is a potentially sensitive issue, resulting from<br />
a loophole in the Code.<br />
9.3.1. Article 31.1 explains how names based on<br />
names <strong>of</strong> persons must be formed (see 1). Art.<br />
31.1.3. says that a name based on a personal name<br />
formed under arts. 31.1.1 and 31.1.2 (see 1) must<br />
be preserved unless it is incorrect. The corollary<br />
is that if the name is incorrect it must not be preserved.<br />
But this article does not say what to do if<br />
a name is not formed correctly. Common sense is<br />
that "a name must be preserved unless incorrect"<br />
implies that an incorrectly formed name is not to<br />
be preserved, and that the only things that can be<br />
done is to render it correct or to replace it. Art.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
11<br />
32.5 lists the spellings that must be corrected but<br />
the list does not include the incorrect names mentioned<br />
in art. 31.1.3. Some authors argue that these<br />
names therefore cannot be corrected. My interpretation<br />
is that, if a name based on a person's<br />
name must be formed following art. 31.1, those<br />
formed incorrectly must be emended. This is implicit<br />
in art. 31.1 and therefore it was not needed<br />
to repeat it in art. 32.5. Therefore, erroneous original<br />
spellings <strong>of</strong> names based on person's names<br />
must be corrected.<br />
9.3.2. This art. 31.1.3 applies only for the aspects <strong>of</strong><br />
the names formed under art. 31.1.1 and 31.1.2,<br />
that is mainly ending (formed under 1 above). This<br />
means that a name formed, for example, on the<br />
name <strong>of</strong> a woman but with the ending -i must be<br />
corrected in -ae. Bleeker created the name Cobitis<br />
pfeifferi explicitly for Ida Pfeiffer, a woman,<br />
and the name must be emended into C. pfeifferae.<br />
If the gender <strong>of</strong> the person is not known the original<br />
ending does not change.<br />
9.3.3. If the name <strong>of</strong> the person is mentioned (correctly<br />
spelt or not) and the name <strong>of</strong> the species is<br />
misspelt, it must be corrected. Example: Sykes<br />
(1839a) described Cobitis rupelli, which he named<br />
for Eduard Rüppell, whose name is explicitly mentioned.<br />
The omission <strong>of</strong> one 'p' is an inadvertent<br />
error. The spelling <strong>of</strong> the name should have been,<br />
and must be corrected into, ruppelli. [The omission<br />
<strong>of</strong> the umlaut is correct; if Sykes had written<br />
"Rüppell", ü should have been corrected into ue,<br />
but as Sykes used "Ruppell", u should not be corrected].<br />
9.3.4. If the name <strong>of</strong> the person is not mentioned in<br />
the paper and can only be guessed to possibly be<br />
a misspelling <strong>of</strong> a known name, it must remain<br />
unchanged.<br />
9.3.5. If there is external evidence (for example by<br />
the author himself in a later publication) or indication<br />
that the name is formed on a misspelling <strong>of</strong><br />
the name <strong>of</strong> a person unambiguously identifiable,<br />
there are two conflicting opinions. Some authors<br />
consider that the Code art. 32.5.1 does not allow<br />
the emendation since it can be known that the name<br />
is misspelt only by recourse to external information.<br />
Others think that art. 32.5.1 allows to correct<br />
the spelling because to misspell the name <strong>of</strong><br />
a person to which the species is dedicated can only<br />
be an inadvertent error. Art. 32.5.1 lists some kinds<br />
<strong>of</strong> inadvertent errors to be corrected but does not<br />
mention misspelling <strong>of</strong> personal names; this is not<br />
a problem since the Code explicitly states that<br />
these are examples ("such as ..."), and a list <strong>of</strong><br />
examples is, by definition, not exhaustive [see 9.2<br />
above].<br />
A classical example is Barbus schwanenfeldii,<br />
originally described by Bleeker (1854a: 517) and<br />
dedicated to the collector "H. W. Schwanenfeld".<br />
Later, Bleeker (1860d: 63) emended it as<br />
schwanefeldi because the correct spelling <strong>of</strong> this
person's name is Schwanefeld. Elsewhere in the<br />
same issue <strong>of</strong> the journal including the 1854 description,<br />
but not in the same paper (pp. 540, 545),<br />
Schwanefeld's name was spelt correctly, as it is<br />
also in lists <strong>of</strong> members <strong>of</strong> societies (<strong>of</strong> which<br />
Bleeker was president or secretary) and subscribers<br />
to journals (edited by Bleeker). In addition<br />
Schwanefeld was a pr<strong>of</strong>essional relation <strong>of</strong> Bleeker.<br />
The misspelling could not have been intentional.<br />
Despite all this, because we can know that<br />
the name was misspelt only from information external<br />
to the original paper, some conclude that<br />
the name cannot be corrected (this was my earlier<br />
point <strong>of</strong> view; Kottelat, 1999: 595). But, in the<br />
case <strong>of</strong> personal names dedicated to a clearly identified<br />
person, when the original author himself<br />
AMNH American <strong>Museum</strong> <strong>of</strong> Natural History, New York, USA<br />
AMS Australian <strong>Museum</strong>, Sydney, Australia<br />
ANSP Academy <strong>of</strong> Natural Sciences, Philadelphia, USA<br />
ASIZB Academia Sinica, Institute <strong>of</strong> Zoology, Beijing, China<br />
BIKU Institute <strong>of</strong> Biology and Ecology, University <strong>of</strong> Kragujevac,<br />
Kragujevac, Serbia<br />
BLG Biological Laboratory, Sun Yat-Sen University, Guangzhou,<br />
China<br />
BMAM Beijing <strong>Museum</strong> <strong>of</strong> Natural History, Beijing, China<br />
BMNH Natural History <strong>Museum</strong>, London, United Kingdom<br />
CAS California Academy <strong>of</strong> Sciences, San Francisco, USA<br />
CMK Collection <strong>of</strong> Maurice Kottelat, Cornol, Switzerland<br />
CNUC Department <strong>of</strong> Biology, Chonbuk National University,<br />
Jeonju, South Korea<br />
CRG-SAC Conservation <strong>Research</strong> Group, St Albert’s College,<br />
Kochi, Kerala, India<br />
DBJU Department <strong>of</strong> Biology, Ji-Nan University, Guangzhou,<br />
China<br />
DBLU Department <strong>of</strong> Biology, Lanzhou University, Lanzhou,<br />
China<br />
DHISUB Department <strong>of</strong> Hydrobiology and Ichthyology, Faculty<br />
<strong>of</strong> Biology, University <strong>of</strong> S<strong>of</strong>ia, S<strong>of</strong>ia, Bulgaria<br />
DVZUT Department <strong>of</strong> Vertebrate Zoology, Vietnam National<br />
University <strong>of</strong> Hanoi, Hanoi, Vietnam<br />
DZAUT Department <strong>of</strong> Zoology, Aristotle University, Thessaloniki,<br />
Greece<br />
EAWAG EAWAG Forschungszentrum für Limnologie, Kastanienbaum,<br />
Switzerland<br />
FACQR Fishery Administration Center <strong>of</strong> Qujing Region,<br />
Qujing, China<br />
FESC Fisheries Experimental Station, Guangdong, China<br />
[now Pearl River Fishery <strong>Research</strong> Institute, Chinese<br />
Academy <strong>of</strong> Fishery Sciences, Guangzhou, China]<br />
FFRIJ Freshwater Fisheries <strong>Research</strong> Institute <strong>of</strong> Jiangsu Province,<br />
Nanjing, China<br />
FM Department <strong>of</strong> Nature, Fujian Provincial <strong>Museum</strong>,<br />
Fuzhou, China<br />
FMNH Field <strong>Museum</strong> <strong>of</strong> Natural History, Chicago, USA<br />
FRCL Fisheries <strong>Research</strong> Center, Likas, Kota Kinabalu, Malaysia<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
ABBREVIATIONS USED<br />
12<br />
corrected the spelling and when it is shown from<br />
numerous sources that it was misspelt, it is obvious<br />
that it was an inadvertent error and it would<br />
be against common sense not to correct it. I would<br />
be unhappy to see a species name dedicated to<br />
me but with a misspelling <strong>of</strong> my name; and if I<br />
have to write a paper on this species I would certainly<br />
not use a misspelling <strong>of</strong> my own name; I<br />
think that everybody can share these feelings. It<br />
therefore seems that nobody should really complain<br />
if the emended but possibly 'illegal' spelling<br />
is used.<br />
Note that Bleeker's change <strong>of</strong> the original ending<br />
-ii into -i is not permitted and the original ending<br />
-ii remains.<br />
FSJF Fischsammlung Jörg Freyh<strong>of</strong>, Berlin, Germany<br />
GCMNH Government College <strong>Museum</strong> (Natural History),<br />
Lahore, Pakistan<br />
GIF Guangxi Institute <strong>of</strong> Fisheries, Nanning, China<br />
GUIC Ichthyological <strong>Museum</strong>, Department <strong>of</strong> Fisheries, Natural<br />
Resources Faculty, University <strong>of</strong> Guilan, Soumahe<br />
Sara, Iran<br />
HACW Department <strong>of</strong> Fisheries, Huazhong Agriculture College,<br />
Wuhan, China<br />
HFRI Heilongjiang Fisheries <strong>Research</strong> Institute, Harbin,<br />
China<br />
HNUE Department <strong>of</strong> Zoology, Faculty <strong>of</strong> Biology and Agricultural<br />
Technology, Hanoi National University <strong>of</strong> Education<br />
[also Hanoi University <strong>of</strong> Pedagogy], Hanoi,<br />
Vietnam<br />
HRAS Heilongtan Reservoir Administration, Shilin, China<br />
HUIC Ichthyology <strong>Museum</strong>, Department <strong>of</strong> Biology, Hacettepe<br />
University, Ankara, Turkey<br />
HUJ Hebrew University, Jerusalem, Israel<br />
IHB Institute <strong>of</strong> Hydrobiology, Academia Sinica, Wuhan,<br />
China<br />
IRSNB Institut Royal des Sciences Naturelles, Bruxelles, Belgium<br />
ISBB Institutul Stiinte Biologice, Bucuresti, Romania<br />
IZA Dipartimento di Scienze Ambientali, Universita,<br />
L'Aquila, Italy<br />
IZSX Institute <strong>of</strong> Zoology <strong>of</strong> Shanxi, Xian, China<br />
KFRI Kerala Forest <strong>Research</strong> Institute, Peechi, India<br />
KIZ Kunming Institute <strong>of</strong> Zoology, Academia Sinica, Kunming,<br />
China<br />
KU University <strong>of</strong> Kansas Natural History <strong>Museum</strong> and<br />
<strong>Biodiversity</strong> <strong>Research</strong> Center, Lawrence, USA<br />
KUMF Kasetsart University <strong>Museum</strong> <strong>of</strong> Fisheries, Bangkok,<br />
Thailand<br />
KUN Kinki University, Nara, Japan<br />
LFZFC Laboratory <strong>of</strong> Fishes, Zhejiang Fisheries College,<br />
Zhoushan, China [now Zhejiang Ocean University,<br />
Zhoushan, China]<br />
LNHSM Lingnan Natural History Survey and <strong>Museum</strong>, Guangzhou,<br />
China
MCSNC Museo Civico di Storia Naturale, Carmagnola, Italy<br />
MCSNG Museo Civico di Storia Naturale, Genova, Italy<br />
MCZ <strong>Museum</strong> <strong>of</strong> Comparative Zoölogy, Harvard, USA<br />
MGAB Musée d'Histoire Naturelle "G. Antipa", Bucuresti, Romania<br />
MGHNL Musée Guimet d'Histoire Naturelle, Lyon, France<br />
MHNG Muséum d'Histoire Naturelle, Genève, Switzerland<br />
MIZASP <strong>Museum</strong>, Institute <strong>of</strong> Zoology, Academy <strong>of</strong> Sciences,<br />
Pyongyang, North Korea<br />
MMNHN Metropolitan <strong>Museum</strong> <strong>of</strong> Natural History, Nanjing,<br />
China [now Nanjing <strong>Museum</strong>, Chinese Academy <strong>of</strong><br />
Sciences, China]<br />
MMNHS Macedonian <strong>Museum</strong> <strong>of</strong> Natural History, Skopje,<br />
Former Yugoslavian Republic <strong>of</strong> Macedonia<br />
MNCN Museo Nacional de Ciencias Naturales, Madrid, Spain<br />
MNH Magyar Nemzeti Muzeum, Budapest, Hungary<br />
MNHN Muséum National d'Histoire Naturelle, Paris, France<br />
MNHV <strong>Museum</strong> <strong>of</strong> Natural History, Varna, Bulgaria<br />
MSINR <strong>Museum</strong> <strong>of</strong> Sichuan Institute <strong>of</strong> Natural Resources,<br />
Chengdu, China<br />
MUMF Manipur University <strong>Museum</strong> <strong>of</strong> Fishes, Canchipur,<br />
India<br />
MUS Muzium Sabah, Kota Kinabalu, Malaysia<br />
MZB <strong>Museum</strong> Zoologicum Bogoriense, Cibinong, Indonesia<br />
MZH Finnish <strong>Museum</strong> <strong>of</strong> Natural History, Zoological <strong>Museum</strong>,<br />
University <strong>of</strong> Helsinki, Helsinki, Finland<br />
MZUF Museo Zoologico "la Specola", Università di Firenze,<br />
Firenze, Italy<br />
MZUP Museo Zoologico, Universita, Pavia, Italy<br />
MZUT Museo Zoologico, Universita, Torino, Italy<br />
NCNTTSI Aquaculture <strong>Research</strong> Institute No. 1 [earlier Dinh<br />
Bang Fish <strong>Research</strong> Station], Bac Ninh, Vietnam<br />
NGI Nanjing Geographical Institute, Academia Sinica, Nanjing,<br />
China [see NIG]<br />
NHMG Natural History <strong>Museum</strong>, Göteborg, Sweden<br />
NHMS Natural History <strong>Museum</strong>, S<strong>of</strong>ia, Bulgaria<br />
NIFI National Inland Fisheries Institute, Bangkok, Thailand<br />
NIG Nanjing Institute <strong>of</strong> Geography and Limnology, Chinese<br />
Academy <strong>of</strong> Sciences, Nanjing, China<br />
NMC Canadian <strong>Museum</strong> <strong>of</strong> Nature, Ottawa, Canada<br />
NMHW <strong>Museum</strong> <strong>of</strong> Natural History, Wroclaw University, Wroclaw,<br />
Poland<br />
NMMB National <strong>Museum</strong> <strong>of</strong> Marine Biology and Aquarium,<br />
Pingtung, Taiwan<br />
NMSL National <strong>Museum</strong> <strong>of</strong> Sri Lanka, Colombo, Sri Lanka<br />
NMW Naturhistorisches <strong>Museum</strong>, Wien, Austria<br />
NPIB Northwest Plateau Institute <strong>of</strong> Biology, Xining, China<br />
NRIBAS National <strong>Research</strong> Institute <strong>of</strong> Biology, Chinese Academy<br />
<strong>of</strong> Sciences, Nanjing, China<br />
NRM Naturhistoriska Riksmuseet, Stockholm, Sweden<br />
NSMT National Science <strong>Museum</strong>, Tokyo, Japan<br />
NTOU National Taiwan Ocean University, Institute <strong>of</strong> Marine<br />
Biology, Keelung, Taiwan<br />
NTUM <strong>Museum</strong> <strong>of</strong> the Department <strong>of</strong> Zoology, National Taiwan<br />
University, Taipei, Taiwan<br />
PMR Rijeka <strong>Museum</strong> <strong>of</strong> Natura History, Rijeka, Croatia<br />
PRFRI Pearl River Fisheries <strong>Research</strong> Institute, Chinese Academy<br />
<strong>of</strong> Fishery Science, Guangzhou, China<br />
PUCMF Pachhunga University College <strong>Museum</strong> <strong>of</strong> Fishes, Aizawl,<br />
India<br />
RMNH Nationaal Natuurhistorisch <strong>Museum</strong> [earlier Rijksmuseum<br />
van Natuurlijke Historie], Leiden, The Netherlands<br />
ROM Royal Ontario <strong>Museum</strong>, Toronto, Canada<br />
SBC Sarawak <strong>Biodiversity</strong> Center, Kuching, Malaysia<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
13<br />
SCNU Department <strong>of</strong> Biology, South China Normal University,<br />
Guangzhou, China<br />
SCUM <strong>Museum</strong> <strong>of</strong> Sichuan University, Chengdu, China<br />
SMF Senckenberg <strong>Museum</strong>, Frankfurt, Germany<br />
SMK Sarawak <strong>Museum</strong>, Kuching, Malaysia<br />
SNM Slovak National <strong>Museum</strong>, Bratislava, Slovakia<br />
SOU Shanghai Ocean University, Shanghai, China<br />
SPNRI Sichuan Provincial Nature <strong>Research</strong> Institute, Chengdu,<br />
China<br />
SSCN <strong>Museum</strong> <strong>of</strong> the Biological Laboratory, Science Society<br />
<strong>of</strong> China, Nanking, China [now NRIBAS]<br />
SU Stanford University; now at CAS (as CAS-SU)<br />
SUBC Department <strong>of</strong> Biology, Seowon University, Cheongju,<br />
Korea<br />
SWFC <strong>Museum</strong> <strong>of</strong> Zoology, Southwest Forestry College, Kunming,<br />
China<br />
TAU Tel Aviv University, Tel Aviv, Israel<br />
TUK Department <strong>of</strong> Zoology, Tribhuvan University, Kathmandu,<br />
Nepal<br />
UBD Universiti Brunei Darussalam, Brunei Darussalam<br />
UF Florida <strong>Museum</strong> <strong>of</strong> Natural History, University <strong>of</strong> Florida,<br />
Gainesville, USA<br />
UMMZ University <strong>of</strong> Michigan <strong>Museum</strong> <strong>of</strong> Zoology, Ann Arbor,<br />
USA<br />
UMSB University Malaysia Sabah, Kota Kinabalu, Malaysia<br />
USNM National <strong>Museum</strong> <strong>of</strong> Natural History, Washington, USA<br />
VUP Vinh University <strong>of</strong> Pedagogy, Vinh, Vietnam<br />
WIRI Watanabe Ichthyological Institute, Tokyo, Japan<br />
YCM Yokosuka City <strong>Museum</strong>, Yokosuka, Japan<br />
YU Department <strong>of</strong> Biology, Yunnan University, Kunming,<br />
China<br />
ZFMK Zoologisches Forschungsinstitut und <strong>Museum</strong> Alexander<br />
Koenig, Bonn, Germany<br />
ZISP Institute <strong>of</strong> Zoology, Academy <strong>of</strong> Sciences, St. Petersburg,<br />
Russia<br />
ZMA Instituut voor Taxonomische Zoölogie, Amsterdam, The<br />
Netherlands; now at RMNH<br />
ZMB Zoologisches <strong>Museum</strong>, Berlin, Germany<br />
ZMCZ Department <strong>of</strong> Biology, Zunyi Medical College, Zhuhai,<br />
China<br />
ZMFIB Zoological <strong>Museum</strong>, Fan Memorial Institute <strong>of</strong> Biology,<br />
Beijing, China; now at ASIZB<br />
ZMH Zoologisches <strong>Museum</strong> und Zoologisches Institut, Hamburg,<br />
Germany<br />
ZMMU Zoological <strong>Museum</strong>, Moscow State University, Moscow,<br />
Russia<br />
ZMNH Zhejiang <strong>Museum</strong> <strong>of</strong> Natural History, Hangzhou,<br />
China<br />
ZMT State <strong>Museum</strong> <strong>of</strong> Georgia, Georgian Academy <strong>of</strong> Sciences,<br />
Tbilisi, Georgia<br />
ZMUC <strong>Museum</strong> <strong>of</strong> Zoology, University, Kobenhavn, Denmark<br />
ZMUU Zoologiska Museet, Uppsala Universitet, Uppsala,<br />
Sweden<br />
ZRC <strong>Raffles</strong> <strong>Museum</strong> <strong>of</strong> <strong>Biodiversity</strong> <strong>Research</strong>, National<br />
University <strong>of</strong> Singapore, Singapore<br />
ZSI Zoological Survey <strong>of</strong> India, Calcutta, India<br />
ZSI/APFS Arunachal Pradesh Regional Station, ZSI, Itanagar,<br />
India<br />
ZSI/ERS Eastern Regional Station, ZSI, Shillong, India<br />
ZSI/NRS Northern Regional Station, ZSI, Dehra Dun, India<br />
ZSI/SRS Southern Regional Station, ZSI, Madras, India<br />
ZSI/WGRS Western Ghat Regional Station, ZSI, Calicut, India<br />
ZSI/WRC Western Regional Centre, ZSI, Pune, India<br />
ZSM Zoologische Staatssammlung, München, Germany
1 Gyrinocheilidae Gill, 1905<br />
Gyrinocheilidae Gill, 1905: 196 (type genus: Gyrinocheilus<br />
Vaillant, 1902: 107)<br />
1.1 Gyrinocheilus Vaillant, 1902<br />
Gyrinocheilus Vaillant, 1902: 107 (type species: Gyrinocheilus<br />
pustulosus Vaillant, 1902: 111, by monotypy).<br />
Gender masculine.<br />
Gyrinocheilops Fowler, 1937: 160 (type species: Gyrinocheilops<br />
pennocki Fowler, 1937: 161, by original<br />
designation). Gender masculine.<br />
Taxonomic notes. Revision by Roberts & Kottelat (1993).<br />
1.1.1 Gyrinocheilus aymonieri (Tirant, 1884)<br />
Psilorhynchus Aymonieri Tirant, 1884: 167, figs. 1–2 (type<br />
locality: Cambodia: tributaries <strong>of</strong> Prek-Tenot in the hills<br />
<strong>of</strong> Samrong Tong, 75 km from Phnom Penh; holotype:<br />
MGHNL 42000056, Kottelat, 1987a: 17, Roberts &<br />
Kottelat, 1993: 378, fig. 3b–c; noun in genitive, indeclinable)<br />
Gyrinocheilus kaznakovi Berg, 1906a: 306, 365 (type locality:<br />
Cambodia: Pailin, between Battambang and Schan-<br />
2 Botiidae Berg, 1940<br />
Botiidae Berg, 1940: 270 (type genus: Botia Gray, 1831b: 8)<br />
Leptobotiini Nalbant, 2002: 315 (type genus: Leptobotia<br />
Bleeker, 1870: 256)<br />
2.1. Ambastaia Kottelat, 2012<br />
Ambastaia Kottelat, 2012: 137 [appendix <strong>of</strong> present work]<br />
(type species: Botia nigrolineata Kottelat & Chu, 1987:<br />
395, by original designation). Gender feminine.<br />
2.1.1 Ambastaia nigrolineata (Kottelat & Chu, 1987)<br />
Botia nigrolineata Kottelat & Chu, 1987: 395, fig. 3 (type<br />
locality: China: Yunnan: Xishuangbanna: Menghan, about<br />
21°50'N 100°23'E; holotype: KIZ 735198; misspelt<br />
niqrolineata p. 395, a type setter error, Code art. 32.5.1;<br />
adjective, -us, -a, -um)<br />
2.1.2 Ambastaia sidthimunki (Klausewitz, 1959)<br />
Botia sidthimunki Klausewitz, 1959: 51, figs. 1–3 (type locality:<br />
brooks <strong>of</strong> northern Thailand ["Loom district, Yom<br />
River"; Eschmeyer & Fricke, 2010, no source stated]<br />
[Phetchaburi Province: probably Lom Sak district,<br />
16°46'46"N 101°14'32"E]; holotype: SMF 4505; noun<br />
in genitive, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Family GYRINOCHEILIDAE<br />
Family BOTIIDAE<br />
14<br />
tabun [Chantaburi, in Thailand]; syntypes: ZISP 11254<br />
[2], Roberts & Kottelat, 1993: 381, fig. 3a; noun in genitive,<br />
indeclinable)<br />
Gyrinocheilus monchadskii Krasyukova & Gusev, 1987: 67,<br />
fig. 1, pl. 2 (type locality: China: Yunnan: Mekong River<br />
near Tchili; holotype: ZISP 48103, Eschmeyer & Fricke,<br />
2010; noun in genitive, indeclinable)<br />
1.1.2 Gyrinocheilus pennocki (Fowler, 1937)<br />
Gyrinocheilops pennocki Fowler, 1937: 161, figs. 98–99<br />
(type locality: Thailand: Kemarat; holotype: ANSP<br />
68102, Böhlke, 1984: 112; noun in genitive, indeclinable)<br />
1.1.3 Gyrinocheilus pustulosus Vaillant, 1902<br />
Gyrinocheilus pustulosus Vaillant, 1902: 111, figs. 30–32,<br />
pls. 1–2 (type locality: Indonesia: Borneo: Kalimantan<br />
Barat: Kapuas River basin: mouth <strong>of</strong> Raoen [Raun,<br />
0°39'N 113°10'E], upper Sibau; syntypes: RMNH 7796<br />
[1], 7797 [2], BMNH 1921.7.28.1 [1], Roberts & Kottelat,<br />
1993: 378, Eschmeyer & Fricke, 2010; adjective,<br />
-us, -a, -um)<br />
2.2 Botia Gray, 1381<br />
Botia Gray, 1831b: 8 (type species: Botia almorhae Gray,<br />
1831b: 8, by monotypy). Gender feminine.<br />
Hymenphysa M'Clelland, 1839: 443 (type species: Cobitis<br />
dario Hamilton, 1822: 354, by subsequent designation<br />
by Jordan, 1919: 195). Gender feminine.<br />
Diacantha Swainson, 1839: 190, 310 (subgenus <strong>of</strong> Canthophrys<br />
Swainson, 1838: 364; type species: Canthophrys<br />
zebra Swainson, 1839: 310, by subsequent designation<br />
by Swain, 1883: 282; also spelt Diacanthus p. 190, first<br />
reviser [Eschmeyer, 1990: 123] retained Diacantha as<br />
correct original spelling). Gender feminine.<br />
Hymenophysa Bleeker, 1858: 303 (incorrect subsequent<br />
spelling <strong>of</strong> Hymenphysa M'Clelland, 1839: 443, see Kottelat,<br />
2004a: 12)<br />
Nomenclatural notes. Bleeker (1858: 303) explicitly listed<br />
Hymenophysa as a name created by M'Clelland, without<br />
explaining the spelling change. Therefore it is not an emendation<br />
but an incorrect subsequent spelling (Code art. 33.1,<br />
33.3).<br />
Nalbant (2002: 317) used the name Hymenophysa for<br />
the species here placed in Syncrossus. Kottelat (2004a: 12)<br />
showed that the name Hymenophysa could not be used for<br />
these species and that, anyway, Hymenophysa being an in-
1.1.1 Gyrinocheilus aymonieri, CMK 22815, 80.5 mm SL; Laos:<br />
Mekong drainage: Nam Theun.<br />
1.1.1 Gyrinocheilus aymonieri, CMK 22799, 128 mm SL.<br />
correct spelling it is not available. Nalbant (2004: 270)<br />
claimed that the name in prevailing usage (Hymenophysa)<br />
should be maintained under Code art. 33.3.1. Nalbant interpreted<br />
Hymenphysa and Hymenophysa as two different<br />
names and therefore art. 33.3.1 is irrelevant since it applies<br />
only in cases <strong>of</strong> alternative spellings <strong>of</strong> a name, and not in<br />
case <strong>of</strong> alternative names for a taxon.<br />
Even though Hymenophysa is not a distinct name but an<br />
incorrect subsequent spelling <strong>of</strong> Hymenphysa, Code art.<br />
33.3.1 could not be used the way interpreted by Nalbant.<br />
This article states that an incorrect subsequent spelling has<br />
to be preserved when it is "in prevailing usage and attributed<br />
to the publication <strong>of</strong> the original spelling". Nalbant wished<br />
to retain Hymenophysa as spelt by Bleeker [supposed prevailing<br />
usage], and to use it for the genus called here Syncrossus,<br />
without attributing it to M'Clelland [original spelling]<br />
but to Bleeker (1858). Anyway, even if the spelling<br />
Hymenophysa had been attributed to M'Clelland, the type<br />
species remains Cobitis dario and the result remains that<br />
Hymenophysa is a junior synonym <strong>of</strong> Botia and cannot be<br />
used instead <strong>of</strong> Syncrossus. Further, Hymenophysa having<br />
almost never been used as the valid name <strong>of</strong> a genus, it is<br />
difficult to qualify it as 'prevailing usage'.<br />
2.2.1 Botia almorhae Gray, 1831<br />
Botia Almorhae Gray, 1831b: 8 (type locality: India: Uttarakhand:<br />
Almorah; holotype: BMNH 2002.9.18.1, Kullander<br />
et al., 1999: 137, Grant, 2007a: fig. 30; noun in genitive,<br />
indeclinable)<br />
Botia grandis Gray, 1832: vol. 1, pl. 94 fig. 3 (type locality:<br />
India: Uttarakhand: Almorah [29°37'12"N 79°40'12"E];<br />
holotype: BMNH 2002.9.18.1; objective junior synonym<br />
<strong>of</strong> Botia almorhae Gray, 1831b: 8; adjective, -is, -is, -e)<br />
Schistura maculata M'Clelland, 1839: 307 (unnecessary replacement<br />
name for Botia grandis Gray, 1832: vol. 1, pl.<br />
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15<br />
2.1.1. Ambastaia nigrolineata; top: CMK16043, 32.8 mm SL;<br />
Thailand: Chao Phraya drainage: Nan; bottom: CMK 21861, 65.1<br />
mm SL; Laos: Mekong drainage: Xekong.<br />
94 fig. 3; adjective, -us, -a, -um)<br />
Botia Blythi Bleeker, 1863a: 39 (type locality: India: Uttarakhand:<br />
Almorah [29°37'12"N 79°40'12"E]; syntypes:<br />
BMNH 2002.9.18.1 [1], ZSI ?; available by indication<br />
to Schistura grandis <strong>of</strong> M'Clelland, 1839: 307, 444 and<br />
Syncrossus grandis <strong>of</strong> Blyth, 1860: 165; actually, C. grandis<br />
<strong>of</strong> M'Clelland is a new combination <strong>of</strong> Botia grandis<br />
Gray, 1832; M'Clelland's description is exclusively based<br />
on drawing in Gray, 1832; Blyth's record is based on a<br />
specimen from Almorah; noun in genitive, indeclinable)<br />
2.2.2 Botia birdi Chaudhuri, 1910<br />
Botia birdi Chaudhuri, 1910a: 339 (type locality: India:<br />
Ambala: Rupar [Rupnagar, 30°57'59"N 76°31'59"E],<br />
where Sirhind Canal diverges from Sutlej River, Indus<br />
River drainage; syntypes: ZSI F 3578–3580/1 [3], Menon<br />
& Yazdani, 1968: 120; noun in genitive, indeclinable)<br />
Botia javedi Mirza & Syed, 1995: 379, fig. (type locality:<br />
Pakistan: Michni, 34°11'N 71°28'E, about 24 km north<br />
<strong>of</strong> Peshawar, Kabul River system, Indus River drainage;<br />
holotype: GCMNH 17F; noun in genitive, indeclinable)<br />
2.2.3 Botia dario (Hamilton, 1822)<br />
Cobitis dario Hamilton, 1822: 354, 394, pl. 29 fig. 95 (type<br />
locality: India: "northern rivers <strong>of</strong> Bengal" [Bangladesh:<br />
Dumdumma, Dinajpur District, 25°37'N 88°38'E; Hora,<br />
1935a: 49]; types: NT; noun in apposition, indeclinable)<br />
Cobitis geto Hamilton, 1822: 355, 394, pl. 11 fig. 96 (type<br />
locality: India: "north-eastern parts <strong>of</strong> Bengal" [Goalpara;<br />
Hora, 1935a: 49]; simultaneous subjective synonym <strong>of</strong><br />
Cobitis dario Hamilton, 1822: 354; first reviser [Günther,<br />
1868: 366] gave precedence to C. dario; types: NT; noun<br />
in apposition, indeclinable)<br />
Canthophrys flavicauda Swainson, 1839: 310 (available by<br />
indication to Hamilton, 1822: pl. 29 fig. 95 [which is<br />
Cobitis dario]; type locality: India: "northern rivers <strong>of</strong><br />
Bengal" [Bangladesh: Dumdumma, Dinajpur District,<br />
25°37'N 88°38'E; Hora, 1935a: 49]; types: NT; compound<br />
noun, indeclinable)<br />
Canthophrys zebra Swainson, 1839: 310 (available by indication<br />
to Hamilton, 1822: pl. 11 fig. 96 [which is Cobitis<br />
geto]; type locality: India: "north-eastern parts <strong>of</strong> Ben-
gal" [Goalpara; Hora, 1935a: 49]; types: NT; noun in<br />
apposition, indeclinable)<br />
Botia macrolineata Teugels, De Vos & Snoeks, 1986: 188,<br />
fig. 2 (type locality: India: reportedly from about 100<br />
km from Bombay; holotype: IRSNB 701; adjective, -us,<br />
-a, -um)<br />
2.2.4 Botia histrionica Blyth, 1860<br />
Botia histrionica Blyth, 1860: 166 (type locality: Burma:<br />
Tenasserim; holotype: ZSI F 2634/1, Menon & Yazdani,<br />
1968: 120; adjective, -us, -a, -um)<br />
2.2.5 Botia kubotai Kottelat, 2004<br />
Botia kubotai Kottelat, 2004a: 2, fig. 1 (type locality: Myanmar:<br />
Kayin State [Karen]: stream "Chon Son" between<br />
Kyondaw and Phadaw, about 20 km northwest <strong>of</strong><br />
Payathouzu [Payathonzu] (at border with Thailand), about<br />
15°25'N 98°15'E; holotype: MHNG 2644.24; noun in<br />
genitive, indeclinable)<br />
2.2.6 Botia lohachata Chaudhuri, 1912<br />
Botia lohachata Chaudhuri, 1912: 441, pl. 40 fig. 2 (type<br />
locality: India: Bihar: Gandak River in Saran; holotype:<br />
ZSI F 8068/1, Menon & Yazdani, 1968: 120; adjective,<br />
-us, -a, -um)<br />
2.2.7 Botia rostrata Günther, 1868<br />
? Botia nebulosa Blyth, 1860: 165 (type locality: India:<br />
Darjeeling; holotype: ZSI F 2637/1, Menon & Yazdani,<br />
1968: 120; adjective, -us, -a, -um)<br />
Botia rostrata Günther, 1868: 367, fig. (type locality: India:<br />
Ganges and Assam; syntypes: BMNH 1855.12.26.694<br />
[1], 1860.3.19.114 [1], Grant, 2007a: figs. 11–12; adjective,<br />
-us, -a, -um)<br />
Botia dayi Hora, 1932b: 571 (type locality: India: Mahanaddi<br />
River [Mahananda, flowing through Siliguri,<br />
26°41'52"N 88°23'34"E] at base <strong>of</strong> Darjiling [Darjeeling]<br />
Himalayas; holotype: ZSI F 11299/1, Menon &<br />
Yazdani, 1968: 120; noun in genitive, indeclinable)<br />
Nomenclatural notes. Botia nebulosa has usually been listed<br />
as a synonym <strong>of</strong> Acanthocobitis botia but Blyth (1860:<br />
165) unambiguously described it as a species <strong>of</strong> Botia, with<br />
the branched suborbital spine. Grant (2007a: unnumb. p. 16)<br />
conjectured that it might be a senior synonym <strong>of</strong> B. rostrata.<br />
Assuming they are synonyms, B. nebulosa cannot be declared<br />
nomen oblitum under Code art. 23.9.1 because it has been<br />
used (though rarely) as the valid name for a species after<br />
1899 (e.g. Negi, 1994: 156). As long as the identity <strong>of</strong> B.<br />
nebulosa is not established by examination <strong>of</strong> the holotype,<br />
B. rostrata should be retained as the valid name <strong>of</strong> the species.<br />
2.2.8 Botia striata Narayan Rao, 1920<br />
Botia striata Narayan Rao, 1920: 60, pl. 2 fig. 4 (type locality:<br />
India: Maharasthra: Thunga River in Shimoga town;<br />
holotype: BMNH 1919.11.19.8–12 [1 <strong>of</strong> 5], Eschmeyer<br />
& Fricke, 2010 [specimens listed as syntypes by Menon<br />
& Yazdani, 1968: 121 are not syntypes]; adjective, -us,<br />
-a, -um)<br />
Botia striata var. kolhapurensis Kalawar & Kelkar, 1958:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
16<br />
677 (type locality: India: Maharashtra: Kolhapur; syntypes:<br />
[LU] 6750; adjective, -is, -is, -e)<br />
2.2.9 Botia udomritthiruji Ng, 2007<br />
Botia udomritthiruji Ng, 2007: 42, figs. 1–3 (type locality:<br />
Myanmar: Taninthayi Division: Tenasserim River drainage<br />
in the vicinity <strong>of</strong> Same, 13°36'N 99°02'E; holotype:<br />
UMMZ 248184; noun in genitive, indeclinable)<br />
2.3 Chromobotia Kottelat, 2004<br />
Chromobotia Kottelat, 2004a: 13 (type species: Cobitis<br />
macracanthus Bleeker, 1852: 603, by original designation).<br />
Gender feminine.<br />
2.3.1 Chromobotia macracanthus (Bleeker, 1852)<br />
Cobitis macracanthus Bleeker, 1852: 603 (type locality:<br />
Indonesia: Sumatra: Kwanten River [Kuantan]; lectotype:<br />
RMNNH 7058, designated by Alfred, 1961: 34; compound<br />
noun, indeclinable)<br />
2.4 Leptobotia Bleeker, 1870<br />
Leptobotia Bleeker, 1870: 256 (type species: Botia elongata<br />
Bleeker, 1870: 254, by monotypy). Gender feminine.<br />
Species inquirendae et incertae sedis<br />
Nemacheilus cheni Herre & Lin, 1936: 20, fig. 6 (type locality:<br />
upper Tsien Tang; holotype: LU [paratypes [2]: CAS-<br />
SU 35257 [1], Böhlke, 1953: 40]; noun in genitive, indeclinable).<br />
Apparently a Leptobotia.<br />
Botia variegata Günther, 1889a: 228 (type locality: China:<br />
Sichuan: Yangtsze-Kiang, Ichang [Hubei: Ychang,<br />
30°43'N 111°17'N]; syntypes: BMNH 1889.6.25.28–29<br />
[2]; secondary junior homonym <strong>of</strong> Cobitis variegata<br />
Dabry de Thiersant, 1872: 191, when treated as valid in<br />
Botia; also in Günther, 1892: 249; adjective, -us, -a,<br />
-um)<br />
2.4.1 Leptobotia elongata (Bleeker, 1870)<br />
Botia elongata Bleeker, 1870: 254, pl. (type locality: China:<br />
Yang-Tse-Kiang [Yangtze]; holotype: MNHN 5930, Bertin<br />
& Estève, 1948: 94; adjective, -us, -a, -um)<br />
Cobitis variegata Dabry de Thiersant, 1872: 191, pl. 49 fig. 5<br />
(type locality: China: Yetchuen; holotype: MNHN ?; adjective,<br />
-us, -a, -um)<br />
Botia citrauratea Nichols, 1925a: 5 (type locality: China:<br />
Hunan: Tungting Lake; holotype: AMNH 8402; adjective,<br />
-us, -a, -um)<br />
2.4.2 Leptobotia flavolineata Wang, 1981<br />
Leptobotia flavolineata Wang, 1981: 1, pl. (type locality:<br />
China: Beijing: Shidu Fangshan County: Juma River;<br />
holotype: BMAM 772196; adjective, -us, -a, -um)<br />
2.4.3 Leptobotia guilinensis Chen, 1980<br />
Leptobotia guilinensis Chen, 1980: 15, fig. 5 (type locality:<br />
China: Guangxi: Guilin; syntypes: IHB 587705, 712, 809,<br />
827, 7541652, 1654–1656, 1705–1707, 1709, 1710,<br />
1712, 2223, 2224 [16]; adjective, -is, -is, -e)
2.2.5 Botia kubotai, MHNG 2664.24, 84.9 mm SL, holotype;<br />
Myanmar: Ataran drainage.<br />
2.3.1 Chromobotia macracanthus, CMK 10304, 54.7 mm SL;<br />
Indonesia: Borneo: Kapuas drainage.<br />
2.4.4 Leptobotia hengyangensis Huang & Zhang, 1986<br />
Leptobotia hengyangensis Huang & Zhang, 1986: 99 (type<br />
locality: China: Hunan: Changjiang River [Xiang-jiang]:<br />
Heng-Yang; syntypes: IHB 80037, 80013 [2]; adjective,<br />
-is, -is, -e)<br />
2.4.5 Leptobotia microphthalma Fu & Ye, 1983<br />
Leptobotia microphthalma Fu & Ye, 1983: 121, fig. 1 (type<br />
locality: China: Sichuan: Leshan County: Min River<br />
drainage; syntypes: Dept. Zool. Sichuan Agric. Coll. 6-<br />
1040 [1], IHB 6-1118 [1]; compound adjective, -us, -a,<br />
-um)<br />
2.4.6 Leptobotia orientalis Xu, Fang & Wang, 1981<br />
Leptobotia orientalis Xu, Fang & Wang, 1981: 379, fig. 1<br />
(type locality: China: Shanxi: Danfeng County: Wuguan<br />
River; holotype: IZSX 80VI0074; adjective, -is, -is, -e)<br />
2.4.7 Leptobotia pellegrini Fang, 1936<br />
Leptobotia pellegrini Fang, 1936: 29 (type locality: China:<br />
Sichuan; holotype: NRIBAS 1779; noun in genitive, indeclinable)<br />
Taxonomic notes. The species figured as L. elongata by<br />
Kottelat (2001a: fig. 102) and Nguyen (2005: 206, fig. 103)<br />
is apparently L. pellegrini.<br />
2.4.8 Leptobotia posterodorsalis Lan & Chen, in Chen &<br />
Lan, 1992<br />
Leptobotia posterodorsalis Lan & Chen, in Chen & Lan,<br />
1992: 106, fig. 3 (type locality: China: Guangxi: Huanjiang<br />
County; holotype: IHB 87067044; compound noun,<br />
indeclinable)<br />
2.4.9 Leptobotia punctata Li, Li & Chen, 2008<br />
Leptobotia punctatus Li, Li & Chen, 2008: 630, fig. 1 (type<br />
locality: China: Guangxi. Qianjiang River in Guiping City,<br />
23°24'N 110°24'E; holotype: ASIZB 07 X PRFRI 001;<br />
adjective, -us, -a, -um)<br />
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17<br />
2.4.7 Leptobotia pellegrini, CMK 21967, 71.6 mm SL; Vietnam:<br />
Pearl River drainage.<br />
2.5.4 Parabotia curtus, RMNH 28128, 79.7 mm SL; Japan: Osaka.<br />
2.4.10 Leptobotia rubrilabris Dabry de Thiersant, 1872<br />
Leptobotia rubrilabris Dabry de Thiersant, 1872: 191, pl. 49<br />
fig. 8 (type locality: China: Yang-tsee-kiang [Yangtze];<br />
holotype: MNHN 5085, Bertin & Estève, 1948: 84; compound<br />
noun, indeclinable)<br />
Botia pratti Günther, 1892: 250, pl. 4 fig. A (type locality:<br />
China: Sichuan: Kia-tiang-fu, foot <strong>of</strong> Omie-shan [Omei<br />
Shan]; syntypes: BMNH 1891.6.13:35–37 [3], Fang,<br />
1936: 43; noun in genitive, indeclinable)<br />
Botia fangi Tchang, 1930: 51, fig. 5 (type locality: China:<br />
Sichuan; holotype: LU or possibly MNHN [paratype:<br />
MNHN 1934.174, Bertin & Estève, 1948: 94]; noun in<br />
genitive, indeclinable)<br />
2.4.11 Leptobotia taeniops (Sauvage, 1878)<br />
Cobitis Cha-ny Dabry de Thiersant, 1872: 191, pl. 49 fig. 9<br />
(name not binominal, not available; locality: China: "rivière<br />
de Kouei-tcheou-fou (Sse-tchuen)" [Sichuan: river<br />
<strong>of</strong> Kweichow-fu = river <strong>of</strong> Guizhou-fu [Guiyang]; this<br />
apparently means "in Sichuan, in river that flows from<br />
Guiyang", i.e. Nanning He, which becomes Wu Jiang])<br />
Parabotia taeniops Sauvage, 1878a: 90 (type locality: China:<br />
Yang-tse-kiang [Yangtze]; holotype: MNHN 5086,<br />
Bertin & Estève, 1948: 84; compound noun, indeclinable)<br />
Leptobotia purpurea Nichols, 1925a: 4, fig. 3 (type locality:<br />
China: Hunan: Huping, Tungting Lake; holotype: AMNH<br />
8401; adjective, -us, -a, -um)<br />
2.4.12 Leptobotia tchangi Fang, 1936<br />
Leptobotia tchangi Fang, 1936: 40 (type locality: China:<br />
Zhejiang: Hsia-chiao-kou near West Tien-mu-shan; holotype:<br />
"West Lake <strong>Museum</strong>" [? paratype: MNHN<br />
1934.175, Bertin & Estève, 1948: 94]; noun in genitive,<br />
indeclinable)<br />
2.4.13 Leptobotia tientainensis (Wu, 1930)<br />
Botia tientainensis Wu, 1930: 258, fig. 3 (type locality: China:<br />
Tche-kiang: brook in Tien-Tai hill; holotype: MNHN<br />
1934.176, Bertin & Estève, 1948: 84; adjective, -is, -is, -e)<br />
Botia compressicauda Nichols, 1931c: 2 (type locality: China:<br />
northwestern Fukien [Fujian]: Chungan Hsien<br />
[Chong'an County]; holotype: AMNH 9682; compound<br />
noun, indeclinable)
Leptobotia tientaiensis hansuiensis Fang & Hsu, 1980: 265,<br />
fig. 1 (type locality: China: Shaanxi: Langao; holotype:<br />
IZSX 790236; adjective, -is, -is, -e)<br />
2.5 Parabotia Dabry de Thiersant, 1872<br />
Parabotia Dabry de Thiersant, 1872: 191 (type species:<br />
Parabotia fasciatus Dabry de Thiersant, 1872: 191, by<br />
subsequent designation by Fang, 1936: 4). Gender masculine<br />
[Code art. 30.2.3].<br />
Nomenclatural notes. Dabry de Thiersant (1872: 191) did<br />
not indicate the gender <strong>of</strong> the genus Parabotia when he created<br />
it. As Parabotia is not and does not end with a Latin,<br />
Greek or Western European word (Code art. 30.2.1), its gender<br />
is determined by that <strong>of</strong> adjectivial species-group names<br />
originally included. Two nominal species were originally<br />
included, P. fasciatus and P. rubrilabris. Rubrilabris is a noun<br />
in apposition and therefore uninformative. Fasciatus is an<br />
adjective with a masculine declension, therefore the gender<br />
<strong>of</strong> Parabotia is masculine. This is unfortunate, since all other<br />
genus names ending in -botia are feminine, but it cannot<br />
be changed.<br />
2.5.1 Parabotia banarescui (Nalbant, 1965)<br />
Leptobotia banarescui Nalbant, 1965: 2, fig. 4 (type locality:<br />
China: Hupeh: Yangtze at Wutchang [Wuhan]; holotype:<br />
ISBB 1332; noun in genitive, indeclinable)<br />
2.5.2 Parabotia bimaculatus Chen, 1980<br />
Parabotia bimaculata Chen, 1980: 11, fig. 3 (type locality:<br />
China: Sichuan: Lu-Xian; syntypes: IHB 58426, 58530,<br />
58532–536, 585243, 585244, 584284, 583023, 583029,<br />
583053, 583057 [13]; adjective, -us, -a, -um)<br />
2.5.3 Parabotia brevirostris Zhu & Zhu, 2012<br />
Parabotia brevirostris Zhu & Zhu, 2012: 448, fig. 1 (type<br />
locality: China: Guangxi: Du'an County: Hongshui river,<br />
Pearl River drainage; holotype: FLFS 2008005; compound<br />
noun, indeclinable)<br />
2.5.4 Parabotia curtus (Temminck & Schlegel, 1846)<br />
Cobitis curta Temminck & Schlegel, 1846: 223, pl. 103 fig. 4<br />
(type locality: Japan [surroundings <strong>of</strong> Nagasaki]; holotype:<br />
RMNH 2708, Boeseman, 1947: 167; adjective,<br />
-us, -a, -um)<br />
2.5.5 Parabotia dubius Kottelat, 2001<br />
Botia elongata Mai, 1978: 240, fig. 110 (type locality: northern<br />
Vietnam: Chay River, Thac Ba reservoir; holotype:<br />
DVZUT; junior primary homonym <strong>of</strong> Botia elongata<br />
Bleeker, 1870: 254; adjective, -us, -a, -um)<br />
Parabotia dubia Kottelat, 2001a: 50 (replacement name for<br />
Botia elongata Mai, 1978: 240; adjective, -us, -a, -um)<br />
? Parabotia kimluani Nguyen, 2005: 553, fig. 5 (type locality:<br />
Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; etymology unknown,<br />
either noun in genitive or in apposition, both indeclinable)<br />
? Parabotia vancuongi Nguyen, 2005: 555, fig. 6 (type locality:<br />
Vietnam: Tuyen Quang Province: Na Hang district:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
18<br />
Gam River; holotype: NCNTTSI; subjective simultaneous<br />
synonym <strong>of</strong> Parabotia kimluani Nguyen, 2005:<br />
553; as first reviser, I give precedence to Parabotia kimluani;<br />
also spelt vacuongi p. 686, as first reviser I select<br />
vancuongi as the correct original spelling; etymology<br />
unknown, either noun in genitive or in apposition, both<br />
indeclinable)<br />
2.5.6 Parabotia fasciatus Dabry de Thiersant, 1872<br />
Parabotia fasciatus Dabry de Thiersant, 1872: 191, pl. 49<br />
fig. 7 (type locality: China: Yang-tsee-kiang [Yangtze];<br />
holotype: MNHN 5088, Bertin & Estève, 1948: 92; adjective,<br />
-us, -a, -um)<br />
Nemachilus xanthi Günther, 1888: 434 (type locality: China:<br />
Sichuan: Yangtsze-Kiang at Ichang [Hubei: Ychang,<br />
30°43'N 111°17'N]; holotype: BMNH 1888.5.15.43;<br />
noun in genitive or in apposition, indeclinable)<br />
Botia multifasciata Regan, 1905c: 389, pl. 5 fig. 3 (type locality:<br />
China [Tung-Kung, east <strong>of</strong> Guangdong, on East River;<br />
Mahnert, 1976: 473]; holotype: MHNG 677.98, Mahnert,<br />
1976: 473; adjective, -us, -a, -um)<br />
Leptobotia intermedia Mori, 1929: 384 (type locality: China:<br />
Tsi-nan; holotype: LU; adjective, -us, -a, -um)<br />
Leptobotia hopeiensis Shaw & Tchang, 1931: 70, fig. 3 (type<br />
locality: China: Shao-ho, about 60 li [then about 25 km]<br />
north <strong>of</strong> Beijing; holotype: ZMFIB 5378; adjective, -is,<br />
-is, -e)<br />
Leptobotia kudoii Mori, 1933: 13, fig. (type locality: China:<br />
Manchuria [now in Jilin Province]: Sungari River near<br />
Kirin [Jilin City]; holotype: LU; noun in genitive, indeclinable)<br />
Botia kwangsiensis Fang, 1936: 13 (type locality: China:<br />
Guangxi: Ling-yüng-shien / Nan-gning / Pai-Sê; syntypes:<br />
NRIBAS 780, 808, 821–824, 907–909, 1085, 1188–1189<br />
[13 ?], MNHN 1940.142, AMNH 12971 [ex NRIBAS<br />
1084], Bertin & Estève, 1948: 92, Eschmeyer & Fricke,<br />
2010; adjective, -is, -is, -e)<br />
Botia wui Chang, 1944: 48, fig. 2 (type locality: China: Sichuan:<br />
Loshan in Min River drainage; syntypes [2]: [LU]<br />
2134 [1], 2135 [1]; noun in genitive, indeclinable)<br />
2.5.7 Parabotia heterocheilus Zhu & Chen, in Ni & Wu,<br />
2006<br />
Parabotia heterocheila Zhu & Chen, in Ni & Wu, 2006:<br />
383, fig. 180 (type locality: China: Jiangsu: Nanjing<br />
Municipality: Xi Jiang stream near its mouth, a small tributary<br />
<strong>of</strong> Chiang Jiang River, at Jiangpu; holotype: FFRIJ<br />
200211001; compound adjective, -us, -a, -um)<br />
2.5.8 Parabotia kiangsiensis Liu & Guo, 1986<br />
Parabotia kiangsiensis Liu & Guo, 1986: 69, fig. 1 (type<br />
locality: China: Jiangxi: Sin River; syntypes: Department<br />
<strong>of</strong> Biology, Jiangxi University 815181, 815326, 840512,<br />
840513 [4]; syntypes because listed as 2 holotypes and 2<br />
paratypes; also spelt kiangensis on p. 70, an obvious inadvertent<br />
error, thus incorrect original spelling [Code art.<br />
32.5.1]; adjective, -is, -is, -e)<br />
2.5.9 Parabotia lijiangensis Chen, 1980<br />
Parabotia lijiangensis Chen, 1980: 11, fig. 2 (type locality:
China: Guangxi: Guilin: syntypes: IHB 587746, 780, 782,<br />
802, 806, 75IV2263, 2264, 2258 [8]; adjective, -is, -is, -e)<br />
2.5.10 Parabotia maculosus (Wu, 1939)<br />
Botia maculosa Wu, 1939: 121, pl. 3 fig. 6 (type locality:<br />
China: Guangxi: Li-Kiang in Yongso; syntypes: [LU]<br />
271–272 [2]; adjective, -us, -a, -um)<br />
2.5.11 Parabotia mantschuricus (Berg, 1907)<br />
Leptobotia mantschurica Berg, 1907b: 420 (type locality:<br />
China: Heilongjiang: Mutang-kiang [Mudan Jiang River]<br />
near E-ho, a tributary <strong>of</strong> Sungari River [Songhua<br />
Jiang], Amur drainage; syntypes: ZISP 14085 [2]; adjective,<br />
-us, -a, -um)<br />
2.5.12 Parabotia parvus Chen, 1980<br />
Parabotia parva Chen, 1980: 12, fig. 4 (type locality: China:<br />
Guangxi: Bopai [? Bobai, 22°13'00"N 109°55'00"E];<br />
syntypes: IHB 75V3614, 3616–3623, 3625–3632, 3634–<br />
3637 [21]; adjective, -us, -a, -um)<br />
2.6 Sinibotia Fang, 1936<br />
Sinibotia Fang, 1936: 19 (subgenus <strong>of</strong> Botia Gray, 1831b: 8;<br />
type species: Botia superciliaris Günther, 1892: 250, by<br />
original designation). Gender feminine.<br />
2.6.1 Sinibotia longiventralis (Yang & Chen, 1992)<br />
Botia longiventralis Yang & Chen, 1992: 344, fig. 3 (type<br />
locality: China: Yunnan: Weixi County: small stream and<br />
Lancangjiang [Mekong] near Baijixun [27°21'N<br />
99°06'E]; holotype: KIZ 748660; compound noun, indeclinable)<br />
2.6.2 Sinibotia pulchra (Wu, 1939)<br />
Botia pulchra Wu, 1939: 124, pl. 2 fig. 4 (type locality:<br />
China: Guangxi: Li-Kiang at Yangso; syntypes: LU, 405–<br />
406 [2]; adjective, -er, -ra, -rum)<br />
Botia gigantea Mai, 1978: 239, fig. 109 (type locality: northern<br />
Vietnam; types: DVZUT; adjective, -us, -a, -um)<br />
2.6.3 Sinibotia reevesae (Chang, 1944)<br />
Botia reevesae Chang, 1944: 49 (type locality: China: Sichuan:<br />
Yaotan at Wachang, Luhsien; syntypes: LU, 3028,<br />
3031, 3051, 3065 [4]; noun in genitive, indeclinable)<br />
2.6.4 Sinibotia robusta (Wu, 1939)<br />
Botia robusta Wu, 1939: 122, pl. 2 fig. 5 (type locality: China:<br />
Guangxi: Li-Kiang at Yangso; syntypes: LU, 278–<br />
285 [8]; adjective, -us, -a, -um)<br />
Botia hexafurca Mai, 1978: 238, fig. 108 (type locality: northern<br />
Vietnam: Tay Giang basin; types: DVZUT; compound<br />
noun, indeclinable)<br />
2.6.5 Sinibotia superciliaris (Günther, 1892)<br />
Botia superciliaris Günther, 1892: 250, pl. 4 fig. B (type<br />
locality: China: Sichuan: Kia-tiang-fu, foot <strong>of</strong> Omie-shan<br />
[Omei Shan]; syntypes: BMNH 1891.6.13.30–34 [5],<br />
Eschmeyer & Fricke, 2010; compound noun, indeclinable)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
19<br />
2.6.6 Sinibotia zebra (Wu, 1939)<br />
Botia zebra Wu, 1939: 126, pl. 3 fig. 9 (type locality: China:<br />
Guangxi: Li-Kiang at Yangso; holotype: LU, 407; noun<br />
in apposition, indeclinable)<br />
Taxonomic notes. Generic placement follows Tang et al.<br />
(2008).<br />
2.7 Syncrossus Blyth, 1860<br />
Syncrossus Blyth, 1860: 166 (type species: Syncrossus berdmorei<br />
Blyth, 1860: 166, by subsequent designation by<br />
Jordan & Fowler, 1903: 772 ["Schistura grandis apud<br />
McClelland" qualifies as a deliberate application <strong>of</strong> a<br />
previous (mis)identification, Code art. 67.2.1, and therefore<br />
two species are included; Cobitis (Schistura) grandis<br />
in M'Clelland, 1839: 307, 444 in fact is a new combination<br />
<strong>of</strong> Botia grandis Gray, 1832: vol. 1, pl. 94 fig. 3;<br />
"apud" means "in an author"). Gender masculine.<br />
Nomenclatural notes. See under Botia Gray, 1831 for discussion<br />
<strong>of</strong> Hymenphysa M'Clelland, 1839 and Hymenophysa<br />
Bleeker, 1858, erroneously used for the present genus.<br />
2.7.1 Syncrossus beauforti (Smith, 1931)<br />
Botia beauforti Smith, 1931: 2, fig. 1 (type locality: Thailand:<br />
Nakon Sritamarat Province: Tadi Stream at Ban<br />
Kiriwong; holotype: USNM 90285; noun in genitive,<br />
indeclinable)<br />
2.7.2 Syncrossus berdmorei Blyth, 1860<br />
Syncrossus Berdmorei Blyth, 1860: 166 (type locality: Burma:<br />
Tenasserim provinces; syntypes: ZSI F 2636/1 [4],<br />
Menon & Yazdani, 1968: 120; noun in genitive, indeclinable)<br />
2.7.3 Syncrossus helodes (Sauvage, 1876)<br />
Botia helodes Sauvage, 1876: 99 (type locality: Cambodia:<br />
Tma-Kré; holotype: MNHN 8595, Kottelat, 1984a: 807;<br />
noun in apposition, indeclinable)<br />
2.7.4 Syncrossus hymenophysa (Bleeker, 1852)<br />
Cobitis hymenophysa Bleeker, 1852: 602 (type locality: Indonesia:<br />
Sumatra: Palembang; holotype: RMNH 7059,<br />
Alfred, 1961: 33; compound noun, indeclinable)<br />
Hymenophysa MacClellandi Bleeker, 1859a: 358 (unnecessary<br />
replacement name for Cobitis hymenophysa Bleeker,<br />
1852: 602; material listed as types by Fricke, 1991:<br />
12 has no type status; noun in genitive, indeclinable)<br />
2.7.5 Syncrossus lucasbahi (Fowler, 1937)<br />
Botia lucas-bahi Fowler, 1937: 154, fig. 70 (type locality:<br />
Thailand: Tachin [Tha Chin, Samut Sakhon; 13°32'22"N<br />
100°15'20"E]; holotype: ANSP 68005, Grant, 2007b: fig.<br />
80; noun in genitive, indeclinable)<br />
Botia Beauforti var. formosa Pellegrin & Fang, 1940: 119,<br />
fig. 5 (type locality: Laos: Ban Nam Khueng, 30 km northwest<br />
<strong>of</strong> Ban Houei Sai, about 6 km from Mekong; syntypes:<br />
MNHN 1939.218–220 [5]; adjective, -us, -a, -um)<br />
Botia yunnanensis Chen, 1980: 6, fig. 1 (type locality: China:<br />
Yunnan: Jinghong; holotype: IHB 638040, Grant,<br />
2007b: fig. 84; adjective, -is, -is, -e)
2.6.4 Sinibotia robusta, CMK10824, 66.3 mm SL; China: aquarium-fish<br />
trade.<br />
2.7.2 Syncrossus berdmorei, CMK 18421, 111 mm SL; Myanmar:<br />
Ataran drainage.<br />
2.7.6 Syncrossus reversus (Roberts, 1989)<br />
Botia reversa Roberts, 1989: 102, fig. 76 (type locality: Indonesia:<br />
Borneo: Kalimantan Barat: Sungei Pinoh, 37 km<br />
south <strong>of</strong> Nangapinoh; MZB 3521; adjective, -us, -a, -um)<br />
2.8 Yasuhikotakia Nalbant, 2002<br />
Yasuhikotakia Nalbant, 2002: 317 (type species: Botia modesta<br />
Bleeker, 1864c: 11, by original designation). Gender<br />
feminine.<br />
2.8.1 Yasuhikotakia caudipunctata (Taki & Doi, 1995)<br />
Botia caudipunctata Taki & Doi, 1995: 150, fig. 3 (type locality:<br />
Laos: Mekong River at Hatsalao, near Pakse; holotype:<br />
NSMT P 35935; adjective, -us, -a, -um)<br />
2.8.2 Yasuhikotakia eos (Taki, 1972)<br />
Botia eos Taki, 1972: 66, fig. 1 (type locality: Laos: Nam<br />
Ngum at mouth <strong>of</strong> Nam Khon at Tha Ngon, about 22 km<br />
north <strong>of</strong> Vientiane; holotype: NSMT P 14537; noun in<br />
apposition, indeclinable)<br />
2.8.3 Yasuhikotakia lecontei (Fowler, 1937)<br />
Botia lecontei Fowler, 1937: 156, figs. 71–74 (type locality:<br />
Thailand: Kemarat; holotype: ANSP 68006; noun in genitive,<br />
indeclinable)<br />
? Botia pulchripinnis Paysan, 1970: 156, pl. 13 fig. 10 (type<br />
locality: "Southeast Asia" [aquarium material imported<br />
from Thailand]; syntypes: at least the specimen on which<br />
the figure is based, not preserved; compound noun, indeclinable)<br />
2.8.4 Yasuhikotakia longidorsalis (Taki & Doi, 1995)<br />
Botia longidorsalis Taki & Doi, 1995: 148, fig. 1 (type locality:<br />
Laos: Mekong River at Hatsalao, near Pakse; holotype:<br />
NSMT P 35940; compound noun, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
20<br />
2.8.2 Yasuhikotakia eos, CMK 17124, 57.7 mm SL; Laos: Mekong<br />
drainage: Xe Bangfai.<br />
2.8.2 Yasuhikotakia eos, CMK 23084, 88.9 mm SL.<br />
2.8.5 Yasuhikotakia modesta (Bleeker, 1864)<br />
Botia modesta Bleeker, 1864c: 11 (type locality: Thailand:<br />
Ayuthaya; syntypes: MNHN 1851 [7], Bertin & Estève,<br />
1948: 91; figured in Bleeker, 1870: pl.; adjective, -us,<br />
-a, -um)<br />
Botia rubripinnis Sauvage, 1876: 100 (type locality: Cambodia:<br />
Phnom Penh; lectotype: MNHN 9545, designated by<br />
Kottelat, 1984a: 809; compound noun, indeclinable)<br />
2.8.6 Yasuhikotakia morleti (Tirant, 1885)<br />
Botia Morleti Tirant, 1885: 155 [1929: 133] (type locality:<br />
Vietnam: Thu-dau-mot; syntypes: MGHNL [lost, Kottelat,<br />
1987a: 17]; named for malacologist L.-J. Morlet, not<br />
to be confused with P.-A. Morelet, see Crosse, 1893: 75,<br />
78; noun in genitive, indeclinable)<br />
Botia horae Smith, 1931: 4, fig. 2 (type locality: Thailand:<br />
Kanchanaburi Province: "west fork (Kwe Noi) <strong>of</strong> the<br />
Meklong" [Kwai Noi River]; holotype: USNM 90286;<br />
Hora is treated as a Latinized name and horae is correct<br />
original spelling [Code art. 31.1.1 and Example]; noun<br />
in genitive, indeclinable)<br />
2.8.7 Yasuhikotakia splendida (Roberts, 1995)<br />
Botia splendida Roberts, 1995: 463, figs. 1–2 (type locality:<br />
Laos: Attapu Province: Xe Pian 5–6 km upstream from<br />
Ban Hin Lat and 1 km downstream from Se Pa waterfall,<br />
14°45'10"N 106°27'50"E; holotype: ZRC 39215; adjective,<br />
-us, -a, -um)
3.1.1 Vaillantella cinnamomea, CMK 21831, 68.3 mm SL; Indonesia:<br />
Borneo: Mahakam drainage.<br />
4.1.1 Acanthopsoides delphax, CMK 16178, 56.5 mm SL; Thailand:<br />
Salween drainage: Mae Hon Son.<br />
4.2.3 Acantopsis spectabilis, CMK 14683, 146 mm SL; Thailand:<br />
Salween drainage: Mae Nam Moei.<br />
3 Vaillantellidae Nalbant & Banarescu, 1977<br />
Vaillantellinae Nalbant & Bănărescu, 1977: 100 (type genus:<br />
Vaillantella Fowler, 1905a: 474)<br />
3.1 Vaillantella Fowler, 1905<br />
Vaillantella Fowler, 1905a: 474 (type species: Nemacheilus<br />
euepipterus Vaillant, 1902: 137, by original designation).<br />
Gender feminine.<br />
3.1.1 Vaillantella cinnamomea Kottelat<br />
Vaillantella cinnamomea Kottelat, 1994a: 428, fig. 1 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Mahakam<br />
drainage: Sungei Behernas, a blackwater tributary<br />
<strong>of</strong> Mahakam River immediately upriver <strong>of</strong> Merimun,<br />
0°05'S 115°47'E; holotype: MZB 5893; adjective, -us,<br />
-a, -um)<br />
4 Cobitidae Swainson, 1838<br />
Cobitidae Swainson, 1838: 360 (type genus: Cobitis Linnaeus,<br />
1758: 303; correct spelling confirmed by ICZN,<br />
1988: 178 [Opinion 1500]). See Steyskal (1980: 170)<br />
for gramatically correct (but nomenclaturally incorrect)<br />
spelling <strong>of</strong> the name.<br />
Acanthopsides Heckel & Kner, 1858: 296 (type genus: Acanthopsis<br />
Agassiz, 1832: 134, not Acantopsis van Hasselt,<br />
1823: 133)<br />
Misgurninae Fowler, 1905a: 474 (type genus: Misgurnus La<br />
Cepède, 1803: 16)<br />
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Family VAILLANTELLIDAE<br />
Family COBITIDAE<br />
21<br />
4.2 Acantopsis sp., CMK 23079, 88.4 mm SL; Laos:Mekong drainage:<br />
Xe Bangfai.<br />
3.1.2 Vaillantella euepiptera (Vaillant, 1902)<br />
Nemacheilus euepipterus Vaillant, 1902: 137, fig. 41 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Pontianak;<br />
lectotype: RMNH 7781, designated by Nalbant &<br />
Bănărescu, 1977: 101; compound adjective, -us, -a, -um)<br />
3.1.3 Vaillantella maassi Weber & de Beaufort, 1912<br />
Vaillantella Maassi Weber & de Beaufort, 1912: 532, pl. 12<br />
fig. 2 (type locality: Indonesia: Sumatra: Gunung Sahilan<br />
on Kampar Kiri; holotype: ZMA 100.993 Nijssen et al.,<br />
1993: 216; noun in genitive, indeclinable)<br />
Vaillantella flav<strong>of</strong>asciata Tweedie, 1956: 59, pl. 6 fig. b (type<br />
locality: Malaysia: Pahang: King George V National Park<br />
[Taman Negara], Tahan River near Kuala Tahan; holotype:<br />
BMNH 1957.1.23.4; adjective, -us, -a, -um)<br />
4.1 Acanthopsoides Fowler, 1934<br />
Acanthopsoides Fowler, 1934: 103 (type species: Acanthopsoides<br />
gracilis Fowler, 1934: 103, by original designation).<br />
Gender masculine.<br />
Neacanthopsis Smith, 1945: 297 (type species: Neacanthopsis<br />
gracilentus Smith, 1945: 297, by original designation).<br />
Gender feminine.<br />
4.1.1 Acanthopsoides delphax Siebert, 1991<br />
Acanthopsoides delphax Siebert, 1991a: 105, fig. 6 (type<br />
locality: Thailand: Salween River at Mae Sahm [Mae
Sahm Leap], west <strong>of</strong> Mae Sariang; holotype: USNM<br />
229043; noun in apposition, indeclinable)<br />
4.1.2 Acanthopsoides gracilentus (Smith, 1945)<br />
Neacanthopsis gracilentus Smith, 1945: 297, fig. 61 (type<br />
locality: Thailand: Mae Ping River, north <strong>of</strong> Chiang Mai;<br />
holotype: USNM 107952; adjective, -us, -a, -um)<br />
Acanthopsoides namromensis Nguyen, 2005: 559, fig. 8<br />
(type locality: Vietnam: Dien Bien Province: Mekong<br />
drainage: Nam Rom River; holotype: NCNTTSI; adjective,<br />
-is, -is, -e)<br />
4.1.3 Acanthopsoides gracilis Fowler, 1934<br />
Acanthopsoides gracilis Fowler, 1934: 103, fig. 55 (type<br />
locality: Thailand: Chiang Mai; holotype: ANSP 56999;<br />
adjective, -is, -is, -e)<br />
4.1.4 Acanthopsoides hapalias Siebert, 1991<br />
Acanthopsoides hapalias Siebert, 1991a: 106, fig. 7 (type<br />
species: Thailand: Nakhon Ratchasima Province: Mekong<br />
basin: Mae Nam Mun about 2 km downstream from Phimai,<br />
15°14'N 102°31'E; holotype: USNM 271723; noun<br />
in apposition, indeclinable)<br />
4.1.5 Acanthopsoides molobrion Siebert, 1991<br />
Acanthopsoides molobrion Siebert, 1991a: 107, fig. 8 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Boh River;<br />
holotype: RMNH 31273; noun in apposition, indeclinable)<br />
4.1.6 Acanthopsoides robertsi Siebert, 1991<br />
Acanthopsoides robertsi Siebert, 1991a: 109, fig. 10 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: small<br />
oxbow <strong>of</strong>f Kapuas River opposite Empangau, 124 km<br />
northeast <strong>of</strong> Sintang; 0°44'N 112°23'E; holotype: CAS<br />
49345; noun in genitive, indeclinable)<br />
4.2 Acantopsis van Hasselt, 1823<br />
Acantopsis van Hasselt, 1823: 133 (type species: Acantopsis<br />
dialuzona van Hasselt, 1823: 133, by monotypy).<br />
Gender feminine.<br />
Acanthopsis van Hasselt, 1824: 376, 377 (incorrect subsequent<br />
spelling <strong>of</strong> Acantopsis van Hasselt, 1823: 133)<br />
? Aperioptus Richardson, 1848: 27 (type species: Aperioptus<br />
pictorius Richardson, 1848: 27, by monotypy). Gender<br />
masculine.<br />
Prostheacanthus Blyth, 1860: 167 (type species: Prostheacanthus<br />
spectabilis Blyth, 1860: 167, by monotypy).<br />
Gender masculine.<br />
4.2.1 Acantopsis dialuzona van Hasselt, 1823<br />
Acantopsis Dialuzona van Hasselt, 1823: 133 (type locality:<br />
Indonesia: Java: Batavia [Jakarta]; syntypes: RMNH<br />
2707 [1]; unpublished van Hasselt's figure reproduced<br />
in Roberts, 1993: fig. 25; compound noun, indeclinable)<br />
? Aperioptus pictorius Richardson, 1848: 27, pl. 10 fig. 4<br />
(type locality: Borneo; syntypes [2]: lost; adjective, -us,<br />
-a, -um)<br />
Acanthopsis biaculeata Rüppell, 1852: 28 (nomen nudum)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
22<br />
Cobitis choirorhynchos Bleeker, 1854b: 95 (type locality:<br />
Indonesia: Sumatra: Palembang: at confluence <strong>of</strong> Lamatang<br />
and Enim Rivers; lectotype: RMNH 4977, designated<br />
by Alfred, 1961: 33; compound noun, indeclinable)<br />
Cobitis macrorhynchos Bleeker, 1854b: 95 (unnecessary<br />
replacement name for Acantopsis dialuzona van Hasselt,<br />
1823: 133 [lectotype designation by Alfred, 1961: 34, is<br />
invalid as this is a replacement name and not a new name];<br />
compound noun, indeclinable)<br />
4.2.2 Acantopsis octoactinotos Siebert, 1991<br />
Acantopsis octoactinotos Siebert, 1991b: 910, fig. 2 (type<br />
locality: Malaysia: Borneo: Sabah: Kinabatangan District:<br />
tributary <strong>of</strong> Kinabantangan River in lowlands near<br />
Sungei Deramakot, approx. 5°18'N 117°33'E; holotype:<br />
FMNH 68148; compound noun, indeclinable)<br />
4.2.3 Acantopsis spectabilis (Blyth, 1860)<br />
Prostheacanthus spectabilis Blyth, 1860: 167 (type locality:<br />
Burma: Tenasserim provinces; types: ? ZSI; adjective,<br />
-is, -is, -e)<br />
? Acantopsis multistigmatus Vishwanath & Laisram, 2005:<br />
433, fig. 1 (type locality: India: Manipur: Chindwin drainage:<br />
Lokchao stream, a tributary <strong>of</strong> Yu River; holotype:<br />
MUMF 3044; also spelt mu/tistigmatus p. 435, an obvious<br />
inadvertent error, thus incorrect original spelling<br />
[Code art. 32.5.1]; adjective, -us, -a, -um)<br />
4.2.4 Acantopsis thiemmedhi Sontirat, 1999<br />
Acanthopsis thiemmedhi Sontirat, 1997: 99 (not available,<br />
name published in meeting material, Code art. 9.9)<br />
Acantopsis thiemmedhi Sontirat, 1999: 66, fig. 1 (type locality:<br />
Thailand: Uthaithani Province: Amphoe Lan Sak:<br />
Huey Kha Khaeng Wildlife Sanctuary, Huey Nam Khoon;<br />
holotype: KUMF 3131; noun in genitive, indeclinable)<br />
4.3 Bibarba Chen & Chen, 2007<br />
Bibarba Chen & Chen, 2007: 105 (type species: Bibarba<br />
bibarba Chen & Chen, 2007: 105, by original designation).<br />
Gender feminine.<br />
4.3.1 Bibarba bibarba Chen & Chen, 2007<br />
Bibarba bibarba Chen & Chen, 2007: 105, fig. A-B (type<br />
locality: China: Guangxi: Du'an County (23°55–56'N<br />
108°06'E): Chengjiang River, tributary <strong>of</strong> Hongshuihe<br />
River, upper reaches <strong>of</strong> Pearl River; holotype: IHB<br />
87024411; compound noun, indeclinable)<br />
4.4 Canthophrys Swainson, 1838<br />
Canthophrys Swainson, 1838: 364 (type species Canthophrys<br />
albescens Swainson, 1839: 310, by subsequent<br />
designation by Swain, 1882: 282; no species originally<br />
included, first inclusion by Swainson, 1839: 190, 310).<br />
Gender feminine.<br />
Somileptus Swainson, 1839: 190, 311 (as subgenus <strong>of</strong> Canthophrys<br />
Swainson, 1839: 364; type species: Somileptes<br />
bispinosa Swainson, 1838: 311, by subsequent designation<br />
by Bleeker, 1863a: 36, 1863c: 3; also spelt Somi-
leptes on p. 311, first reviser [Nalbant, 1963: 364] gave<br />
precedence to Somileptus, see Kottelat, 1998c: 118).<br />
Gender masculine.<br />
4.4.1 Canthophrys gongota (Hamilton, 1822)<br />
Cobitis gongota Hamilton, 1822: 351, 394 (type locality:<br />
India: "rivers <strong>of</strong> Northern Bengal towards the mountains"<br />
[Bangladesh: Patgong, Rangpur District, 25°45'N<br />
89°15'E; Hora, 1935a: 49]; types: lost; Hamilton's unpublished<br />
figure reproduced in M'Clelland, 1839: pl. 51<br />
fig. 1; noun in apposition, indeclinable)<br />
Cobitis cucura Hamilton, 1822: 352, 394 (type locality: India:<br />
Kosi River [at Nathpur near the Nepal frontier; Hora,<br />
1935a: 49]; types: lost; Hamilton's unpublished figure<br />
reproduced in M'Clelland, 1839: pl. 51 fig. 2; simultaneous<br />
subjective synonym <strong>of</strong> Cobitis gongota Hamilton,<br />
1822: 351, first reviser [apparently Günther, 1868: 363]<br />
gave precedence to C. gongota; noun in apposition, indeclinable)<br />
Cobitis oculata M'Clelland, 1839: 303, 433, pl. 51 fig. 1<br />
(type locality: India: Upper Assam / northern parts <strong>of</strong><br />
Bengal near the foot <strong>of</strong> the mountains; syntypes: SMF<br />
412 [4], 8979 [1], Eschmeyer & Fricke, 2010 [plus basis<br />
<strong>of</strong> Cobitis gongota Hamilton, 1822: 351]; adjective, -us,<br />
-a, -um)<br />
Canthophrys albescens Swainson, 1839: 310 (available by<br />
indication to Hamilton, 1822: "Cob. No. 3" [which is<br />
Cobitis cucura]; type locality: India: Kosi River [at Nathpur<br />
near the Nepal frontier; Hora, 1935a: 49]; types: NT;<br />
Hamilton's unpublished figure reproduced in M'Clelland,<br />
1839: pl. 51 fig. 2; participle, indeclinable)<br />
Somileptes bispinosa Swainson, 1839: 311 (available by indication<br />
to Hamilton, 1822: 351 [which is Cobitis gongota];<br />
type locality: India: "rivers <strong>of</strong> Northern Bengal towards<br />
the mountains" [Bangladesh: Patgong, Rangpur<br />
District, 25°45'N 89°15'E; Hora, 1935a: 49]; types: lost;<br />
Hamilton's unpublished figure reproduced in M'Clelland,<br />
1839: pl. 51 fig. 1; adjective, -us, -a, -um)<br />
? Cobitis amnicola Valenciennes, in Cuvier & Valenciennes,<br />
1846: 68 (type locality: India: Bengal; syntypes: MNHN<br />
B.2171 [3], Spillmann, 1962: 587; noun in apposition,<br />
indeclinable)<br />
4.5 Cobitis Linnaeus, 1758<br />
Cobitis Linnaeus, 1758: 303 (type species: Cobitis taenia<br />
Linnaeus, 1758: 303, designated by ICZN, 1988 [Opinion<br />
1500]). Gender feminine.<br />
Acantophthalmus van Hasselt, 1823:132 (type species: Cobitis<br />
taenia Linnaeus, 1758: 303, by monotypy [see Kottelat,<br />
1987b: 371]; on Official Index <strong>of</strong> Rejected Names,<br />
ICZN, 1988: 178 [Opinion 1500]). Gender masculine.<br />
Acanthophthalmus van Hasselt, 1824: 377 (incorrect subsequent<br />
spelling <strong>of</strong> Acantophthalmus van Hasselt, 1823:<br />
132; on Official Index <strong>of</strong> Rejected Names, ICZN, 1992b:<br />
248 [Opinion 1695])<br />
? Acanthopsis Agassiz, 1832: 134 (type species: Acanthopsis<br />
angustus Agassiz, 1835: vol. 5, pl. 50 fig. 2, by subsequent<br />
monotypy in Agassiz, 1835: pl. 50; not a junior<br />
homonym <strong>of</strong> Acanthopsis van Hasselt, 1824: 377, which<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
23<br />
is an incorrect subsequent spelling <strong>of</strong> Acantopsis van<br />
Hasselt, 1823: 133, thus does not enter into homonymy<br />
[Code art. 33.3]; based on a fossil species, possibly not a<br />
cobitid (Kottelat, 1987b: 372)). Gender feminine.<br />
Cobitinula Hankó, 1924: 152 (type species: Cobitinula anatoliae<br />
Hankó, 1924: 152, by monotypy). Gender feminine.<br />
Acanestrinia Băcescu, 1962b: 435 (subgenus <strong>of</strong> Cobitis Linnaeus,<br />
1758: 303; type species: Cobitis elongata Heckel &<br />
Kner, 1858: 305, by original designation). Gender feminine.<br />
Bicanestrinia Băcescu, 1962b: 436 (subgenus <strong>of</strong> Cobitis<br />
Linnaeus, 1758: 303; type species: Cobitis simplicispina<br />
Hankó, 1924: 153, by original designation). Gender<br />
feminine.<br />
Iberocobitis Băcescu, 1962b: 438 (subgenus <strong>of</strong> Cobitis Linnaeus,<br />
1758: 303; type species: Acanthopsis taenia paludica<br />
de Buen, 1930: 34, by original designation). Gender<br />
feminine.<br />
Beyshehiria Erk'akan, Atalay-Ekmekçi & Nalbant, 1999: 20<br />
(subgenus <strong>of</strong> Cobitis Linnaeus, 1758: 303; type species:<br />
Cobitis bilseli Battalgil, 1942: 292, by original designation).<br />
Gender feminine.<br />
Taxonomic notes. Cobitis is obviously a polyphyletic genus.<br />
Several chromosome and molecular studies (e.g. Šlechtová<br />
et al., 2008; Kitagawa et al., 2011; Perdices et al., 2012)<br />
show a very complex situation and future research will certainly<br />
result in important changes in the taxonomy <strong>of</strong> Cobitis<br />
and the related genera (Iksookimia, Niwaella, Kichulchoia, etc.).<br />
Nomina nuda<br />
Cobitis longimanus Dybowski, in Sinicyn, 1900: 49 (nomen<br />
nudum; material: “Imperial Warsaw University” 2773;<br />
locality: Onon River)<br />
Cobitis poecilopleura Bleeker, 1872: 146 (nomen nudum;<br />
locality: China)<br />
Nomen nudum<br />
Cobitis yeni Nguyen, 1982: 26 (nomen nudum)<br />
Cobitis yeni Nguyen (T. T.), in Nguyen (V. H.), 2005: 227,<br />
fig. 114 (not available; locality: Vietnam: Ha Tinh Province:<br />
stream Ngam Pho at Huong Son)<br />
Nomenclatural notes. Nguyen (V. H.; 2005: 227) listed<br />
Nguyen (T. T., 1983: 83, pl. 3 fig. 2) as author <strong>of</strong> C. yeni.<br />
Nguyen (1983) is an unpublished thesis and the name is not<br />
available from it. As the description in Nguyen (2005) is<br />
from Nguyen (1983), I treat the author <strong>of</strong> the nomen nudum<br />
as Nguyen (T. T.), in Nguyen (V. H.). The Code art. 16.1<br />
requires that, after 1999, a new name must be explicitly indicated<br />
as intentionally new. This is not the case for C. yeni<br />
in Nguyen (2005) and the name is not available. Further, to<br />
be available, a new specific name published after 1999 must<br />
be accompanied by the explicit designation <strong>of</strong> a holotype or<br />
syntypes (Code art. 16.4). Nguyen (2005) mentioned that<br />
the description is based on 19 specimens but they are not<br />
mentioned as a holotype or syntypes.<br />
Species inquirenda<br />
4.5.1 Cobitis lachnostoma (Rutter, 1897)<br />
Acanthopsis lachnostoma Rutter, 1897: 60 (type locality:<br />
China: Guangdong: Swatow; holotype: CAS-SU 1812,<br />
Böhlke, 1953: 39; compound noun, indeclinable)
4.3 Bibarba sp., GIF 08090070, 56.7 mm SL; China: Guangxi:<br />
cave in Luocheng County. (Photograph by Lan Jia-Hu).<br />
4.3 Bibarba sp., GIF 08090070, 56.7 mm SL. (Photograph by Lan<br />
Jia-Hu).<br />
Species incertae sedis<br />
4.5.2 'Cobitis' arenae (Lin, 1934)<br />
Misgurnus arenae Lin, 1934: 227, figs. 5–6 (type locality:<br />
China: Guangdong: Hwei-yang County: shallow stream<br />
near Western Lake; holotype: FESC M 10; noun in genitive,<br />
indeclinable)<br />
Taxonomic notes. Apparently represents an unnamed genus<br />
close to Cobitis.<br />
Species incertae sedis<br />
4.5.3 'Cobitis' guttata (Nguyen, 2005)<br />
Acantopsis guttatus Nguyen, 2005: 557, fig. 7 (type locality:<br />
Vietnam: Ha Giang Province: Lo River, Bac Quang<br />
district, Suoi Mu stream in Tan Thanh village; holotype:<br />
NCNTTSI; adjective, -us, -a, -um)<br />
Taxonomic notes. Apparently the species tentatively identified<br />
as Acantopsis arenae by Kottelat (2001a: 49, fig. 95).<br />
Apparently represents an unnamed genus close to Cobitis.<br />
Nomenclatural notes. If placed in the genus Cobitis, then it<br />
is a junior secondary homonym <strong>of</strong> Cobitis guttata M'Clelland,<br />
1839. I do not propose a new replacement name because it<br />
will soon be transferred to another genus and the original<br />
name will then again be valid.<br />
4.5.4 Cobitis amphilekta Vasil’eva & Vasil’ev, 2012<br />
Cobitis amphilekta Vasil’eva & Vasil’ev, 2012: 177, figs. 2–<br />
3 [p. 200 <strong>of</strong> translation] (type locality: Azerbaijan: Caspian<br />
Sea near Port Il'icha between old bed <strong>of</strong> Kumbashi<br />
River and state farm; holotype: ZMMU P-22794; Greek<br />
adjective, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
24<br />
4.4.1 Canthophrys gongota, CMK 14401, 65.1 mm SL; India:<br />
Brahmaputra drainage.<br />
4.5.21 Cobitis hellenica, CMK 16951, 82.8 mm SL; Greece: Louros<br />
drainage.<br />
4.5.5 Cobitis arachthosensis Economidis & Nalbant, 1997<br />
Cobitis hellenica arahthosensis Economidis, 1991: 28, 1992:<br />
75 (nomen nudum; locality: Greece: Epiros: Louros and<br />
Arachthos Rivers)<br />
Cobitis arachthosensis Economidis & Nalbant, 1997: 313,<br />
fig. 12a–f (type locality: Greece: Epiros: dead branch <strong>of</strong><br />
Arachthos River near Akropotamia village; holotype:<br />
DZAUT 1988-137; adjective, -is, -is, -e)<br />
4.5.6 Cobitis bilineata Canestrini, 1865<br />
Cobitis taenia var. bilineata Canestrini, 1865: 305 (type<br />
locality: Italy: Modenese and Treviso; syntypes: LU; also<br />
in Canestrini, 1866: 148; adjective, -us, -a, -um)<br />
Cobitis taenia var. puta Cantoni, 1882: 362, pl. 1 fig. 1 (type<br />
locality: Italy: Torino and Vercellese; syntypes: LU; adjective,<br />
-us, -a, -um)<br />
Cobitis taenia var. septa Cantoni, 1882: 363, pl. 1 fig. 2<br />
(type locality: Italy; types: NT ?; participle, -us, -a, -um)<br />
4.5.7 Cobitis bilseli Battalgil, 1942<br />
Cobitis bilseli Battalgil, 1942: 292, fig. 4 (type locality: Turkey:<br />
Lake Beysehir; syntypes: LU; noun in genitive, indeclinable)<br />
4.5.8 Cobitis biwae Jordan & Snyder, 1901<br />
Cobitis taenia japonica Temminck & Schlegel, 1846: 222,<br />
pl. 103 fig. 3 (type locality: Japan [surroundings <strong>of</strong> Nagasaki];<br />
lectotype: RMNH 2703a, designated by Boeseman,<br />
1947: 167; junior primary homonym <strong>of</strong> Cobitis<br />
japonica Houttuyn, 1782: 337; adjective, -us, -a, -um)<br />
Cobitis Schlegeli Bleeker, 1863a: 41 (replacement name for<br />
Cobitis taenia japonica Temminck & Schlegel, 1846:<br />
222; here declared nomen oblitum under Code art. 23.9.2,<br />
as it has not been used as a valid name since 1899 [Code<br />
art. 23.9.1.1], and Cobitis biwae Jordan & Snyder, 1901:<br />
748 has been used in at least 25 works in the last 50 years<br />
[Code art. 23.9.1.2]; noun in genitive, indeclinable)<br />
Cobitis biwae Jordan & Snyder, 1901: 748 (replacement<br />
name for Cobitis taenia japonica Temminck & Schlegel,<br />
1846: pl. 103; here declared nomen protectum under Code<br />
art. 23.9.2, used in at least 25 works in the last 50 years,<br />
listed under Remarks [Code art. 23.9.1.2]; noun in genitive,<br />
indeclinable)<br />
Nomenclatural notes. Cobitis schlegeli has not been used<br />
as the valid name <strong>of</strong> a species after 1899 (Code art. 23.9.1.1)
and C. biwae has been used in a large number <strong>of</strong> works.<br />
Cobitis biwae is declared a nomen protectum and C. schlegeli<br />
is declared a nomen oblitum (Code art. 23.9.2). List <strong>of</strong> 28<br />
works in which Cobitis biwae Jordan & Snyder, 1901: 748<br />
is used as a valid name, published by at least 10 authors, in<br />
the immediately preceeding 50 years and encompassing a<br />
span <strong>of</strong> not less than 10 years (Code art. 23.9.2): Arai, 2003:<br />
113; Kawanabe et al., 2001: 392; Kimizuka & Kobayasi,<br />
1983: 308; Kitagawa et al., 2001: 250, 2003a: 93, 2003b:<br />
318, 2004: 117, 2005: 116, 2009: 13; Masuda et al., 1984:<br />
58; Matsuura et al., 2000: 162; Nakabo, 1993: 232, 2002:<br />
274; Nakajima, 1987: 208; Sawada, 1976: 176, 1982: 68;<br />
Sawada & Azizawa, 1983: 318; Sezaki & Kobayasi, 1978:<br />
851; Sezaki et al., 1988: 149; Shimizu, 2003a: 85, 2003b:<br />
155, 2008: 102; Shimizu et al., 1998: 381, 2004: 241; Suzawa,<br />
2006: 315; Tominaga et al., 2009: 195; Ueno & Ojima,<br />
1976: 446; Watanabe, 1998: 263.<br />
4.5.9 Cobitis calderoni Bacescu, 1962<br />
Cobitis Calderóni Băcescu, 1962b: 440, pl. 1 figs. 2–3, pl. 2<br />
figs. 6–7 (type locality: Spain: Burgos: stream Arlanzón,<br />
tributary <strong>of</strong> Duero at Cardena-Jimeno, Charcas de San<br />
Medel and La Gravera; holotype: MGAB 49415, Kottelat,<br />
1997: 87; must be emended in calderoni, Code art.<br />
32.5.2.1; noun in genitive, indeclinable)<br />
4.5.10 Cobitis choii Kim & Son, 1984<br />
Cobitis choii Kim & Son, 1984: 50, fig. 1 (type locality:<br />
South Korea: Chungcheongbug-do province: Cheongwon-gun<br />
district: Ochang-myon municipality: Miheocheon<br />
stream, a tributary <strong>of</strong> Geum River at Yeocheon-ri;<br />
holotype: CNUC 4854; noun in genitive, indeclinable)<br />
4.5.11 Cobitis dalmatina Karaman, 1928<br />
Cobitis taenia dalmatina Karaman, 1928: 163 (type locality:<br />
Croatia: Dalmatia: Cetina River; syntypes: MMNHS,<br />
lost; also in Karaman, 1929: 173; adjective, -us, -a, -um)<br />
4.5.12 Cobitis dolichorhynchus Nichols, 1918<br />
Cobitis dolichorhynchus Nichols, 1918: 16 (type locality:<br />
China: Fukien [Fujian]: Futsing; holotype: AMNH 7026;<br />
compound noun, indeclinable)<br />
4.5.13 Cobitis elazigensis Coad & Sarieyyüpoglu, 1988<br />
Cobitis elazigensis Coad & Sarieyyüpoglu, 1988: 426, fig.<br />
1 (type locality: Turkey: Elazig Province: a creek at Cip,<br />
15 km west <strong>of</strong> Elazig, in drainage <strong>of</strong> Murat Nehri, a tributary<br />
<strong>of</strong> Euphrates River; 38°42'N 39°05'E; holotype:<br />
NMC 85-0679A; adjective, -is, -is, -e)<br />
Taxonomic notes. "Provisionally considered" as synonym<br />
<strong>of</strong> C. simplicispina by Erkakan et al. (1999: 24).<br />
4.5.14 Cobitis elongata Heckel & Kner, 1858<br />
Cobitis elongata Heckel & Kner, 1858: 305, fig. 164 (type<br />
locality: Slovenia: Krain [Carniola]: Sala River near Idria;<br />
syntypes: NMW 48609 [2], 48610 [3], Kottelat, 1997:<br />
88; adjective, -us, -a, -um)<br />
4.5.15 Cobitis elongatoides Bacescu & Maier, 1969<br />
Cobitis taenia var. elongatoïdes Băcescu, 1962a: 294 (in-<br />
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25<br />
frasubspecific, name not available; locality: southwestern<br />
Romania: Neajlov, Jiu and Sîi Rivers)<br />
Cobitis taenia elongatoides Băcescu & Maier, 1969: 60 [p.<br />
39 <strong>of</strong> translation], figs. 3a, 7-1 (type locality: Romania:<br />
Argesel River, Danube drainage; syntypes: LU; adjective,<br />
indeclinable)<br />
Cobitis taenia danubialis Nalbant, 1993: 109, fig. 29 (type<br />
locality: Romania: Transilvania: Mures River at Sarmas,<br />
Harghita; holotype: MGAB 49923; adjective, -is, -is, -e)<br />
Cobitis megaspila Nalbant, 1993: 108, fig. 24 (type locality:<br />
Romania: delta <strong>of</strong> Danube River at Caraorman; holotype:<br />
ISBB 4497; compound noun, indeclinable)<br />
4.5.16 Cobitis evreni Erk'akan, Özeren & Nalbant, 2008<br />
Cobitis evreni Erk'akan, Özeren & Nalbant, 2008a: 112, fig.<br />
1 (type locality: Turkey: Kömür stream, Göksun, Kahramanmaras;<br />
38°00'52.24"N 36°30'31.11"E; holotype:<br />
HUIC CEY-2; noun in genitive, indeclinable)<br />
4.5.17 Cobitis fahireae Erk'akan, Atalay-Ekmekçi &<br />
Nalbant, 2008<br />
Cobitis fahireae Erk'akan, Atalay-Ekmekçi & Nalbant, 1998:<br />
10, fig. 2 (type locality: Turkey: Küçük Menderes, Selçuk-<br />
Aydin; holotype: HUIC uncat.; spelled fahirae on p. 10,<br />
fahireae on p. 11, as first reviser I select fahireae as the<br />
correct original spelling; noun in genitive, indeclinable)<br />
4.5.18 Cobitis faridpaki Mousavi-Sabet, Vasil’eva, Vatandoust<br />
& Vasil’ev, 2011<br />
Cobitis faridpaki Mousavi-Sabet, Vasil’eva, Vatandoust &<br />
Vasil’ev, 2011: 928, figs. 3–4 (type locality: Iran: Mazandaran<br />
region: Siahrud River, 36°26'85.05"N<br />
52°56'70.08"E; holotype: GUIC CC1403MA; noun in<br />
genitive, indeclinable)<br />
4.5.19 Cobitis gladkovi Vasil'ev & Vasil'eva, 2008<br />
Cobitis melanoleuca gladkovi Vasil'ev & Vasil'eva, 2008: 9,<br />
fig. 9 (type locality: Russia: Severskii Donets River, Don<br />
drainage, Starodubovka village, Staraya Stanitsa; holotype:<br />
ZMMU R-21654; noun in genitive, indeclinable)<br />
Taxonomic notes. Vasil'ev & Vasil'eva (2008) treated C.<br />
gladkovi as a subspecies <strong>of</strong> C. melanoleuca. They are treated<br />
as distinct species as they satisfy the criteria <strong>of</strong> species<br />
under the Evolutionary Species Concept.<br />
4.5.20 Cobitis hankugensis Kim, Park, Son & Nalbant,<br />
2003<br />
Cobitis hankugensis Kim, Park, Son & Nalbant, 2003: 2,<br />
fig. 1 (type locality: Korea: Gyongsangnam-do province:<br />
Sanchong-gun district: Dojeon-ri in Sengbirryang-myon<br />
municipality; holotype: CNUC 33912; adjective, -is, -is,<br />
-e)<br />
4.5.21 Cobitis hellenica Economidis & Nalbant, 1997<br />
Cobitis hellenica Economidis, 1991: 28, 41 (figs. 1–2), 1992:<br />
75 (nomen nudum; locality not stated)<br />
Cobitis hellenica Economidis & Nalbant, 1997: 311, figs.<br />
10, 12g-i (type locality: Greece: Epiros: Barbanakos<br />
spring near Stephani village, Louros drainage; holotype:<br />
DZAUT 1989-115; adjective, -us, -a, -um)
4.5.34 Cobitis melanoleuca, CMK 18807, 71.3 mm SL; Mongolia:<br />
Selenge drainage: Eg Gol.<br />
4.5.22 Cobitis illyrica Freyh<strong>of</strong> & Stelbrink, 2007<br />
Cobitis illyrica Freyh<strong>of</strong> & Stelbrink, 2007a: 270, figs. 1–2<br />
(type locality: Croatia: small lake south <strong>of</strong> Lake Proložak,<br />
48°28'13"N 17°07'20"E, Imotsko polje; holotype: ZMB<br />
33753; adjective, -us, -a, -um)<br />
4.5.23 Cobitis jadovaensis Mustafic & Mrakovcic, in<br />
Mustafic, Marcic, Duplic, Mrakovcic, Caleta, Zanella,<br />
Buj, Podnar & Dolenec, 2008<br />
Cobitis jadovaensis Mustafić & Mrakovčić, in Mustafić,<br />
Marčić, Duplić, Mrakovčić, Caleta, Zanella, Buj, Podnar<br />
& Dolenec, 2008: 5, fig. 2 (type locality: Croatia:<br />
Jadova River at 44°30'14.6"N 15°32'42.5"E; holotype:<br />
PMR VP1654; adjective, -is, -is, -e)<br />
4.5.24 Cobitis kellei Erk'akan, Atalay-Ekmekçi & Nalbant,<br />
1998<br />
Cobitis kellei Erk'akan, Atalay-Ekmekçi & Nalbant, 1998:<br />
10, fig. 1 (type locality: Turkey: Tigris drainage: Göksu<br />
stream, Cinar, Diyarbakir; holotype: ISBB 4682; noun<br />
in genitive, indeclinable)<br />
4.5.25 Cobitis keyvani Mousavi-Sabet, Yerli, Vatandoust,<br />
Özeren & Moradkhani, 2012<br />
Cobitis keyvani Mousavi-Sabet, Yerli, Vatandoust, Özeren<br />
& Moradkhani, 2012: 8, figs. 3–4 (type locality: Iran:<br />
Mazandaran Province: Keselian stream, Talar River,<br />
southeast <strong>of</strong> Caspian Sea basin, 36°11'74.09"N<br />
53°00'92.01"E, 473 masl; holotype: GUIC CC1389MA;<br />
noun in genitive, indeclinable)<br />
4.5.26 Cobitis laoensis (Sauvage, 1878)<br />
Misgurnus laoensis Sauvage, 1878b: 241 (type locality: Laos<br />
[apparently erroneous; possibly Vietnam; Freyh<strong>of</strong> & Serov,<br />
2000b: 206]; holotype: MNHN A.840, Kottelat,<br />
1984a: 809, fig. 5; adjective, -is, -is, -e)<br />
? Cobitis longitaeniatus Ngo, 2008: 66, fig. 1 (type locality:<br />
Vietnam: Quang Binh Province: Bo Trach District, Phong<br />
Nha, Son Trach, 17°30'N 106°15'E; holotype: NCNTTSI;<br />
also spelt longitaenia p. 67, an obvious inadvertent error,<br />
thus incorrect original spelling [Code art. 32.5.1];<br />
adjective, -us, -a, -um)<br />
? Cobitis phongnhaensis Ngo, 2008: 68, fig. 3 (type locality:<br />
Vietnam: Quang Binh Province: Bo Trach District:<br />
Phong Nha, Son Trach, Khe Mon, 17°30'N 106°15'E;<br />
holotype: NCNTTSI; adjective, -is, -is, -e)<br />
4.5.27 ? Cobitis lebedevi Vasil'eva & Vasil'ev, 1985<br />
Cobitis lebedevi Vasil'eva & Vasil'ev, 1985: 464, fig. 1A (type<br />
locality: Russia: Russian Far East: Khabarovsk Krai:<br />
Amur River near Elabuga [48°48'N 135°52'E]; holotype:<br />
ZMMU P-16252; noun in genitive, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
26<br />
4.5.44 Cobitis punctilineata, CMK 12914, 46.5 mm SL; Greece:<br />
Angitis drainage.<br />
Taxonomic notes. Treated as synonym <strong>of</strong> C. choii by, e.g.<br />
Bogutskaya et al. (2008: 336). The ranges <strong>of</strong> C. choii and<br />
C. lebedevi are widely disjunct and it does not seem that Amur<br />
and Korean material has yet been compared side by side.<br />
4.5.28 Cobitis levantina Krupp & Moubayed, 1992<br />
Cobitis levantina Krupp & Moubayed, 1992: 14, figs. 1–2<br />
(type locality: Syria: outflow <strong>of</strong> Buhairat Hims near Qattina,<br />
34°40'N 36°37'E; holotype: SMF 24371; adjective,<br />
-us, -a, -um)<br />
4.5.29 Cobitis linea (Heckel, 1848)<br />
Acanthopsis linea Heckel, 1848b: 267 (type locality: Iran:<br />
Persepolis [Marvdasht Plain, about 70 km northeast <strong>of</strong><br />
Shiraz; 29°56'04"N 52°53'29"E]; syntypes: NMW 48560<br />
or 48450 [7], Eschmeyer & Fricke, 2010; noun in apposition,<br />
indeclinable)<br />
4.5.30 Cobitis lutheri Rendahl, 1935<br />
Cobitis taenia lutheri Rendahl, 1935: 330, figs. 1–4 (type<br />
locality: Russia: Santachesa River, near its confluence<br />
with lake Chanka [Khanka]; syntypes: MZH, NRM 10810<br />
[1], 10811 [1], 10812 [1], 10813 [1]; noun in genitive,<br />
indeclinable)<br />
4.5.31 Cobitis macrostigma Dabry de Thiersant, 1872<br />
Cobitis macrostigma Dabry de Thiersant, 1872: 191, pl. 49<br />
fig. 4 (type locality: lakes <strong>of</strong> central China; holotype:<br />
MNHN 5089, Bertin & Estève, 1948: 92; noun in apposition,<br />
indeclinable)<br />
4.5.32 Cobitis maroccana Pellegrin, 1929<br />
Cobitis taenia var. maroccana Pellegrin, 1929: 525, fig. 1<br />
(type locality: Morocco: Oued Tiflet at Tiflet and Sidi<br />
Yahia; syntypes: MNHN 1922.84 [1], 1922.85 [1],<br />
1925.372 [1]; adjective, -us, -a, -um)<br />
4.5.33 Cobitis matsubarae Okada & Ikeda, 1939<br />
Cobitis taenia matsubarae Okada & Ikeda, 1939: 91, 101<br />
(type locality: Japan: Kagosima-ken: Hisikari-Mura /<br />
Fukuoka-Ken: Siba-Mura, Jôzima, and Naka River /<br />
Nagasaki-Ken: near Nagasaki; syntypes: LU [26], Eschmeyer<br />
& Fricke, 2010; Matsubara is treated as a Latinized<br />
name and matsubarae is correct original spelling<br />
[Code art. 31.1.1 and Example]; noun in genitive, indeclinable)<br />
4.5.34 Cobitis melanoleuca Nichols, 1925<br />
Cobitis taenia melanoleuca Nichols, 1925d: 3 (type locality:<br />
China: Shansi: Chin-ssu; holotype: AMNH 8403; treated<br />
as an adjective, -us, -a, -um)
4.5.54 Cobitis strumicae, CMK 17331, 86.7 mm SL; Greece: Evros<br />
drainage.<br />
Taxonomic notes. Vasil'ev & Vasil'eva (2008) recognised<br />
three subspecies on the basis <strong>of</strong> karyotypes: C. m. melanoleuca<br />
in China and the Amur drainage, C. m. sibirica (as<br />
granoei) in Siberia, and C. m. gladkovi in Europe. They are<br />
treated as valid species as they satisfy the criteria <strong>of</strong> species<br />
under the Evolutionary Species Concept.<br />
4.5.35 Cobitis meridionalis Karaman, 1924<br />
Cobitis taenia meridionalis Karaman, 1924: 75 (type locality:<br />
FYROM: Lake Prespa and its small tributaries; syntypes:<br />
MMNHS, lost; adjective, -is, -is, -e)<br />
4.5.36 Cobitis microcephala Chen & Chen, 2011<br />
Cobitis microcephala Chen & Chen, 2011: 147, fig. 2 (type<br />
locality: China: Guangxi: Bobai County: Nanliu River<br />
drainage, 22°28'N 109°95'E; holotype: IHB 0605135;<br />
compound adjective, -us, -a, -um)<br />
4.5.37 Cobitis multimaculata Chen & Chen, 2011<br />
Cobitis multimaculata Chen & Chen, 2011: 145, fig. 2 (type<br />
locality: China: Guangxi: Bobai County: Nanliu River<br />
drainage, 22°28'N 109°95'E; holotype: IHB 75v3203; an<br />
obvious adjective but declared to be used as a noun in<br />
original description, therefore indeclinable)<br />
4.5.38 Cobitis narentana Karaman, 1928<br />
Cobitis taenia narentana Karaman, 1928: 163 (type locality:<br />
Dalmatia [Croatia and Bosnia-Herzegovina]: Neretva<br />
River; syntypes: MMNHS, lost; also in Karaman,<br />
1929: 173; adjective, -us, -a, -um)<br />
4.5.39 Cobitis ohridana Karaman, 1928<br />
Cobitis taenia ohridana Karaman, 1928: 162, figs. 4a, 5 (type<br />
locality: FYROM: Lake Ohrid; syntypes: MMNHS, lost;<br />
also in Karaman, 1929: 173; adjective, -us, -a, -um)<br />
4.5.40 Cobitis paludica (de Buen, 1930)<br />
Acanthopsis taenia forma paludica de Buen, 1930: 34, fig. 48<br />
(type locality: Spain: Fuente del Roble, Talayuela,<br />
Cáceres; holotype: LU; available by indication, Code art.<br />
12.2.7; adjective, -us, -a, -um)<br />
Cobitis taenia paludicola Berg, 1932c: 155 (incorrect subsequent<br />
spelling <strong>of</strong> C. taenia paludica de Buen, 1930: 34)<br />
Cobitis taenia haasi Klausewitz, 1955: 42, fig. 2 (type locality:<br />
Spain: Albufera near Valencia; holotype: SMF 3232;<br />
noun in genitive, indeclinable)<br />
? Cobitis vitoriae Fernandez de la Cigoña Núñez & Ferreira,<br />
1996: 70, fig. (not available, name proposed conditionally<br />
after 1960 [Code art. 15.1]; locality: Spain: Galicia:<br />
lagoon next to Miño River at Caldelas, 7 km upstream<br />
<strong>of</strong> Tuy, Pontevedra)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
27<br />
4.5.62 Cobitis vardarensis, CMK 12982, 49.4 mm SL; Greece:<br />
Pinios drainage.<br />
? Cobitis victoriae Fernandez de la Cigoña Núñez, 1999:<br />
21, fig. (type locality: Spain: Miño River; syntypes: model<br />
<strong>of</strong> figure and specimens examined by Fernandez de la<br />
Cigoña Núñez & Ferreira, 1996: 66 [by reference on<br />
unnumbered p. 1 <strong>of</strong> Preface]; also spelt vicroriae, p. 1 <strong>of</strong><br />
Preface, an obvious inadvertent error, thus incorrect original<br />
spelling [Code art. 32.5.1]; noun in genitive, indeclinable)<br />
4.5.41 Cobitis phrygica Battalgazi, 1944<br />
Cobitis phrygica Battalgazi, 1944: 300, fig. 1 (type locality:<br />
Turkey: Vilâyet Afyonkarahisar, Aci Göl [Lake Acigöl,<br />
37°49'N 29°53'E]; syntypes: LU; adjective, -us, -a, -um)<br />
Cobitis Battalgili Băcescu, 1962b: 436, 437, fig. 1B (type<br />
locality: Turkey: Lakes Beysehir Gölü [37°41'0"N<br />
31°44'0"E] and Gölhisar Gölü [37°06'00"N 29°36'00"E];<br />
syntypes: MCSNG, ZMH 4744 [1], 4745 [4], Wilkens &<br />
Dohse, 1993: 412 [as holotype and paratypes]; noun in<br />
genitive, indeclinable [should have been battalgilae])<br />
Taxonomic notes. "Provisionally considered" as synonyms<br />
<strong>of</strong> C. simplicispina by Erkakan et al. (1999: 24).<br />
4.5.42 Cobitis pontica Vasil'eva & Vasil'ev, 2006<br />
Cobitis pontica Vasil'eva & Vasil'ev, 2006: S17, fig. (type<br />
locality: Bulgaria: Veleka River; holotype: ZMMU<br />
P-21363; adjective, -us, -a, -um)<br />
4.5.43 Cobitis puncticulata Erk'akan, Atalay-Ekmekçi &<br />
Nalbant, 1998<br />
Cobitis puncticulata Erk'akan, Atalay-Ekmekçi & Nalbant,<br />
1998: 12, fig. 4 (type locality: Turkey: Karaderetream<br />
[Karade stream ?] ath [at ?] outlet <strong>of</strong> Manyas (Kus) Lake;<br />
HUIC uncat.; adjective, -us, -a, -um)<br />
4.5.44 Cobitis punctilineata Economidis & Nalbant, 1997<br />
Cobitis punctilineata Economidis, 1991: 28, 41 (fig. 4), 1992:<br />
75 (nomen nudum; locality: Greece: Macedonia: Philippi<br />
Plain, stream Aggitis, tributary <strong>of</strong> Strymon)<br />
Cobitis punctilineata Economidis & Nalbant, 1997: 314, figs.<br />
11, 12j-l (type locality: Greece: Macedonia: Drama: channels<br />
near Kalambaki village, Strymon-Aggitis drainage;<br />
holotype: DZAUT 1992-39; adjective, -us, -a, -um)<br />
4.5.45 Cobitis rara Chen, 1981<br />
Cobitis rarus Chen, 1981: 24, fig. 1 (type locality: China:<br />
Shanxi: Lüeyang and Fengxian; syntypes [7]: IHB<br />
596070–74, 731121–122; adjective, -us, -a, -um)<br />
4.5.46 Cobitis satunini Gladkov, 1935<br />
Cobitis taenia satunini Gladkov, 1935: 73 (type locality:<br />
Georgia: Kintrich River [entering Black Sea at
41°48'14"N 41°46'09"E]; holotype: ZMMU P 2852,<br />
Mousavi-Sabet et al., 2011: 930, fig. 5; noun in genitive,<br />
indeclinable)<br />
4.5.47 Cobitis shikokuensis Suzawa, 2006<br />
Cobitis shikokuensis Suzawa, 2006: 316, figs. 2, 6 (type locality:<br />
Japan: Shikoku Island: Kochi Prefecture: Takaoka<br />
County: Shimanto Town, Morigauchi, Hinoji River at<br />
33°10'N 133°03'E, Shimanto River drainage; holotype:<br />
YCM-P 21981; adjective, -is, -is, -e)<br />
4.5.48 Cobitis sibirica Gladkov, 1935<br />
Cobitis taenia sibirica Gladkov, 1935 [between 1 Aug and<br />
19 Nov]: 73 (type locality: Russia: Lake Turgoïak, southern<br />
Urals [Chelyabinsk oblast: Lake Turgoyak, 55°09'N<br />
60°04'E]; holotype: ZMMU P2853; adjective, -us, -a,<br />
-um)<br />
Cobitis taenia granoei Rendahl, 1935 [before 16 Oct]: 332,<br />
figs. 5–6 (type locality: Russia: Siberia: Irtysch River near<br />
Omsk; holotype: MZH; noun in genitive, indeclinable;<br />
subjective simultaneous synonym <strong>of</strong> Cobitis taenia sibirica<br />
Gladkov, 1935: 73; first reviser [Berg, 1949: 894]<br />
gave precedence to C. t. sibirica.)<br />
Cobitis taenia sibirica morpha elongata Berg, 1949: 894<br />
(infrasubspecific, name not available; locality: [Mongolia<br />
?]: Lake Kosogol [? Khuvsgul])<br />
Cobitis granoei olivai Nalbant, Holčík & Pivnička, 1970:<br />
121, fig. 2, pl. fig. 1c (type locality: Mongolia: Archangaj<br />
County: Lake Ögijn-nuur and Narijn-gol River, an<br />
upper right tributary <strong>of</strong> Orkhon River, Selenge drainage;<br />
holotype: SNM RY-2093; noun in genitive, indeclinable)<br />
Taxonomic notes. Nalbant (1993: 108) treated C. sibirica<br />
(as C. granoei) as a synonym <strong>of</strong> C. melanoleuca but convincing<br />
evidence was not provided. Chen (1981) treated them<br />
as two distinct species. Prok<strong>of</strong>iev (2007c: 111) considered<br />
C. olivai as valid based on colour pattern, but Kottelat (2006:<br />
56) had commented that both colour patterns are present in<br />
different specimens from the same location in Mongolia.<br />
Vasil'ev & Vasil'eva (2008) treated C. sibirica (as C. granoei)<br />
as a subspecies <strong>of</strong> C. melanoleuca. They are treated<br />
here as distinct species as they satisfy the criteria <strong>of</strong> species<br />
under the Evolutionary Species Concept.<br />
Nomenclatural notes. The exact dates <strong>of</strong> publication <strong>of</strong><br />
Gladkov (1935) and Rendahl (1935) are not known. They<br />
are discussed by Bogutskaya & Naseka (2004: 105). Rendahl<br />
(1935) is recorded in the issue <strong>of</strong> Naturae Novitates (57 (10):<br />
129–144) received at ZISP on 16 October 1935 and could<br />
have precedence over Gladkov (1935), which is recorded in<br />
Naturae Novitates (57 (11): 145–150) received at ZISP on<br />
19 November 1935. On the other hand, Gladov (1935) is<br />
dated as sent to the press on 1 August 1935, but the actual<br />
printing and distribution dates are not known. These dates<br />
do not allow determination <strong>of</strong> which work appeared first;<br />
they are only indicative <strong>of</strong> the sequence in which they were<br />
received by Naturae Novitates in Berlin. It is possible that<br />
Gladkov (1935) was published before Rendahl (1935) but<br />
dispatched later or travelled more slowly because <strong>of</strong> some<br />
administrative burden, given the political situation in that<br />
period. This provides information on the latest possible publication<br />
date, but no objective information allowing deter-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
28<br />
mination <strong>of</strong> which work appeared first and which was first<br />
available from its respective publisher. For the time being it<br />
is better to consider them as simultaneous subjective synonyms,<br />
in which case Berg (1949: 894) is the first reviser<br />
and he gave precedence to C. sibirica. This is also the conclusion<br />
reached by a strict application <strong>of</strong> Code art. 21.7 using<br />
the data available today. [Naturae Novitates was a journal<br />
published in Berlin between 1879 and 1944 that listed<br />
new literature on various fields <strong>of</strong> natural sciences: Schmid,<br />
1984].<br />
4.5.49 Cobitis simplicispina Hankó, 1924<br />
Cobitis simplicispina Hankó, 1924: 153, pl. 3 fig. 7 (type<br />
locality: Turkey: Kötschke-Kissik [Gökçekısık (39°39'N<br />
30°24'E), a town 15 km southwest <strong>of</strong> Eskişehir<br />
(39°46'36"N 30°31'14"E); Porsuk River drainage]; syntypes:<br />
MNH [3], lost; invalid neotype designation by<br />
Erkakan, Atalay-Ekmekçi & Nalbant, 1999: 24; compound<br />
noun, indeclinable)<br />
Nomenclatural notes. Erk'akan, Atalay-Ekmekçi & Nalbant<br />
(1999: 24) designated a neotype for C. simplicispina. The<br />
neotype designation does not fulfill conditions <strong>of</strong> the Code<br />
art. 75.3.2, 75.3.3, 75.3.4 and is invalid. Further, at top <strong>of</strong><br />
p. 24, specimen ISBB 4694 is listed as neotype, and at bottom<br />
<strong>of</strong> same column, specimen HUIC 1 is listed as "selected<br />
here as neotype". This last specimen is the one in Fig. 9 but<br />
it is not labelled as neotype. In my reprint <strong>of</strong> the paper, the<br />
mention 'neotype' for ISBB 4694 had been crossed out (by<br />
Nalbant) indicating that ISBB 4694 was not intended as neotype;<br />
however, this information is external and formally irrelevant<br />
and, therefore, the simultaneous 'designation' <strong>of</strong> two<br />
neotypes makes the designation invalid in any case. The locality<br />
<strong>of</strong> the purported neotype (HUIC 1) is: Turkey: Eskişehir:<br />
Kocadere, Porsuk stream, Sakarya River drainage.<br />
4.5.50 Cobitis sinensis Sauvage & Dabry de Thiersant, 1874<br />
Cobitis sinensis Sauvage & Dabry de Thiersant, 1874: 16<br />
(type locality: China: brooks <strong>of</strong> western Sichuan [probably<br />
Pingwu County: Long'an; on Fu Jiang river, Jialing<br />
Jiang system, Yangtze drainage; 32°24'31"N<br />
104°31'36"E; see below]; syntypes: MNHN 6779 [4],<br />
Son & Kim, 2002: 241, fig. 1a, Bertin & Estève, 1948:<br />
92; adjective, -is, -is, -e)<br />
Taxonomic notes. The type locality <strong>of</strong> the syntypes <strong>of</strong> C.<br />
sinensis is given as Western Sichuan. The specimens were<br />
collected by A. David. In Sichuan, David visited Chengdu,<br />
then travelled to Muping. Muping is now in Western Sichuan,<br />
but at David's time it was an independant princedom and<br />
David's map shows it in Tibet (see Oreias dabryi). After returning<br />
to Chengdu, David wanted to visit Kokonoor [now<br />
China: Qinghai]. He reached to Lon-gan [Sichuan: Pingwu<br />
County: Long'an; also as Lunyan, Lung-yan, Lungan;<br />
32°24'31"N 104°31'36"E], explored that area, returned to<br />
Chengdu and Shanghai (David, 1871: 97). This makes Pingwu<br />
the only area where he collected within what he called<br />
Western Sichuan. Unfortunately, David did not publish a<br />
detailed account <strong>of</strong> this part <strong>of</strong> his second travel. His map<br />
(1875) shows that he made a number <strong>of</strong> small trips around<br />
Pingwu.
4.5.51 Cobitis splendens Erk'akan, Atalay-Ekmekçi &<br />
Nalbant, 1998<br />
Cobitis splendens Erk'akan, Atalay-Ekmekçi & Nalbant,<br />
1998: 11, fig. 3 (type locality: Turkey: small stream tributary<br />
to Black Sea, about 200 m from sea shore, 16 km<br />
east <strong>of</strong> Akçakoca [41°05'00"N 31°07'00"E] and about<br />
30 km south-west <strong>of</strong> Eregli; holotype: HUIC uncat.; adjective,<br />
indeclinable)<br />
4.5.52 Cobitis stephanidisi Economidis, 1992<br />
Cobitis stephanidisi Economidis, 1991: 27, 41 (fig. 7)<br />
(nomen nudum; locality: Greece: Thessaly: Kefalovrissos<br />
spring in Velestino village (Feres))<br />
Cobitis stephanidisi Economidis, 1992: 60 (type locality:<br />
Greece: Thessaly: Lake Karla and Kefalovrissos spring<br />
in Velestino village (Feres); syntypes [material listed by<br />
Economidis & Nalbant, 1997: 303]; lectotype: DZAUT<br />
1992-40, by present designation; noun in genitive, indeclinable)<br />
Cobitis stephanidisi Economidis & Nalbant, 1997: 303, fig. 6<br />
(type locality: Greece: Thessaly: spring in village Velestino,<br />
Lake Karla basin; holotype: DZAUT 1992-40; noun<br />
in genitive, indeclinable)<br />
Nomenclatural notes. The complete description <strong>of</strong> Cobitis<br />
stephanidisi appeared first in Economidis & Nalbant (1997);<br />
however, the use <strong>of</strong> the name accompanied by a diagnosis<br />
and a figure in Economidis (1992) makes the name available<br />
from this publication and with Economidis as author.<br />
As Economidis (1992) referred to the then in-press 1997<br />
paper, the specimens listed as holotype and paratypes by<br />
Economidis & Nalbant (1997) are in fact syntypes. Specimen<br />
DZAUT 1992-40 is here designated as lectotype <strong>of</strong><br />
C. stephanidisi Economidis, 1992.<br />
4.5.53 Cobitis striata Ikeda, 1936<br />
Cobitis taenia striata Ikeda, 1936: 984, 991, figs. 10-2,<br />
11-4 (type locality: Japan: Shikoku Island: Kagawa-Ken /<br />
Honshu Island: Osaka-fu; syntypes [240]: LU, Eschmeyer<br />
& Fricke, 2010; adjective, -us, -a, -um)<br />
4.5.54 Cobitis strumicae Karaman, 1955<br />
Cobitis taenia strumicae Karaman, 1955: 190, fig. 4 (type<br />
locality: FYROM: Monospitovo swamp and Strumica<br />
River; syntypes: MMNHS, lost; noun in genitive, indeclinable)<br />
Cobitis peschevi Sivkov & Dobrovolov, 1984: 1673, fig. 1<br />
(type locality: Bulgaria: Eleshnitsa River, a tributary <strong>of</strong><br />
Kamchiya River; holotype: MNHV 1627; noun in genitive,<br />
indeclinable)<br />
Cobitis rhodopensis Vassilev, 1998: 280, fig. 1 (type locality:<br />
Bulgaria: Biala River, a tributary <strong>of</strong> Maritsa River,<br />
downstream <strong>of</strong> Meden buk village; holotype: DHISUB<br />
199501C; adjective, -is, -is, -e)<br />
4.5.55 Cobitis taenia Linnaeus, 1758<br />
Cobitis Taenia Linnaeus, 1758: 303 (based on Artedi [1738:<br />
gen. 2 [4], syn. 3, spec. 4 [2], Cobitis aculeo bifurco ...],<br />
Linnaeus [1746: 126, n. 333, idem] and Gronovius<br />
[1754: 2, n. 5, idem]; type locality: "in Europae aquis<br />
dulcibus"; syntype: ? ZMUU 205, Wheeler, 1991: 163;<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
29<br />
noun in apposition, indeclinable)<br />
Cobitis spilura Holandre, 1837: 253 (type locality: Belgium:<br />
Liège / France: Moselle River, stream Nied below Malroy,<br />
Meuse River below Saint Mihiel; holotype: LU [description<br />
explicitly based on a single specimen; "syntypes":<br />
<strong>Museum</strong> <strong>of</strong> Metz, France, Blanchard, 1866: 288];<br />
author stated as Carlier, but Holandre is responsible for<br />
conditions making name available and thus is author;<br />
compound adjective, -us, -a, -um)<br />
Cobitis taenioides Băcescu & Maier, 1969: 52 [p. 39 <strong>of</strong> translation]<br />
(type locality: western Europe and Danube basin;<br />
syntypes: LU; adjective, indeclinable)<br />
4.5.56 Cobitis takatsuensis Mizuno, 1970<br />
Cobitis takatsuensis Mizuno, 1970: 133, figs. 1–2 (type locality:<br />
Japan: Honshu: Shimane-ken Prefecture: Kanoashigun<br />
District: Kabatani-gawa River, a tributary <strong>of</strong> Takatsu-gawa,<br />
at Kurobuchi in Kakinori-mura village; holotype:<br />
Collection Mizuno; adjective, -is, -is, -e)<br />
4.5.57 Cobitis tanaitica Bacescu & Maier, 1969<br />
Cobitis taenia tanaitica Băcescu & Maier, 1969: 52 [p. 39<br />
<strong>of</strong> translation], fig. 1 (type locality: Ukraine: Don River<br />
below Rostov; syntypes: LU [6]; adjective, -us, -a, -um)<br />
Cobitis rossomeridionalis Vasil'eva & Vasil'ev, 1998: 607,<br />
fig. 2 [p. 582 <strong>of</strong> translation] (type locality: Russia: Rostovskaya<br />
oblast: Don River near Rogozhkin [49°24'22"N<br />
41°37'57"E]; holotype: ZMMU P-20240; adjective, -is,<br />
-is, -e)<br />
4.5.58 Cobitis taurica Vasil'eva, Vasil'ev, Janko, Ráb &<br />
Rábová, in Janko, Vasil'ev, Ráb, Rábová, Šlechtová<br />
& Vasil'eva, 2005<br />
Cobitis taurica Vasil'eva, Vasil'ev, Janko, Ráb & Rábová, in<br />
Janko, Vasil'ev, Ráb, Rábová, Šlechtová & Vasil'eva,<br />
2005: 417, fig. 4a (type locality: Ukraine: Crimea: Chernaya<br />
River; holotype: ZMMU P-21358; adjective, -us,<br />
-a, -um)<br />
4.5.59 Cobitis tetralineata Kim, Park & Nalbant, 1999<br />
Cobitis tetralineata Kim, Park & Nalbant, 1999: 379, fig. 10<br />
(type locality: Korea: Deonranam-do province: Guraegun<br />
district: Somjin River at Guraegu; holotype: CNUC<br />
723, Kim, 2009: 11; adjective, -us, -a, -um)<br />
4.5.60 Cobitis trichonica Stephanidis, 1974<br />
Cobitis trichonica Stephanidis, 1974: 227, figs. 1–3 (type<br />
locality: Greece: Lake Trichonis; syntypes [34]: lost,<br />
Kottelat, 1997: 90; adjective, -us, -a, -um)<br />
4.5.61 Cobitis turcica Hankó, 1924<br />
Cobitis taenia turcica Hankó, 1924: 154, pl. 3 fig. 8 (type<br />
locality: Turkey: Eregli [apparently the one at 37°31'00"N<br />
34°03'00"E]; syntypes: MNH [2]; adjective, -us, -a, -um)<br />
? Cobitinula anatoliae Hankó, 1924: 152, pl. 3 fig. 6 (type<br />
locality: Turkey: Lake Ak-Göl [several lakes Akgöl in<br />
Turkey, probably the one near Ereli, since many <strong>of</strong><br />
Hankó's (1924) samples are from Ereli; 37°30'51"N<br />
33°43'02"E]; syntypes: MNH [2], lost; noun in genitive,<br />
indeclinable; simultaneous subjective synonym <strong>of</strong> Cobi-
4.6.2 Iksookimia koreensis, CMK 21096, 56.1 mm SL; South<br />
Korea: Mangyeong drainage.<br />
4.7.2 Kichulchoia multifasciata, CMK 21107, 99.3 mm SL; South<br />
Korea: Naktong drainage.<br />
tis taenia turcica Hankó, 1924: 154; as first reviser I give<br />
precedence to C. t. turcica)<br />
4.5.62 Cobitis vardarensis Karaman, 1928<br />
Cobitis taenia vardarensis Karaman, 1928: 163 (type locality:<br />
FYROM: Vardar River drainage; syntypes: MMNHS,<br />
lost; also in Karaman, 1929: 172; adjective, -is, -is, -e)<br />
Cobitis vardarensis kurui Erk'akan, Atalay-Ekmekçi & Nalbant,<br />
1998: 13, fig. 6 (type locality: Turkey: Menderes<br />
River, Selçuk-Aydin, Saplik Bridge; holotype: HUIC<br />
uncat.; noun in genitive, indeclinable)<br />
4.5.63 Cobitis vettonica Doadrio & Perdices, 1997<br />
Cobitis vettonica Doadrio & Perdices, 1997: 53, fig. 1 (type<br />
locality: Spain: Arrago River, Cadalso de Gata, Cáceres,<br />
Tagus River drainage; holotype: MNCM 18271; adjective,<br />
-us, -a, -um)<br />
4.5.64 Cobitis ylengensis Ngo, 2003<br />
Cobitis ylengensis Ngo, 2003: 18, fig. 2 (type locality: Vietnam:<br />
Quang Binh Province: Minh Hoa district: Dan Hoa<br />
village: Bai Dinh; holotype: NCNTTSI; adjective, -is,<br />
-is, -e)<br />
? Cobitis squataeniatus Ngo, 2008: 67, fig. 2 (type locality:<br />
Vietnam: Quang Binh Province: Minh Hoa District, Dan<br />
Hoa, Quy Dat; holotype: NCNTTSI; adjective, -us, -a,<br />
-um)<br />
Taxonomic notes. Figure 2 in the description <strong>of</strong> C. ylengensis<br />
is distorted and shows the fish more slender than described<br />
in the text.<br />
4.5.65 Cobitis zanandreai Cavicchioli, 1965<br />
Cobitis taenia zanandreai Cavicchioli, 1965: 1152, fig. 1<br />
(type locality: Italy: Caserta: Volturno River at Ponte<br />
Annibale, Capua; holotype: Istituto di Anatomia Comparata,<br />
Università di Ferrara; noun in genitive, indeclinable)<br />
4.5.66 Cobitis zhejiangensis Son & He, 2005<br />
Cobitis zhejiangensis Son & He, 2005: 237, figs. 2–3 (type<br />
locality: China: Zhejiang: Taizhou city: Xianju District:<br />
Ling River; holotype: SUBC 9619; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
30<br />
4.8.1 Koreocobitis naktongensis; CMK 27108, 119.6 mm SL;<br />
South Korea: Naktong drainage.<br />
4.6 Iksookimia Nalbant, 1993<br />
Iksookimia Nalbant, 1993: 101 (type species: Cobitis koreensis<br />
Kim, 1975: 51, by original designation). Gender<br />
feminine.<br />
Taxonomic notes. Iksookimia is possibly polyphyletic according<br />
to the molecular tree in Šlechtova et al. (2008).<br />
4.6.1 Iksookimia hugowolfeldi Nalbant, 1993<br />
Iksookimia hugowolfeldi Nalbant, 1993: 106, fig. 1 (type<br />
locality: South Korea: Yung San River; holotype: ISBB<br />
4495; noun in genitive, indeclinable)<br />
4.6.2 Iksookimia koreensis (Kim, 1975)<br />
Cobitis koreensis Kim, 1975: 51, fig. 1 (type locality: South<br />
Korea: Gyongi-do province: Gapyong-gun district: Jojong<br />
River, tributary <strong>of</strong> Han River at Sang-myon; holotype:<br />
CNUC 21674, Kim, 2009: 13; adjective, -is, -is, -e)<br />
4.6.3 Iksookimia longicorpus (Kim, Choi & Nalbant, 1977)<br />
Cobitis longicorpus Kim, Choi & Nalbant, 1977: 172, figs.<br />
1–2 (type locality: South Korea: Jeonrabug Do province:<br />
Sunchang Gun district: tributary <strong>of</strong> Seomjin River at<br />
Bogheung Myon; holotype: CNUC 505; compound noun,<br />
indeclinable)<br />
4.6.4 Iksookimia pacifica (Kim, Park & Nalbant, 1999)<br />
Cobitis pacifica Kim, Park & Nalbant, 1999: 380, fig. 12<br />
(type locality: South Korea: Kangwon-do province<br />
[Gangwon]: Kangreung-shi [Gangneung]: Yonkokhon<br />
stream about 2 km upstream <strong>of</strong> confluence with Eastern<br />
(Japan) Sea, Shingwhang-ri in Yonkok-myon municipality;<br />
holotype: CNUC 25111, Kim, 2009: 15; junior primary<br />
homonym <strong>of</strong> Cobitis pacifica Forster, in Lichtenstein,<br />
1844: 235; adjective, -us, -a, -um)<br />
Nomenclatural notes. The name Cobitis pacifica was first<br />
made available in a manuscript <strong>of</strong> Forster posthumously<br />
published by Lichtenstein (1844: 235). Lichtenstein (footnote)<br />
commented that this is Eleotris fusca (Schneider, 1801:<br />
453). The name has never been used since the original description<br />
and it could probably be declared nomen oblitum<br />
under Code art. 23.9.2. The case was discovered too late for<br />
researching usages <strong>of</strong> the junior homonym.<br />
4.6.5 Iksookimia pumila (Kim & Lee, 1987)<br />
Cobitis koreensis pumilus Kim & Lee, 1987: 58, figs. 1–2<br />
(type locality: South Korea: Chollabuk-do province:<br />
Puan-gun district: stream Paikchon at Sangso-myon; holotype:<br />
CNUC 9374; adjective, -us, -a, -um)<br />
4.6.6 Iksookimia yongdokensis Kim & Park, 1997<br />
Iksookimia yongdokensis Kim & Park, 1997: 250, fig. 2 (type<br />
locality: South Korea: Kyongsangbuk-do province: Yong-
4.9.2 Kottelatlimia katik, ZRC uncat., 12.3 mm SL; Indonesia:<br />
Borneo: Pembuang drainage. (Photograph by Tan Heok Hui).<br />
dok-gun district: Dalsan-myon municipality: Yongdokoship<br />
River in Yongjeon-ri village; 36°23'47"N<br />
129°15'20"E; holotype: CNUC 21827; adjective, -is,<br />
-is, -e)<br />
4.7 Kichulchoia Kim, Park & Nalbant, 1999<br />
Choia Kim, Park & Nalbant, 1997: 192 (type species: Niiwaella<br />
brevifasciata Kim & Lee, 1995: 285, by original<br />
designation; junior homonym <strong>of</strong> Choia Walcott, 1920:<br />
291 in Spongiae). Gender feminine.<br />
Kichulchoia Kim, Park & Nalbant, 1999: 374 (replacement<br />
name for Choia Kim, Park & Nalbant, 1997: 192). Gender<br />
feminine.<br />
4.7.1 Kichulchoia brevifasciata (Kim & Lee, 1995)<br />
Niwaella brevifasciata Kim & Lee, 1995: 285, figs. 1–2 (type<br />
locality: South Korea: Chollanam-do province: Kohûnggun<br />
district: Pungyang-myon municipality: Koûp stream<br />
in Yamag-ri village, 34°34'06"N 127°14'16"E; holotype:<br />
CNUC 19907; adjective, -us, -a, -um)<br />
4.7.2 Kichulchoia multifasciata (Wakiya & Mori, 1929)<br />
Cobitis multifasciatus Wakiya & Mori, 1929: 31, pl. 2 fig. 1<br />
(type locality: Korea: Rakuto River at Taikyu; holotype:<br />
LU, Böhlke, 1953: 39; adjective, -us, -a, -um)<br />
Taxonomic notes. Generic position follows Šlechtová et al.<br />
(2008) and Kim (2009).<br />
4.8 Koreocobitis Kim, Park & Nalbant, 1997<br />
Koreocobitis Kim, Park & Nalbant, 1997: 191 (type species:<br />
Cobitis rotundicaudata Wakiya & Mori, 1929: 32,<br />
by original designation). Gender feminine.<br />
4.8.1 Koreocobitis naktongensis Kim, Park & Nalbant, 2000<br />
Koreocobitis naktongensis Kim, Park & Nalbant, 2000: 90,<br />
figs. 1a, 2a (type locality: Korea: Chollabuk-do province:<br />
Namwon-gun district: Inwol-myon municipality:<br />
Naktong River at Jugkun-ri village; holotype: CNUC<br />
21229; adjective, -is, -is, -e)<br />
4.8.2 Koreocobitis rotundicaudata (Wakiya & Mori, 1929)<br />
Cobitis rotundicaudata Wakiya & Mori, 1929: 32, pl. 2 fig. 2<br />
(type locality: Korea: Chung-chonbuk-do province: Dangyang-gun<br />
district: Yonsangchon-myon municipality:<br />
Youm Hyon stream, tributary <strong>of</strong> South Han River, at Shingok-ri<br />
village [original type locality: Korea: Soun Kan<br />
River at Tanyo]; neotype: CNUC uncat., designated by<br />
Kim, Park & Nalbant, 1997: 192 but questionable validity<br />
since reasons to believe that the original type materi-<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
31<br />
4.9.3 Kottelatlimia pristes, CMK 11116, 41.9 mm SL; Indonesia:<br />
Sumatra: Batang Hari drainage.<br />
al is lost are not explained, Code art. 75.3.4; adjective, -<br />
us, -a, -um)<br />
4.9 Kottelatlimia Nalbant, 1994<br />
Kottelatlimia Nalbant, 1994: 377 (type species: Lepidocephalichthys<br />
katik Kottelat & Lim, 1992: 212, by original<br />
designation). Gender feminine.<br />
4.9.1 Kottelatlimia hipporhynchos Kottelat & Tan, 2008<br />
Kottelatlimia hipporhynchos Kottelat & Tan, 2008: 64, fig. 1<br />
(type locality: Indonesia: Borneo: Kalimantan Tengah:<br />
Kahayan drainage, blackwater stream at km 80 on road<br />
from Palangka Raya to Tumbang Telakian (35 km after<br />
turn-<strong>of</strong>f at km 45 on road from Palangka Raya to Kasongan);<br />
1°37.324'S 113°37.569'E; holotype: MZB 10977;<br />
compound noun, indeclinable)<br />
4.9.2 Kottelatlimia katik (Kottelat & Lim, 1992)<br />
Lepidocephalichthys katik Kottelat & Lim, 1992: 212, fig. 6<br />
(type locality: Malaysia; holotype: ZRC 9344; noun in<br />
apposition, indeclinable)<br />
4.9.3 Kottelatlimia pristes (Roberts, 1989)<br />
Lepidocephalichthys pristes Roberts, 1989: 105, fig. 80 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: 30 km<br />
west <strong>of</strong> Sintang on road from Sanggau to Sintang; holotype:<br />
MZB 3527; noun in apposition, indeclinable)<br />
4.10 Lepidocephalichthys Bleeker, 1863<br />
Cobitichthys Bleeker, 1858: 304 (type species: Cobitis barbatuloides<br />
Bleeker, 1851: 435, by monotypy; declared<br />
nomen oblitum under Code art. 23.9.2 by Kottelat & Tan,<br />
2008b: 70). Gender masculine.<br />
Lepidocephalichthys Bleeker, 1863a: 38 (type species: Cobitis<br />
hasselti Valenciennes, in Cuvier & Valenciennes,<br />
1846: 74, by original designation; also in Bleeker, 1863c:<br />
4, pl. 103; declared nomen protectum under Code art.<br />
23.9.2 by Kottelat & Tan, 2008b: 70). Gender masculine.<br />
Platacanthus Day, 1865a: 296 (type species: Platacanthus<br />
agrensis Day, 1865a: 296, by monotypy; junior homonym<br />
<strong>of</strong> Platacanthus Fischer von Waldheim, 1849: 80, in fossil<br />
fishes; also in Day, 1865b: 204). Gender masculine.<br />
Jerdonia Day, 1871: 700 (type species: Platacanthus maculatus<br />
Day, 1868: 941, by monotypy; junior homonym <strong>of</strong><br />
Jerdonia Blanford & Blanford, 1862: 348, in Mollusca).<br />
Gender feminine.<br />
Enobarbus Whitley, 1928: 296 (replacement name for Jerdonia<br />
Day, 1871: 700; not a junior homonym <strong>of</strong> Aenobarbus<br />
Agassiz, 1842: 2, in Aves). Gender masculine.
Enobarbichthys Whitley, 1931: 107 (unnecessary replacement<br />
name for Enobarbus Whitley, 1928: 296). Gender<br />
masculine.<br />
Madrasia Nalbant, 1963: 364 (replacement name for Jerdonia<br />
Day, 1871: 700). Gender feminine.<br />
Nomenclatural notes. Whitley (1931: 107) proposed<br />
Enobarbichthys as a replacement name for Enobarbus which<br />
he considered preoccupied by "Aenobarbus Temminck,<br />
1835". As the spellings <strong>of</strong> the two names differ, they are not<br />
homonyms and Enobarbus is available.<br />
Whitley could not trace the original description <strong>of</strong><br />
Aenobarbus. The reference given by Neave (1939: 73) is<br />
Agassiz (1842: 2) but in Agassiz (1842), the name is listed<br />
simply as "Aenobarbus Temm. Pl. Col. d'Ois. II, 1838. Nom.<br />
propr. (aenas; barba.) Columbinae." This refers to Temminck<br />
& Laugier de Chartrouse (1836), which I have examined.<br />
This is an unpaginated collection <strong>of</strong> plates and text and the<br />
name Aenobarbus does not appear in volume 2. Volume 1<br />
has a 109-pages "Tableau méthodique" (1839) listing all<br />
birds, including those in the work, and I could not find<br />
Aenobarbus. The word aenobarbus only appears as Allotrius<br />
aenobarbus on plate 589 (dated 1836 by Dickinson, 2001:<br />
46). The genus name Aenobarbus is made available by its<br />
mention by Agassiz (1842: 2), in which "nom. prop." indicates<br />
it is a proper noun, which means that the genus name is<br />
based on an existing noun, which can only refer to Allotrius<br />
aenobarbus. In conclusion, the author <strong>of</strong> Aenobarbus is Agassiz<br />
(1842: 2) and it is not a senior synonym <strong>of</strong> Enobarbus.<br />
4.10.1 Lepidocephalichthys alkaia Havird & Page, 2010<br />
Lepidocephalichthys alkaia Havird & Page, 2010: 140, fig. 4<br />
(type locality: Myanmar: Kachin State: Hpa-Lap stream,<br />
<strong>of</strong> Nam Chim Chaung, <strong>of</strong> Nan Kwe Chaung, northwest<br />
<strong>of</strong> Myitkyina, 25°31'36"N 97°22'45"E; holotype: USNM<br />
372169; noun in apposition, indeclinable)<br />
4.10.2 Lepidocephalichthys annandalei Chaudhuri, 1912<br />
Lepidocephalichthys annandalei Chaudhuri, 1912: 442,<br />
pl. 40 fig. 3 (type locality: India: Mahananda River at<br />
Siliguri and Tista River near Jalgaipuri; syntypes [total<br />
7]: ZSI F 5593–5595/1 [3], Menon & Yazdani, 1968:<br />
121; noun in genitive, indeclinable)<br />
Lepidocephalichthys menoni Pillai & Yazdani, 1976: 13,<br />
fig. 1 (type locality: India: Meghalaya: South Garo Hills<br />
District: Someswari River at Baghmara [22°15'N<br />
88°52'E]; holotype: ZSI/ETS V/ERS 551; noun in genitive,<br />
indeclinable)<br />
4.10.3 ? Lepidocephalichthys arunachalensis (Datta &<br />
Barman, 1984)<br />
Noemacheilus arunachalensis Datta & Barman, 1984: 275,<br />
pl. 14 (type locality: India: Arunachal Pradesh: Tirap<br />
District: Namdapha River, Namdapha Wild Life Sanctuary;<br />
holotype: ZSI FF 1713; adjective, -is, -is, -e)<br />
Taxonomic notes. Apparently a species <strong>of</strong> Lepidocephalichthys<br />
(see Kottelat, 1990a: 118). Validity unclear.<br />
4.10.4 Lepidocephalichthys barbatuloides (Bleeker, 1851)<br />
Cobitis barbatuloïdes Bleeker, 1851: 435 (type locality: Indonesia:<br />
Borneo: Kalimantan Barat: Sambas; holotype:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
32<br />
RMNH 4960, Alfred, 1961: 33; adjective, indeclinable)<br />
Taxonomic notes. Tentatively recognised as valid (Kottelat<br />
& Tan, 2008b: 70). Not recorded since original description.<br />
4.10.5 Lepidocephalichthys berdmorei (Blyth, 1860)<br />
Acanthopis [sic] Berdmorei Blyth, 1860: 168 (type locality:<br />
Burma: Tenasserim provinces; holotype: ZSI F 2646/1,<br />
Kottelat & Lim, 1992: 205 [contra statement in Eschmeyer<br />
& Fricke, 2010, a single locality is stated in original<br />
description]; noun in genitive, indeclinable)<br />
Lepidocephalus cataractus Fowler, 1939: 60, fig. 10 (type<br />
locality: Thailand: waterfall at Trang; holotype: ANSP<br />
68470; treated as a noun in apposition, indeclinable<br />
[should have been cataracta])<br />
Lepidocephalus guntea birmanicus Rendahl, 1948: 64,<br />
fig. 30 (original description; type locality: Myanmar:<br />
Shweli Kyaung, 24 miles east <strong>of</strong> 24°N 96°E; holotype:<br />
NRM 20829; adjective, -us, -a, -um)<br />
? Acanthophthalmus longipinnis Menon, 1992: 93, fig. 10<br />
(type locality: India: Manipur: Kharungpat Lake, 20 km<br />
south <strong>of</strong> Imphal; holotype: ZSI/SRS 3371; compound<br />
noun, indeclinable)<br />
4.10.6 Lepidocephalichthys coromandelensis Menon, 1992<br />
Lepidocephalichthys coromandelensis Menon, 1992: 67,<br />
fig. 4, pl. 6 fig. 1 (type locality: India: Andhra Pradesh:<br />
Araku Valley; holotype: ZSI/SRS F 3369; adjective, -is,<br />
-is,-e)<br />
4.10.7 Lepidocephalichthys furcatus (de Beaufort, 1933)<br />
Lepidocephalus furcatus de Beaufort, 1933: 32 (type locality:<br />
Malaysia: Perak: Bukit Merah Reservoir; syntypes<br />
[6]: ZRC 1445 [3], ZMA 100.979 [1], BMNH<br />
1969.11.13.1 [1], Kottelat & Lim, 1992: 207, Nijssen et<br />
al., 1993: 214, Eschmeyer & Fricke, 2010; adjective,<br />
-us, -a, -um)<br />
4.10.8 Lepidocephalichthys goalparensis Pillai & Yazdani,<br />
1976<br />
Lepidocephalichthys goalparensis Pillai & Yazdani, 1976:<br />
15, fig. 2 (type locality: India: Assam: 6 km west <strong>of</strong> Dudhna<br />
Inspection Bungalow, Goalpara; holotype: ZSI/ERS<br />
V/ERS 519; adjective, -is, -is, -e)<br />
Lepidocephalus caud<strong>of</strong>urcatus Tilak & Husain, 1979: 60,<br />
fig. 2 (type locality: India: Uttar Pradesh: Dehra Dun<br />
District: Kalapani nala Rishikesh; holotype: ZSI/NRS<br />
V-1186; adjective, -us, -a, -um)<br />
4.10.9 Lepidocephalichthys guntea (Hamilton, 1822)<br />
Cobitis guntea Hamilton, 1822: 353, 394 (type locality:<br />
Bengal [Goalpara; Hora, 1935a: 49]; types: NT; Hamilton's<br />
unpublished figure reproduced in M'Clelland, 1839:<br />
pl. 51 fig. 3; noun in apposition, indeclinable)<br />
Cobitis balgara Hamilton, 1822: 356, 394 (type locality:<br />
India: Kosi River [at Nathpur, near the Nepal frontier;<br />
Hora, 1935a: 49]; types: NT; Hamilton's unpublished figure<br />
reproduced in M'Clelland, 1839: pl. 53 fig. 2; simultaneous<br />
subjective synonym <strong>of</strong> Cobitis guntea Hamilton,<br />
1822: 353; first reviser [Day, 1878a: 609] gave precedence<br />
to C. guntea; noun in apposition, indeclinable)
4.10.5 Lepidocephalichthys berdmorei, CMK 22043, 62.8 mm SL;<br />
Thailand: Ranong.<br />
4.10.7 Lepidocephalichthys furcatus, CMK 16516, 21.5 mm SL;<br />
Thailand: Tapi drainage.<br />
Cobitis Maya Sykes, 1839a: 162 (type locality: India: Deccan<br />
[Mola Mola River at Poona; Sykes, 1841: 367 [Pune,<br />
18°28'N 73°48'E]]; syntypes: LU; also in Sykes, 1839b:<br />
59, 1841: 367; noun in apposition, indeclinable)<br />
Canthophrys vittatus Swainson, 1839: 310 (available by indication<br />
to Hamilton, 1822 "Cob. No. 4" [p. 353, Cobitis<br />
guntea]; type locality: Bengal [Goalpara; Hora, 1935a:<br />
49]; types: NT; Hamilton's unpublished figure reproduced<br />
in M'Clelland, 1839: pl. 51 fig. 3; adjective, -us, -a, -um)<br />
Cantophrys olivaceus Swainson, 1839: 310 (available by<br />
indication to Hamilton, 1822 "Cob. No. 8" [p. 356, Cobitis<br />
balgara]; type locality: India: Kosi River [at Nathpur,<br />
near the Nepal frontier; Hora, 1935a: 49]; types: NT;<br />
Hamilton's unpublished figure reproduced in M'Clelland,<br />
1839: pl. 53 fig. 2; adjective, -us, -a, -um)<br />
? Cobitis guttata M'Clelland, 1839: 305, 438, pl. 52 figs. 5–<br />
6 (type locality: India: tanks in vicinity <strong>of</strong> Joorhath; syntypes:<br />
SMF 409 [1], Eschmeyer & Fricke, 2010 [as holotype];<br />
adjective, -us, -a, -um)<br />
Cobitis phoxocheila M'Clelland, 1839: 305, 439, pl. 52 fig. 4<br />
(type locality: India: Arunachal Pradesh: Mishmee<br />
moutains; syntypes: ZSI [2]; compound adjective, -us,<br />
-a, -um)<br />
? Schistura aculeata M'Clelland, 1839: 307 (type locality:<br />
India: Assam; types: LU; adjective, -us, -a, -um)<br />
Misgurnus lateralis Günther, 1868: 346 (type locality: India:<br />
Bengal; holotype: BMNH 1858.8.15.49, Eschmeyer<br />
& Fricke, 2010; adjective, -is, -is, -e)<br />
Lepidocephalus dibruensis Sen, 1979: 35, fig. 1 (type locality:<br />
India: Assam: Dibru River, Guijan, 60 km from Dibrugarh;<br />
holotype: ZSI FF 1203; adjective, -is, -is, -e)<br />
Lepidoce-phalichthys nepalensis Shrestha, 1978: 37 (nomen<br />
nudum)<br />
Lepidocephalichthys nepalensis Shrestha, 1981: 129, fig. 63<br />
(type locality: Nepal: Biratnagar: Singhia River; holotype:<br />
TUK; adjective, -is, -is, -e)<br />
4.10.10 Lepidocephalichthys hasselti (Valenciennes, in<br />
Cuvier & Valenciennes, 1846)<br />
Cobitis Octocirrhus Kuhl & van Hasselt, in van Hasselt,<br />
1823: 133, 1824: 376 (nomen nudum, see Kottelat,<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
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4.10.12 Lepidocephalichthys jonklaasi, CMK 7117, 53.7 mm SL;<br />
Sri Lanka: Wilpita.<br />
4.10.14 Lepidocephalichthys lorentzi, CMK 6967, 27.5 mm SL;<br />
Indonesia: Borneo: Kapuas drainage.<br />
4.11.1 Lepidocephalus cf. macrochir, about 60 mm SL; Thailand:<br />
Chao Phraya drainage. (Photograph by Jörg Bohlen).<br />
1987b: 371)<br />
Cobitis hasselti Valenciennes, in Cuvier & Valenciennes,<br />
1846: 74 (type locality: Indonesia: Java: Tjelankakan<br />
River [Tjelankahan; Kottelat & Lim, 1992: 210]; holotype:<br />
LU; based on a drawing sent by Kuhl and van Hasselt<br />
[reproduced in Roberts, 1993: fig. 27]; noun in genitive,<br />
indeclinable)<br />
Lepidocephalichthys nudus Machan, 1931: 221 (type locality:<br />
Indonesia: Java: Kalen reservoir near Surabaya; syntypes:<br />
NMW 16151–16156 [6], Kottelat & Lim, 1992:<br />
211; adjective, -us, -a, -um)<br />
Lepidocephalus taeniatus Fowler, 1939: 63, figs. 11–12 (type<br />
locality: Thailand: waterfall at Trang; holotype: ANSP<br />
68486; adjective, -us, -a, -um)<br />
Acanthophthalmus unistriatus Roberts, 1993: 25 (not available,<br />
name published in synonymy)<br />
4.10.11 Lepidocephalichthys irrorata Hora, 1921<br />
Lepidocephalichthys irrorata Hora, 1921b: 196, pl. 9 fig. 5<br />
(type locality: India: Manipur [exact locality not explicitly<br />
stated, probably Loktak Lake]; holotype: ZSI F 9904/<br />
1; participle, -us, -a, -um)<br />
4.10.12 Lepidocephalichthys jonklaasi (Deraniyagala,<br />
1956)<br />
Lepidocephalus jonklaasi Deraniyagala, 1956: 35 (type<br />
locality: Sri Lanka: Southern Province: Akuressa; holotype:<br />
NMSL FF 727 A; noun in genitive, indeclinable)<br />
4.10.13 Lepidocephalichthys kranos Havird & Page, 2010<br />
Lepidocephalichthys kranos Havird & Page, 2010: 149,<br />
fig. 8 (type locality: Thailand: Ubon Ratchathani: Mun<br />
River (tributary <strong>of</strong> Mekong River), marsh near University<br />
Ubon Ratchathani campus, 15°10'45.84"N
104°45'44.76"E; holotype: UF 171980; noun in apposition,<br />
indeclinable)<br />
4.10.14 Lepidocephalichthys lorentzi (Weber & de Beaufort,<br />
1916)<br />
Acanthophthalmus lorentzi Weber & de Beaufort, 1916: 32,<br />
fig. 12 (type locality: Indonesia: Borneo: Kalimantan<br />
Barat: Upper Kapuas [at Putussibau; Roberts, 1989: 104];<br />
holotype: ZMA 103.259, Nijssen et al., 1993: 213; noun<br />
in genitive, indeclinable)<br />
4.10.15 Lepidocephalichthys micropogon (Blyth, 1860)<br />
Acanthopis [sic] micropogon Blyth, 1860: 168 (type locality:<br />
Burma: Tenasserim provinces; types: lost, Kottelat &<br />
Lim, 1992: 209; compound noun, indeclinable)<br />
Lepidocephalichthys manipurensis Arunkumar, 2000b: 1097,<br />
fig. 4 (type locality: India: Manipur: Chindwin drainage:<br />
Chandel District: Lairok Maru, tributary <strong>of</strong> Lokchao River<br />
near Moreh; holotype: MUMF2250/1A; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. The figure <strong>of</strong> the mouth <strong>of</strong> L. manipurensis<br />
in the original description (Arunkumar, 2000b: 1099,<br />
fig. 5a) is a crude copy <strong>of</strong> the drawing <strong>of</strong> the mouth <strong>of</strong> Kottelatlimia<br />
katik in Kottelat & Lim (1992: 213, fig. 7c). The<br />
figure <strong>of</strong> the pectoral fin (fig. 3d) is based on that <strong>of</strong> K. katik<br />
in Kottelat & Lim (1992: fig. 7) in which the serrae have<br />
been removed, a second simple ray has been added (unlikely<br />
in Cobitidae and contradicting the text) and a structure<br />
added on branched ray 5 (not 6–7 as in text, and in all other<br />
species <strong>of</strong> the genus).<br />
4.10.16 Lepidocephalichthys thermalis (Valenciennes, in<br />
Cuvier & Valenciennes, 1846)<br />
Cobitis thermalis Valenciennes, in Cuvier & Valenciennes,<br />
1846: 78 (type locality: Sri Lanka: hot waters <strong>of</strong> Cania;<br />
syntypes: MNHN 1082 [7], Bertin & Estève, 1948: 93;<br />
adjective, -is, -is, -e)<br />
Cobitis Carnaticus Jerdon, 1849: 331 (type locality: India:<br />
Carnatic; types: NT; adjective, -us, -a, -um)<br />
Cobitis Mysorensis Jerdon, 1849: 332 (type locality: India:<br />
Mysore; syntypes: NT; adjective, -is, -is, -e)<br />
Cobitis rubripinnis Jerdon, 1849: 332 (type locality: India:<br />
Malabar; syntypes: NT; junior homonym <strong>of</strong> Cobitis rubripinnis<br />
Temminck & Schlegel, 1846: 222, pl. 103 fig. 1;<br />
compound noun, indeclinable)<br />
Platacanthus agrensis Day, 1865a: 296 (type locality: India:<br />
Kerala: Trichoo near Cochin; holotype: BMNH<br />
1865.7.17.25 [1], 1867.5.10.17–19 [3] or 1975.9.30.12<br />
[1], Whitehead & Talwar, 1976: 156, Eschmeyer &<br />
Fricke, 2010; also in Day, 1865b: 204, pl. 14 fig. 1; adjective,<br />
-is, -is, -e)<br />
Platacanthus maculatus Day, 1868: 941 (type locality: India:<br />
Madras; holotype: BMNH 1868.10.28:36, Harant<br />
& Bohlen, 2010: 2446, fig. 1a, Whitehead & Talwar,<br />
1976: 157, Menon, 1992: 85; adjective, -us, -a, -um)<br />
4.10.17 Lepidocephalichthys tomaculum Kottelat & Lim,<br />
1992<br />
Lepidocephalichthys tomaculum Kottelat & Lim, 1992: 216,<br />
fig. 9 (type locality: Malaysia: Selangor: Sungai Bernam<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
34<br />
basin: North Selangor peat swamp forest, stream at 34<br />
km mark on road from Sungai Besar to Tanjong Malim;<br />
approx. 3°40'N 101°20'E; holotype: ZRC 14938; noun<br />
in apposition, indeclinable)<br />
4.10.18 Lepidocephalichthys zeppelini Havird & Tangjitjaroen,<br />
in Havird, Page, Tangjitjaroen, Vidthayanon,<br />
Grudpan & Udduang, 2010<br />
Lepidocephalichthys zeppelini Havird & Tangjitjaroen, in<br />
Havird, Page, Tangjitjaroen, Vidthayanon, Grudpan &<br />
Udduang, 2010: 6, fig. 1 (type locality: Thailand: Ubon<br />
Ratchathani Province: Mun River (Mekong drainage),<br />
isolated pools in rice field, Ubon Rajathani University<br />
campus, 15°08'03.18"N 104°55'27.78"E; holotype: UF<br />
174131; noun in genitive, indeclinable)<br />
4.11 Lepidocephalus Bleeker, 1858<br />
Lepidocephalus Bleeker, 1858: 303 (type species: Cobitis<br />
macrochir Bleeker, 1854b: 97, by subsequent designation<br />
by Bleeker, 1863a: 38, 1863c: 4). Gender masculine.<br />
4.11.1 Lepidocephalus macrochir (Bleeker, 1854)<br />
Cobitis macrochir Bleeker, 1854b: 97 (type locality: Indonesia:<br />
Sumatra: Palembang: confluence <strong>of</strong> Lamatang<br />
[Lematang] and Enim Rivers / Central Java: Surakarta,<br />
in Pepeh River; syntypes [5, 64–91 mm TL]: RMNH,<br />
BMNH 1866.5.2.55 [1], Eschmeyer & Fricke, 2010; compound<br />
noun, indeclinable)<br />
Lepidocephalichthys pallens Vaillant, 1902: 153, fig. 47 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Kapuas<br />
River, possibly at Sintang; holotype: RMNH 7783, Eschmeyer<br />
& Fricke, 2010; adjective, indeclinable)<br />
Acanthophthalmus pahangensis de Beaufort, 1933: 31 (type<br />
locality: Malaysia: 'fish-drive' <strong>of</strong>f Mentakab, Pahang River;<br />
holotype: ZRC 490, Alfred, 1970: 70; adjective, -is,<br />
-is, -e)<br />
4.11.2 Lepidocephalus spectrum Roberts, 1989<br />
Lepidocephalus spectrum Roberts, 1989: 106, fig. 81 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Sungai<br />
Melawi near confluence with Kapuas, about 0.5 km upstream<br />
from Sintang; holotype: MZB 3533; noun in apposition,<br />
indeclinable)<br />
4.12 Microcobitis Bohlen & Harant, 2011<br />
Microcobitis Bohlen & Harant, 2011: 296 (type species:<br />
Cobitis misgurnoides Rendahl, 1944: 21, by original<br />
designation). Gender feminine.<br />
4.12.1 Microcobitis misgurnoides (Rendahl, 1944)<br />
Cobitis misgurnoides Rendahl, 1944: 21, fig. 10 (type locality:<br />
Vietnam: Annam: Thua Luu, 50 km south-east <strong>of</strong> Hué;<br />
holotype: NHMG; adjective, indeclinable)<br />
? Cobitis nuicocensis Nguyen & Vo, in Nguyen, 2005: 560,<br />
fig. 9 (type locality: Vietnam: Thai Nguyen Province: Dai<br />
Tu district: Cong River, flowing to Nui Coc lake; holotype:<br />
NCNTTSI; adjective, -is, -is, -e)
4.13 Misgurnus La Cepède, 1803<br />
Misgurnus La Cepède, 1803: 16 (type species: Cobitis fossilis<br />
Linnaeus, 1758: 303, by monotypy). Gender masculine.<br />
Ussuria Nikolski, 1904: 362 (type species: Ussuria leptocephala<br />
Nikolski, 1904: 362, by monotypy). Gender feminine.<br />
Mesomisgurnus Fang, 1935b: 129 (type species: Nemacheilus<br />
bipartitus Sauvage & Dabry de Thiersant, 1874: 16,<br />
by original designation). Gender masculine.<br />
Taxonomic notes. Several chromosome and molecular studies<br />
(e.g. Šlechtová et al., 2008; Kitagawa et al., 2011; Perdices<br />
et al., 2012) show that the genus Misgurnus is polyphyletic.<br />
4.13.1 Misgurnus anguillicaudatus (Cantor, 1842)<br />
Cobitis anguillicaudata Cantor, 1842: 485 (type locality:<br />
China: Chusan Island; syntypes: BMNH 2007.9.17.2 [1],<br />
2007.9.17.3–10 [8], Günther, 1868: 345; adjective, -us,<br />
-a, -um)<br />
Cobitis bifurcata M'Clelland, 1844: 400, pl. 23 fig. 1 (type<br />
locality: China: Chusan Island; types: LU; adjective,<br />
-us, -a, -um)<br />
Cobitis pectoralis M'Clelland, 1844: 400, pl. 23 fig. 3 (type<br />
locality: China: Chusan Island; types: LU; noun in apposition,<br />
indeclinable)<br />
Cobitis psammismus Richardson, 1846a: 300 (type locality:<br />
China: Canton; holotype: specimen on which is based<br />
Reeves's unpublished drawing, reproduced in Whitehead,<br />
1970: 210, pl. 19 fig. a; noun in apposition, indeclinable)<br />
Cobitis haematopterus Richardson, 1846a: 301 (type locality:<br />
Japan; holotype: ? BMNH; compound adjective,<br />
-us, -a, -um)<br />
Cobitis rubripinnis Temminck & Schlegel, 1846: 220, pl. 103<br />
fig. 1 (type locality: Japan: surroundings <strong>of</strong> Nagasaki;<br />
lectotype: RMNH 2705b, designated by Boeseman, 1947:<br />
166; compound noun, indeclinable)<br />
Cobitis maculata Temminck & Schlegel, 1846: 221, pl. 103<br />
fig. 2 (type locality: Japan [surroundings <strong>of</strong> Nagasaki];<br />
holotype: ? RMNH 2705f, Boeseman, 1947: 166 [Boeseman<br />
commented that the specimen on the plate is not the<br />
specimen described in the text and that RMNH 2705f is<br />
very similar to the plate; if the plate appeared before the<br />
text and if the plate is really based on this specimen, it is<br />
the holotype; if the text and plate appeared together, it is<br />
possibly a syntype]; adjective, -us, -a, -um)<br />
Cobitis micropus Valenciennes, in Cuvier & Valenciennes,<br />
1846: 29 (type locality; China; holotype: MNHN 5688,<br />
Bertin & Estève, 1948: 95; compound noun, indeclinable)<br />
Cobitis decemcirrosus Basilewsky, 1855: 239, pl. 7 fig. 2<br />
(type locality: China: stagnant waters near Beijing; types:<br />
ZISP ?; adjective, -us, -a, -um)<br />
Cobitichthys enalios Bleeker, 1859b: 259 (nomen nudum)<br />
Cobitichthys dichachrous Bleeker, 1859b: 259 (nomen nudum)<br />
Cobitichthys polynema Bleeker, 1859b: 259 (nomen nudum)<br />
Cobitichthys enalios Bleeker, 1860b: 88, pl. 2 fig. 4 (type<br />
locality: Japan: Kaminoseki; holotype: BMNH<br />
1862.5.2.26 [1], Günther, 1868: 346; treated as noun in<br />
apposition, indeclinable)<br />
Cobitichthys dichachrous Bleeker, 1860b: 89, pl. 2 fig. 2<br />
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35<br />
(type locality: Japan: Jedo [Tokyo]; holotype: BMNH<br />
1866.5.2.96 [1], Günther, 1868: 347; etymology unknown,<br />
indeclinable)<br />
Cobitichthys polynema Bleeker, 1860b: 90, pl. 2 fig. 3 (type<br />
locality: Japan: Jedo [Tokyo]; holotype: BMNH<br />
1866.5.2.24, Günther, 1868: 346; compound noun, indeclinable)<br />
Cobitis erythropterus Günther, 1868: 345 (not available,<br />
unpublished name in collection)<br />
Misgurnus maculatus Bleeker, 1872: 146 (nomen nudum;<br />
locality: China)<br />
Cobitis He-tsieou Dabry de Thiersant, 1872: 191, pl. 49 fig. 1<br />
(name not binominal, not available; locality: Central<br />
China)<br />
? Cobitis cirrhifurcata Dabry de Thiersant, 1872: 191, pl. 49<br />
fig. 2 (type locality: China: lakes <strong>of</strong> the inner <strong>of</strong> China,<br />
Yang-tsee-kiang [Yangtze]; holotype: NT; adjective, -us,<br />
-a, -um)<br />
Cobitis ny-tsieou Dabry de Thiersant, 1872: 191, pl. 49 fig. 3<br />
(name not binominal, not available; China: Yang-tseekiang<br />
[Yangtze], lakes and ponds <strong>of</strong> China)<br />
Nemachilus lividus Sauvage & Dabry de Thiersant, 1874:<br />
15 (type locality: China; syntypes: MNHN 5237 [2],<br />
Fang, 1935b: 139, fig. 9, Bertin & Estève, 1948: 95; adjective,<br />
-us, -a, -um)<br />
Misgurnus crossochilus Sauvage, 1878a: 89 (type locality:<br />
China: Fujian: high mountains <strong>of</strong> Koaten [Fujian: Kuatun,<br />
27°51'N 117°48'E; see Crossostoma davidi]; syntypes:<br />
MNHN 9822 [4], Bertin & Estève, 1948: 95; compound<br />
adjective, -us, -a, -um)<br />
Misgurnus anguillicaudatus tungting Nichols, 1925f: 3 (type<br />
locality: China: Hunan: Huping, Tungting Lake; holotype:<br />
AMNH 8393; noun in apposition, indeclinable)<br />
Misgurnus mizolepis fukien Nichols, 1925f: 4 (type locality:<br />
China: Fukien [Fujian]: Yenping [Yanping]; holotype:<br />
AMNH 8394; noun in apposition, indeclinable)<br />
Misgurnus mizolepis hainan Nichols & Pope, in Nichols,<br />
1925f: 4 (type locality: China: Hainan: Nodoa; holotype:<br />
AMNH 8363; also in Nichols & Pope, 1927: 337, fig. 9,<br />
pl. 16 fig. 1; noun in apposition, indeclinable)<br />
Misgurnus mizolepis grangeri Nichols, 1925f: 5 (type locality:<br />
China: Sichuan: Yen-ching-kao; holotype: AMNH<br />
8395; noun in genitive, indeclinable)<br />
Misgurnus mohoity yunnan Nichols, 1925f: 5 (type locality:<br />
China: Yunnan: Yunnanfu [Kunming]; holotype:<br />
AMNH 8396; noun in apposition, indeclinable)<br />
Misgurnus mohoity leopardus Nichols, 1925f: 6 (type locality:<br />
China: Hunan: Tungting Lake; holotype: AMNH<br />
8397; noun in apposition, indeclinable)<br />
Misgurnus punctatus Oshima, 1926: 5 (type locality: China:<br />
Hainan: Kachek River near Kachek; holotype: LU; adjective,<br />
-us, -a, -um)<br />
Misgurnus anguillicaudatus formosanus Rendahl, 1936: 302<br />
(type locality: Taiwan: Lake Candidius; holotype: NRM<br />
10354; adjective, -us, -a, -um)<br />
Misgurnus mizolepis heungchow Lin, 1932a: 66 (type locality:<br />
China: Guangdong: Heungchow; syntypes: LU; noun<br />
in apposition, indeclinable)<br />
Misgurnus mizolepis unicolor Lin, 1932a: 66 (type locality:<br />
China: Guangdong: East Fisheries Experiment Station
4.12.1 Microcobitis cf. misgurnoides, CMK 19846, 31.1 mm SL;<br />
Laos: Mekong drainage: Nakai reservoir.<br />
4.13.1 Misgurnus anguillicaudatus, CMK 14918, 81.3 mm SL;<br />
Vietnam: Mong Cai.<br />
Canton; holotype: LU; adjective, indeclinable)<br />
Misgurnus elongatus Kimura, 1934: 158, pl. 5 fig. 1 (type<br />
locality: China: Shanghai; holotype: LU; adjective, -us,<br />
-a, -um)<br />
Taxonomic notes. Synonymy largely follows Chen (1981)<br />
but needs re-evaluation. Chromosome and genetic data (e.g.<br />
Yu et al., 1989: 160; Dong et al., 1999; Shimizu & Takagi,<br />
2010; Kitagawa et al., 2011; Perdices et al., 2012) suggest<br />
that several species are confused under the name M. anguillicaudatus.<br />
See also comments under Paramisgurnus dabryanus.<br />
4.13.2 Misgurnus buphoensis Kim & Pak, 1995<br />
Misgurnus buphoensis Kim & Pak, 1995: 54, fig. (type locality:<br />
North Korea: North Hamgyong province: Sonbong<br />
county: Bupori; holotype: MIZASDPRK 93-65; adjective,<br />
-is, -is, -e)<br />
4.13.3 Misgurnus fossilis (Linnaeus, 1758)<br />
Cobitis fossilis Linnaeus, 1758: 303 (based on J. F. Gronovius<br />
[1748: 79, pl. 3, Cobitis aculeo bifurco ...], Artedi [1738:<br />
gen. [spec.] 2, syn. 3, Cobitis caerulescens ...] and Linnaeus<br />
[1754: 76, idem]; type locality: "in Europa"; syntypes:<br />
BMNH 1853.11.12:121 [1], NRM 69 [1], Wheeler,<br />
1958: 213, Fernholm & Wheeler, 1983: 218; adjective,<br />
-is, -is, -e)<br />
? Petromizon [sic] variegatus Wulff, 1765: 16 (type locality:<br />
Borussia [Prussia]; type material: NT; adjective, -us,<br />
-a, -um)<br />
4.13.4 Misgurnus mohoity (Dybowski, 1869)<br />
Cobitis fossilis var. mohoity Dybowski, 1869: 957 (type locality:<br />
Russia: Siberia: Zabaykalsky Krai: muddy lakes <strong>of</strong><br />
Onon and Ingoda drainages [near Duldurga, on Ila River;<br />
Vasil'eva, 2001: 553]; syntypes: NMW 80070 [4],<br />
83462 [1], Eschmeyer & Fricke, 2010; noun in genitive,<br />
indeclinable)<br />
Nemachilus bipartitus Sauvage & Dabry de Thiersant, 1874:<br />
16 (type locality: North China and Central China [Sichuan];<br />
syntypes: MNHN 3931 [2], 4655 [9], Fang,<br />
1935b: 136, fig. 7, Bertin & Estève, 1948: 95; adjective,<br />
-us, -a, -um)<br />
? Misgurnus maculatus Bleeker, 1872: 146 (nomen nudum;<br />
locality: China)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
36<br />
4.13.4 Misgurnus mohoity, CMK 19646, 103.4 mm SL; Mongolia:<br />
Amur drainage: Bayan Gol.<br />
4.13.7 Misgurnus tonkinensis, 82.5 mm SL; Vietnam: Vinh.<br />
? Misgurnus spilurus Bleeker, 1872: 146 (nomen nudum;<br />
locality: China)<br />
Misgurnus cestoideus Kessler, 1876: 34 (type locality: China:<br />
Nei Mongol: Lake Dalai-Nor [Dalai Nur; 43°18'N<br />
116°40'E; not Hu-Lun Lake, which is another Dalai Nur];<br />
holotype: ZISP 2488, Eschmeyer & Fricke, 2010; adjective,<br />
-us, -a, -um)<br />
Cobitis ajdan Dybowski, in Sinicyn, 1900: 49 (nomen nudum;<br />
material: "Imperial Warsaw University" 2769 [Onon<br />
River], ZMB 7116 [2, Dauria])<br />
Ussuria leptocephala Nikolski, 1904: 362 (type locality:<br />
Russia: Ussuri River [Ussury River near Khabarovsk] /<br />
Mongolia: Cherulu River [Kherlen; Vasil'eva, 2001: 553];<br />
syntypes: ZISP 10655 [1], ZISP 12791 [1]; compound<br />
adjective, -us, -a, -um)<br />
Misgurnus erikssoni Rendahl, 1922: 3 (type locality: Mongolia:<br />
Djaggaste [about 120 km on road from Kalgan<br />
[Zhangjiakou] to Urga [Ulaan Baatar], 41.7039°N<br />
113.839°E, NRM catalogue]; syntypes [18]: NRM 10339<br />
[13], MNHN 1933.115 [1], AMNH [2], Bertin & Estève,<br />
1948: 95, Eschmeyer & Fricke, 2010; noun in genitive,<br />
indeclinable)<br />
4.13.5 Misgurnus multimaculatus Rendahl, 1944<br />
Misgurnus mizolepis multimaculatus Rendahl, 1944: 13, fig. 6<br />
(type locality: Vietnam: Annam: Thua Luu, 50 km southeast<br />
<strong>of</strong> Hué; holotype: NHMG; adjective, -us, -a, -um)<br />
4.13.6 Misgurnus nikolskyi Vasil'eva, 2001<br />
Misgurnus nikolskyi Vasil'eva, 2001: 589, fig. 3 [p. 560 <strong>of</strong><br />
translation] (type locality: Russia: Primorsky Kray: a lake<br />
near Lefu River [Ilistaya; a tributary <strong>of</strong> Lake Khanka],<br />
near Novokorovinskaya [p. 561]; holotype: ZMMU<br />
P-20792; noun in genitive, indeclinable)<br />
4.13.7 Misgurnus tonkinensis Rendahl, 1937<br />
Misgurnus mizolepis tonkinensis Rendahl, 1937: 2, fig. 1<br />
(type locality: Vietnam: Tonkin: Hanoi; holotype: NRM<br />
10653; adjective, -is, -is, -e)<br />
4.14 Neoeucirrhichthys Banarescu & Nalbant, 1968<br />
Neoeucirrhichthys Bănărescu & Nalbant, 1968: 349 (type<br />
species: Neoeucirrhichthys maydelli Bănărescu & Nalbant,<br />
1968: 349, by original designation). Gender masculine.
4.14.1 Neoeucirrhichthys maydelli, CMK 21126, 31.8 mm SL;<br />
Bangladesh: Brahmaputra drainage.<br />
4.15.1 Niwaella delicata, 82 mm SL; Japan: Shiga Prefecture: Yasu<br />
River. (Photograph by Kazumi Hosoya).<br />
4.14.1 Neoeucirrhichthys maydelli Banarescu & Nalbant,<br />
1968<br />
Neoeucirrhichthys maydelli Bănărescu & Nalbant, 1968:<br />
349, fig. 15 (type locality: India: Assam: Goalpara District:<br />
Janali River at Raimona; holotype: ZMH H 3609,<br />
Wilkens, 1977: 158; noun in genitive, indeclinable)<br />
4.15 Niwaella Nalbant, 1963<br />
Niwaëlla Nalbant, 1963: 362 (type species: Cobitis delicata<br />
Niwa, 1937: 72, by original designation). Gender feminine.<br />
Taxonomic notes. Niwaella laterimaculata, N. longibarba<br />
and N. xinjiangensis have a colour pattern somewhat similar<br />
to that <strong>of</strong> Iksookimia but do not have sexually dimorphic<br />
pectoral fins. They were placed in Niwaella by Chen & Chen<br />
(2005), which, besides being geographically distant, has a<br />
different appearance and colour pattern. They are tentatively<br />
retained in Niwaella but may represent an unnamed lineage.<br />
4.15.1 Niwaella delicata (Niwa)<br />
Cobitis delicata Niwa, 1937: 72 (type locality: Japan: Agi<br />
River, a tributary <strong>of</strong> Kiso River; holotype: LU; adjective,<br />
-us, -a, -um)<br />
4.15.2 ? Niwaella laterimaculata (Yan & Zheng, 1984)<br />
Cobitis laterimaculata Yan & Zheng, 1984: 82, fig. 1 (type<br />
locality: China: Zhejiang: Fenghua County: Yongjiang<br />
River at Xikou; holotype: LFZFC 821138; adjective,<br />
-us, -a, -um)<br />
4.15.3 ? Niwaella longibarba Chen & Chen, 2005<br />
Niwaella longibarba Chen & Chen, 2005: 1647, figs. 1d, 5<br />
(type locality: China: Zhejiang: Chengxian County:<br />
Cao'ejiang River, Huangzezhen, 29°36.3'N 120°54.4'E;<br />
holotype: IHB 9607011; compound noun, indeclinable)<br />
4.15.4 ? Niwaella xinjiangensis Chen & Chen, 2005<br />
Niwaella xinjiangensis Chen & Chen, 2005: 1649, figs. 1e, 6<br />
(type localoity: China: Jiangxi: Guafeng County: Xinjiang<br />
River, "main upper stream <strong>of</strong> Boyanghu Lake",<br />
Yangtze drainage, 28°26.1'N 119°10.2'E; holotype: IHB<br />
90V1610; adjective, -is, -is, -e)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
37<br />
4.16.16 Pangio kuhlii, CMK 10045, 66.2 mm SL; Indonesia: Java:<br />
Bantan.<br />
4.16.17 Pangio lidi, CMK 21800, 62.1 mm SL; Indonesia: Borneo:<br />
Mahakam drainage.<br />
4.16 Pangio Blyth, 1860<br />
Acanthophthalmus Bleeker, 1858: 304 (incorrect subsequent<br />
spelling <strong>of</strong> Acantophthalmus van Hasselt, 1823: 132; not<br />
a junior homonym <strong>of</strong> Acanthophthalmus van Hasselt,<br />
1824: 376 as this is also an incorrect subsequent spelling<br />
<strong>of</strong> Acantophthalmus van Hasselt, 1823: 132 and on Official<br />
Index <strong>of</strong> Rejected and Invalid Generic Names in<br />
Zoology, ICZN, 1992b: 248 [Opinion 1695])<br />
Pangio Blyth, 1860: 169 (type species: Cobitis cinnamomea<br />
M'Clelland, 1839: 304, by monotypy). Gender feminine<br />
[Code art. 30.2.3].<br />
Apua Blyth, 1860: 169 (type species: Apua fusca Blyth, 1860:<br />
169, by monotypy; simultaneous subjective synonym <strong>of</strong><br />
Pangio Blyth, 1860: 169, first reviser [Kottelat, 1987b:<br />
371] gave precedence to Pangio). Gender feminine.<br />
Eucirrhichthys Perugia, 1892: 1009 (type species: Eucirrhichthys<br />
doriae Perugia, 1892: 1009, by monotypy).<br />
Gender masculine.<br />
Cobitophis Myers, 1927: 4 (type species: Acanthophthalmus<br />
anguillaris Vaillant, 1902: 151, by original designation).<br />
Gender masculine.<br />
Nomen nudum:<br />
Pangio bashai Easa & Shaji, 1997: 181 (nomen nudum;<br />
locality: India: Kerala: Nilgiri Biosphere Reserve); Raju<br />
Thomas et al., 1999: 479 (nomen nudum); Anna Mercy,<br />
Gopalakrishan, Kapoor & Lakra, 2007: 142, fig. (nomen<br />
nudum; locality: none stated, but India: Western Ghat)<br />
4.16.1 Pangio agma (Burridge, 1992)<br />
? Acanthophthalmus borneensis Boulenger, 1894a: 251 (type<br />
locality: Malaysia: Borneo: Sarawak: Baram River; syntypes:<br />
BMNH 1889.7.31.14 [1], MNHN 1894.18 [1],<br />
Bertin & Estève, 1948: 91, Eschmeyer & Fricke, 2010,<br />
Kottelat et al., 1993: pl. 29; adjective, -is, -is, -e)<br />
Acanthophthalmus agmus Burridge, 1992: 180, fig. 6 (type<br />
locality: Malaysia: Borneo: Sarawak: Fourth Division:<br />
Sungai Lawa, 1 km upstream <strong>of</strong> confluence with Baram<br />
River; holotype: ROM 58726; adjective, -us, -a, -um)<br />
4.16.2 Pangio alcoides Kottelat & Lim, 1993<br />
Pangio alcoides Kottelat & Lim, 1993: 208, fig. 1 (type local-
ity: Malaysia: Terengganu: Rantau Abang, stream at<br />
154-km stone on road from Kuala Terengganu to Kuantan;<br />
holotype: ZRC 17209; adjective, indeclinable)<br />
4.16.3 Pangio alternans Kottelat & Lim, 1993<br />
Pangio alternans Kottelat & Lim, 1993: 210, fig. 3 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Mahakam<br />
drainage: blackwater stream entering Mahakam<br />
River on northern side near Mujub; 0°01'S 115°43'E;<br />
holotype: MZB 5895; participle, indeclinable)<br />
4.16.4 Pangio ammophila Britz, Ali & Raghavan, 2012<br />
Pangio ammophila Britz, Ali & Raghavan, 2012: 46, figs. 1–<br />
2 (type locality: India: Karnataka: Subramanya, Kumaradhara<br />
river, 12°00'19"N 75°53'24" E, 811 masl; holotype:<br />
CRG-SAC.2012.1.1; adjective, -us, -a, -um)<br />
4.16.5 Pangio anguillaris (Vaillant, 1902)<br />
Acanthophthalmus anguillaris Vaillant, 1902: 151, fig. 46<br />
(type locality: Indonesia: Borneo: Kalimantan Barat:<br />
Kapuas River, possibly at Sintang; holotype: RMNH<br />
7788, Eschmeyer & Fricke, 2010 [a single specimen implied<br />
on p. 153]; adjective, -is, -is, -e)<br />
Acanthophthalmus vermicularis Weber & de Beaufort, 1916:<br />
34 (type locality: Indonesia: Sumatra: Kampar Kiri River;<br />
holotype: ZMA 100.260, Nijssen et al., 1982: 26;<br />
adjective, -is, -is, -e)<br />
Cobitophis perakensis Herre, 1940: 8, pl. 2 (type locality:<br />
Malaysia: Perak: lake above Chenderoh Dam; holotype:<br />
CAS-SU 33004, Böhlke, 1953: 39; adjective, -is, -is, -e)<br />
Taxonomic notes. Several species are confused under this<br />
name. See also molecular data in Bohlen et al. (2011).<br />
4.16.6 Pangio apoda Britz & Maclaine, 2007<br />
Pangio apoda Britz & Maclaine, 2007: 28, fig. 8 (type locality:<br />
India: West Bengal: Brahmaputra drainage: Tista<br />
River at Tista Barrage; holotype: UMMZ 247779; adjective,<br />
-us, -a, -um)<br />
4.16.7 Pangio atactos Tan & Kottelat, 2009<br />
Pangio atactos Tan & Kottelat, 2009: 57, fig. 42 (type locality:<br />
Indonesia: Sumatra: Jambi: Danau Kamining near<br />
Kampung Trasos, ca. 5 km southwards on unpaved road<br />
turning-<strong>of</strong>f from Muara Bungo - Muara Tebo road at<br />
km 36; holotype: MZB 10995; treated as noun in apposition,<br />
indeclinable)<br />
4.16.8 Pangio bitaimac Tan & Kottelat, 2009<br />
Pangio bitaimac Tan & Kottelat, 2009: 59, fig. 43 (type locality:<br />
Indonesia: Sumatra: Jambi: Sungai Alai; holotype:<br />
MZB 10997; noun in apposition, indeclinable)<br />
4.16.9 Pangio cuneovirgata (Raut, 1957)<br />
Acanthophthalmus cuneovirgatus Raut, 1957: 30, figs. (type<br />
locality: unknown, possibly "Jahore/Hinterindien"<br />
[Malaysia: Johor]; holotype: ZMB 21332; adjective, -us,<br />
-a, -um)<br />
4.16.10 Pangio doriae (Perugia, 1892)<br />
Eucirrhichthys doriae Perugia, 1892: 1009 (type locality:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
38<br />
Malaysia: Borneo: Sarawak; syntypes: MCSNG 9231 [4],<br />
ZMA 114.898 [1], Tortonese, 1961: 188, Nijssen et al.,<br />
1982: 28; noun in genitive, indeclinable)<br />
4.16.11 Pangio elongata Britz & Maclaine, 2007<br />
Pangio elongata Britz & Maclaine, 2007: 18, fig. 1 (type<br />
locality: Myanmar: Tenasserim: mouth <strong>of</strong> Mingaw Kloh<br />
[Min Ngaw Chaung, stream entering Tenasserim River<br />
at 13°25'N 99°01'E]; holotype: BMNH 1992.11.16.13;<br />
adjective, -us, -a, -um)<br />
4.16.12 Pangio filinaris Kottelat & Lim, 1993<br />
Pangio filinaris Kottelat & Lim, 1993: 219, fig. 9 (type locality:<br />
Malaysia: Terengganu: Sungai Tersat, a tributary <strong>of</strong><br />
Sungai Terengganu, immediately downriver <strong>of</strong> Sekayu<br />
Waterfall Park, 4°57'51"N 102°57'45"E; holotype: ZRC<br />
34915; compound noun, indeclinable)<br />
4.16.13 Pangio fusca (Blyth, 1860)<br />
Apua fusca Blyth, 1860: 169 (type locality: Burma: Tenasserim;<br />
syntypes [3]: ZSI F 2647/1 [2], ? AMS B.7500<br />
[1], Hora, 1921d: 32, Menon & Yazdani, 1968: 120, Eschmeyer<br />
& Fricke, 2010; adjective, -us, -a, -um)<br />
4.16.14 Pangio goaensis (Tilak, 1972)<br />
Acanthophthalmus goaensis Tilak, 1972: 62, fig. 1 (type<br />
locality: India: Goa: Colem River; holotype: ZSI F 6150/<br />
2; adjective, -is, -is, -e)<br />
4.16.15 Pangio incognito Kottelat & Lim, 1993<br />
Pangio incognito Kottelat & Lim, 1993: 221, fig. 11 (type<br />
locality: Borneo: Sarawak: km 42 on road from Lindu to<br />
Kuching, west <strong>of</strong> Sungei Stinggang; holotype: ZRC<br />
34913; treated as noun in apposition, indeclinable)<br />
4.16.16 Pangio kuhlii (Valenciennes, in Cuvier & Valenciennes,<br />
1846)<br />
Acantophthalmus fasciatus van Hasselt, 1823: 133 (nomen<br />
nudum, Kottelat, 1987b: 371)<br />
Acanthophthalmus fasciatus van Hasselt, 1824: 377 (nomen<br />
nudum)<br />
Cobitis kuhlii Valenciennes, in Cuvier & Valenciennes, 1846:<br />
77 (type locality: Indonesia: Java: Krawang; neotype:<br />
RMNH 2688, designated by Burridge, 1992: 182 [holotype<br />
listed by Bertin & Estève, 1948: 91, Eschmeyer &<br />
Fricke, 2010, has no type status, see Burridge, 1992: 181];<br />
noun in genitive, indeclinable)<br />
Acanthophthalmus fasciatus Bleeker, 1860c: 74 (type locality:<br />
Indonesia: Java: Krawang; neotype: RMNH 2688,<br />
designated by Kottelat & Lim, 1993: 224; objective junior<br />
synonym <strong>of</strong> Cobitis kuhlii Valenciennes, in Cuvier &<br />
Valenciennes, 1846: 77; adjective, -us, -a, -um)<br />
4.16.17 Pangio lidi Hadiaty & Kottelat, 2009<br />
Pangio lidi Hadiaty & Kottelat, 2009b: 65, figs. 1–2 (type<br />
locality: Indonesia: Kalimantan Timur: Mahakam drainage,<br />
Ulu Belayan, Sungai Petung Kanan, 0°32'06"N<br />
116°10'55"E; holotype: MZB 16528; noun in apposition,<br />
indeclinable)
4.16.18 Pangio longimanus Britz & Kottelat, 2010<br />
Pangio longimanus Britz & Kottelat, 2010: 373, fig. 1 (type<br />
locality: Laos: Vientiane Province: confluence <strong>of</strong> Nam<br />
Leuk and Nam Gnong, 18°22'04"N 103°05'27"E; holotype:<br />
ZRC 51933; compound noun, indeclinable)<br />
4.16.19 Pangio lumbriciformis Britz & Maclaine, 2007<br />
Pangio lumbriciformis Britz & Maclaine, 2007: 22, fig. 4<br />
(type locality: Myanmar: Irrawaddy basin: Nan Kwe<br />
stream [near Myitkyna]; holotype: BMNH 2006.9.29.6;<br />
adjective, -is, -is, -e)<br />
4.16.20 Pangio malayana (Tweedie, 1956)<br />
Acanthophthalmus kuhlii malayanus Tweedie, 1956: 58,<br />
pl. 6a (type locality: Malaysia: Pahang: Kuala Tahan; holotype:<br />
BMNH 1957.1.23.1; adjective, -us, -a, -um)<br />
4.16.21 Pangio mariarum (Inger & Chin, 1962)<br />
Acanthophthalmus mariae Inger & Chin, 1962: 118, fig. 56<br />
(type locality: Malaysia: Borneo: Sabah: Kinabatangan<br />
District: small tributary <strong>of</strong> Kinabatangan River at Deramakot;<br />
holotype: FMNH 68161; incorrect original spelling,<br />
emended into mariarum [Code art. 31.1.2], Kottelat,<br />
1989: 13; noun in genitive, indeclinable)<br />
Nomenclatural notes. The species was named for two women<br />
named Mary. The original spelling mariae must be emended<br />
into mariarum (Code art. 31(a)(ii); Kottelat, 1989: 13).<br />
4.16.22 Pangio muraeniformis (de Beaufort, 1933)<br />
Acanthophthalmus muraeniformis de Beaufort, 1933: 32<br />
(type locality: Singapore: Thomson Road; syntypes: ZRC<br />
1052 [2], ZMA 103.185 [3], CAS-SU 32602 [1], Nijssen<br />
et al., 1993: 213, Böhlke, 1953: 39; adjective, -is,<br />
-is, -e)<br />
4.16.23 Pangio myersi (Harry, 1949)<br />
Acanthophthalmus myersi Harry, 1949: 69 (type locality:<br />
Southeastern Thailand: Nong Khor; holotype: USNM<br />
103300; noun in genitive, indeclinable)<br />
4.16.24 Pangio oblonga (Valenciennes, in Cuvier & Valenciennes,<br />
1846)<br />
Acantophthalmus Javanicus van Hasselt, 1823: 133 (nomen<br />
nudum, Kottelat, 1987b: 371)<br />
Acanthophthalmus javanicus van Hasselt, 1824: 377 (nomen<br />
nudum)<br />
Cobitis oblonga Valenciennes, in Cuvier & Valenciennes,<br />
1846: 76 (type locality: Indonesia: Java: Bogor; neotype:<br />
ZRC 35047, designated by Kottelat & Lim, 1993: 234<br />
[RMNH 2710 listed as possible syntypes by Eschmeyer<br />
& Fricke, 2010 cannot be syntype as Valenciennes based<br />
his description on a drawing <strong>of</strong> a single specimen]; adjective,<br />
-us, -a, -um)<br />
Acanthophthalmus javanicus Bleeker, 1860c: 75 (type locality:<br />
Indonesia: Java: Bogor; neotype: ZRC 35047, designated<br />
by Kottelat & Lim, 1993: 235 [RMNH 7061 listed<br />
by Eschmeyer & Fricke, 2010 are not syntypes; a lectotype<br />
designation by Kottelat & Lim, 1993: 235 made them<br />
paralectotypes if they really were syntypes; the lectotype<br />
is lost and a neotype was designated]; objective junior<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
39<br />
synonym <strong>of</strong> Cobitis oblonga Valenciennes, in Cuvier &<br />
Valenciennes, 1846: 76; adjective, -us, -a, -um)<br />
Taxonomic notes. Several species are confused under this<br />
name. See also molecular data in Bohlen et al. (2011).<br />
4.16.25 Pangio pangia (Hamilton, 1822)<br />
Cobitis pangia Hamilton, 1822: 355, 394 (type locality: India:<br />
"north-eastern parts <strong>of</strong> Bengal" [Goalpara; Hora,<br />
1935a: 49]; types: NT; on Official List <strong>of</strong> Specific Names<br />
in Zoology, ICZN 1992b: 248 [Opinion 1695]; Hamilton's<br />
unpublished figure reproduced in M'Clelland, 1839:<br />
pl. 51 fig. 5, Britz & Maclaine, 2007: 25, fig. 6; noun in<br />
apposition, indeclinable)<br />
Cobitis cinnamomea M'Clelland, 1839: 304, 435, pl. 51 fig. 5<br />
(unnecessary replacement name for Cobitis pangia<br />
Hamilton, 1822: 335; adjective, -us, -a, -um)<br />
Canthophrys rubiginosus Swainson, 1839: 310 (available<br />
by indication to Hamilton, 1822: "Cob. No. 6" [which is<br />
Cobitis pangia]; type locality: India: "north-eastern parts<br />
<strong>of</strong> Bengal" [Goalpara; Hora, 1935a: 49]; types: NT;<br />
Hamilton's unpublished figure reproduced in M'Clelland,<br />
1839: pl. 51 fig. 5; adjective, -us, -a, -um)<br />
Taxonomic notes. At least two species are confused under<br />
this name. See also molecular data in Bohlen et al. (2011).<br />
4.16.26 Pangio piperata Kottelat & Lim, 1993<br />
Pangio piperata Kottelat & Lim, 1993: 236, fig. 20 (type locality:<br />
Malaysia: Terengganu: stream at about km 6 on road<br />
from Kuala Brang to Kuala Terengganu, 6°04'25"N<br />
103°03'20"E; holotype: ZRC 35003; adjective, -us, -a, -um)<br />
Taxonomic notes. At least two species are confused under<br />
this name. See also molecular data in Bohlen et al. (2011).<br />
4.16.27 Pangio pulla Kottelat & Lim, 1993<br />
Pangio pulla Kottelat & Lim, 1993: 238, fig. 21 (type locality:<br />
Indonesia: Kalimantan Tengah: heath forest northwest<br />
<strong>of</strong> Palankaraya; blackwater stream, 28–29 km on<br />
road to Tangkiling; tributary <strong>of</strong> Sungei Rungan, Sungei<br />
Kahajan drainage; holotype: ZRC 35022; adjective, -us,<br />
-a, -um)<br />
4.16.28 Pangio robiginosa (Raut, 1957)<br />
Acanthophthalmus robiginosus Raut, 1957: 31, figs. (type<br />
locality: Indonesia: Java: stream Tjipaja-en and tributaries<br />
near Rangkas-Betong village, Bantean area; holotype:<br />
ZMB 21334; adjective, -us, -a, -um)<br />
4.16.29 Pangio semicincta (Fraser-Brunner, 1940)<br />
Acanthophthalmus semicinctus Fraser-Brunner, 1940: 172,<br />
fig. 3 (type locality: Malaysia: Johor: Mawai District;<br />
holotype: BMNH 1938.12.1.113; adjective, -us, -a, -um)<br />
Acanthophthalmus kuhlii sumatranus Fraser-Brunner, 1940:<br />
175, fig. 4B (type locality: Sumatra: Palembang Province:<br />
Lahat; holotype: BMNH 1866.5.2.41; simultaneous<br />
subjective synonym <strong>of</strong> Acanthophthalmus semicinctus<br />
Fraser-Brunner, 1940: 172; first revisers [Tan & Kottelat,<br />
2009: 61] gave precedence to A. semicinctus; adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Several species are confused under this<br />
name. See also molecular data in Bohlen et al. (2011).
4.16.23 Pangio myersi, CMK 20250, 64.1 mm SL; Thailand: Trat.<br />
4.16.28 Pangio robiginosa, CMK 10046, 39.0 mm SL; Indonesia:<br />
Java: Bantan.<br />
4.16.30 Pangio shelfordii (Popta, 1903)<br />
Acanthophthalmus shelfordii Popta, 1903: 231, fig. (type<br />
locality: Malaysia: Borneo: Sarawak: Sarawak River near<br />
Kuching; holotype: RMNH 7661; noun in genitive, indeclinable)<br />
Taxonomic notes. Several species are confused under this<br />
name. See also molecular data in Bohlen et al. (2011).<br />
4.16.31 Pangio signicauda Britz & Maclaine, 2007<br />
Pangio signicauda Britz & Maclaine, 2007: 26, fig. 7 (type<br />
locality: Myanmar: Irrawaddy basin: Nanmate stream,<br />
25°23'03"N 97°00'38"E, between Myitkina and Hopin;<br />
holotype: USNM 378386; compound noun, indeclinable)<br />
4.16.32 Pangio superba (Roberts, 1989)<br />
Acantophthalmus superbus Roberts, 1989: 98, fig. 74 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: forest<br />
stream about 1 km up Sungai Tajan from Tajan, 87 km<br />
east <strong>of</strong> Pontianak; holotype: MZB 3499; adjective, -us, -<br />
a, -um)<br />
4.17 Paralepidocephalus Tchang, 1935<br />
Paralepidocephalus Tchang, 1935: 17 (type species: Paralepidocephalus<br />
yui Tchang, 1935: 17, by monotypy).<br />
Gender masculine.<br />
Species inquirenda<br />
Paralepidcephalus [sic] guishanensis Li, 2004: 94, fig. 3<br />
(type locality: China: Yunnan: Shilin County: Fenglongtan<br />
in Guishan Xiang; holotype: HRAS 980514001; adjective,<br />
-is, -is, -e)<br />
4.17.1 Paralepidocephalus yui Tchang, 1935<br />
Paralepidocephalus yui Tchang, 1935: 17, fig. 1 (type locality:<br />
China: Yunnan: Shiping; holotype: ASIZB 11678,<br />
Zhang, 1996: 500; noun in genitive, indeclinable)<br />
4.18 Paramisgurnus Dabry de Thiersant, 1872<br />
Paramisgurnus Dabry de Thiersant, 1872: 191 (type species:<br />
Paramisgurnus dabryanus Dabry de Thiersant,<br />
1872: 191, by monotypy). Gender masculine.<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
40<br />
4.17.1 Paralepidocephalus yui, KIZ 2006009175, 74.3 mm SL;<br />
China: Yunnan: Pearl River drainage: Niujie River. (Photograph<br />
by Chen Xiao-Yong).<br />
4.18.1 Paramisgurnus dabryanus, KIZ JWS2012000, 146.0 mm<br />
SL; China: Yunnan: Yangtze drainage: Lake Bita. (Photograph by<br />
Chen Xiao-Yong).<br />
4.19.2 Protocobitis typhlops, KIZ 9180001, 54.0 mm SL, lectotype;<br />
China: Guangxi: Xia’ao. (Photograph by Ren Qiu).<br />
Paramisgurnus Sauvage, 1878a: 90 (type species: Paramisgurnus<br />
dabryanus Sauvage, 1878a: 90, by monotypy;<br />
junior homonym and objective synonym <strong>of</strong> Paramisgurnus<br />
Dabry de Thiersant, 1872: 191). Gender masculine.<br />
4.18.1 Paramisgurnus dabryanus Dabry de Thiersant, 1872<br />
Paramisgurnus dabryanus Dabry de Thiersant, 1872: 191,<br />
pl. 49 fig. 6 (type locality: China: Yang-tsee-kiang<br />
[Yangtze]; holotype: MNHN 5087, Fang, 1935b: 148,<br />
fig. 11, Bertin & Estève, 1948: 100; adjective, -us, -a,<br />
-um)<br />
Paramisgurnus dabryanus Sauvage, 1878a: 90 (type locality:<br />
China: Yang-tse-Kiang [Yangtze]; holotype: MNHN<br />
5087, Fang, 1935b: 148, fig. 11, Bertin & Estève, 1948:<br />
100; junior homonym and objective synonym <strong>of</strong> Paramisgurnus<br />
dabryanus Dabry de Thiersant, 1872: 191; adjective,<br />
-us, -a, -um)<br />
Misgurnus mizolepis Günther, 1888: 434 (type locality: China:<br />
Yangtsze-Kiang [Yangtze] at Kiu-Kiang; syntypes:<br />
BMNH 1888.3.23.83–85 [3]; compound noun, indeclinable)<br />
Misgurnus oligolepos Chen, Shen & Li, 1994: 20, fig. (type<br />
locality: China: Tianjin Municipality: Tanggu area, south<br />
<strong>of</strong> the mouth <strong>of</strong> Beitang River, fyke net fishing grounds;<br />
holotype: 880902, but location not stated, probably Tianjin<br />
Natural History <strong>Museum</strong> or Tianjin Fisheries <strong>Research</strong><br />
Institute; compound noun, indeclinable)<br />
Taxonomic notes. Genetic data (e.g. Shimizu & Takagi,<br />
2010) suggest that several species are confused under the<br />
name P. dabryanus. A review <strong>of</strong> the literature and published<br />
illustrations shows variation in the body shape <strong>of</strong> populations<br />
referred to P. dabryanus. This is, however, not conclusive<br />
because most <strong>of</strong> the Chinese faunal works include fig-
ures copied from earlier works and not representative <strong>of</strong> the<br />
concerned populations.<br />
It seems that a group <strong>of</strong> populations have a short and<br />
tapering caudal peduncle with high dorsal and ventral crests<br />
originating immediately behind dorsal and anal fins, a deep<br />
body and a somewhat pointed caudal fin. This agrees with<br />
the holotype <strong>of</strong> P. dabryanus (figured by Fang, 1935b: fig. 11)<br />
and possibly M. oligolepos. The original description <strong>of</strong><br />
M. mizolepis also mentions the high crests on the caudal peduncle.<br />
Chen (1981) placed M. mizolepis in the synonymy<br />
<strong>of</strong> P. dabryanus.<br />
A second group has a more slender body, shorter and<br />
less deep crests and a more rounded caudal fin, which has<br />
<strong>of</strong>ten been called M. mizolepis. This may correspond to several<br />
<strong>of</strong> the subspecies <strong>of</strong> M. mizolepis described by Nichols<br />
and others and which Chen (1981) placed in the synonymy<br />
<strong>of</strong> M. anguillicaudatus. This is tentatively followed here but<br />
clearly it should be re-evaluated after examination <strong>of</strong> more<br />
material and the the type material <strong>of</strong> all.<br />
Nomenclature notes. The earliest description <strong>of</strong> M. oligolepos<br />
as a new species that I could examine is by Chen,<br />
Shen & Li (1994). Wang et al. (2001: 179) referred to a paper<br />
by Li (1993a), which could not be found. There are also<br />
mentions <strong>of</strong> the description <strong>of</strong> a Misgurnus olygoskolopos<br />
in a paper by Li (1993b), which also cannot be found (Chen<br />
Xiao-Yong, pers. comm.).<br />
4.19 Protocobitis Yang & Chen, 1993<br />
Protocobitis Yang & Chen, 1993: 125 (type species: Protocobitis<br />
typhlops Yang & Chen, 1993: 125, by monotypy;<br />
also in Yang, Chen & Lan, 1994: 92). Gender feminine.<br />
Taxonomic notes. It remains to be demonstrated that the<br />
two cave species P. polylepis and P. typhlops are really congerenic,<br />
that is, that they share an immediate common ancestor.<br />
This implies either that the ancestor was an epigean<br />
species and is now missing or still uncollected, or that this<br />
ancestor was a cave species that colonised two disjunct cave<br />
areas. Each species likely has a surface ancestor.<br />
Nomenclatural notes. Protocobitis and P. typhlops, both<br />
by Yang & Chen (1993), predate the same names in Yang et<br />
al. (1994) (Kottelat & Bréhier, 1999: 356).<br />
4.19.1 'Protocobitis' polylepis Zhu, Lü, Yang & Zhang, 2008<br />
Protocobitis polylepis Zhu, Lü, Yang & Zhang, 2008: 453,<br />
fig. 1 (type locality: China: Guangxi: Wuming County:<br />
underground source, 5 km from Wuming, 23°10'N<br />
108°17'E; holotype: KIZ 20060001; compound noun,<br />
indeclinable)<br />
4.19.2 Protocobitis typhlops Yang & Chen, 1993<br />
Protocobitis typhlops Yang & Chen, 1993: 125 (type locality:<br />
China: Guangxi: Du'an County: Xia'ao town, cave,<br />
24°15'N 107°05'E; lectotype: KIZ 918001, designated<br />
by Kottelat & Bréhier, 1999: 356; also in Yang, Chen &<br />
Lan, 1994: 93; compound noun, indeclinable)<br />
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41<br />
4.20 Sabanejewia Vladykov, 1829<br />
Sabanejewia Vladykov, 1929: 86 (type species: Cobitis balcanica<br />
Karaman, 1922: 307, by original designation).<br />
Gender feminine.<br />
4.20.1 Sabanejewia aurata (Filippi, 1863)<br />
Cobitis aurata Filippi, 1863: 391 (type locality: Iran: stream<br />
near Sartschem [apparently Sarcham-e S<strong>of</strong>la, 39°07'N<br />
47°54'E, B. W. Coad, pers. comm.]; lectotype: MZUT<br />
674, designated by Tortonese, 1961: 188; adjective, -us,<br />
-a, -um)<br />
Cobitis hohenackeri Kessler, 1877: 177 (type locality: Azerbaidjan:<br />
Kura River, Caspian Sea basin [Berg, 1949: 894];<br />
syntypes: ZISP 2854 [5], Eschmeyer & Fricke, 2010;<br />
author indicated as Brandt, but Kessler is actual author;<br />
noun in genitive, indeclinable)<br />
? Cobitis aralensis Kessler, 1877: 184 (type locality: Kazakhstan:<br />
mouth <strong>of</strong> Syr-Darya River / Uzbekistan: mouth<br />
<strong>of</strong> Amu-Darya River [Berg, 1949: 895]; syntypes [5]:<br />
ZISP 10932 [1], Eschmeyer & Fricke, 2010; adjective,<br />
-is, -is, -e)<br />
4.20.2 Sabanejewia balcanica (Karaman, 1922)<br />
Cobitis balcanica Karaman, 1922: 307 (type locality: Serbia:<br />
Save River near Zagreb / Bosnia-Herzegovina: Miljacka<br />
Stream, tributary <strong>of</strong> Bosna / FYROM: Vardar River<br />
(that is, its tributaries near Skopje (Üskub) and Veles)<br />
[type locality restricted by Bănărescu et al., 1972: 1 but<br />
irrelevant as this can only be done by lectotype designation];<br />
syntypes: MMNHS lost, ? MNHN 28.222 [1], Bertin<br />
& Estève, 1948: 93; adjective, -us, -a, -um)<br />
Cobitis montana Vladykov, 1925: 320, pl. 8 (type locality:<br />
Ukraine: streams Apica, Tereovka, Teresulka, Luzanka<br />
and Terebla, tributaries <strong>of</strong> Theiss River [Tisza] / Theiss<br />
River between villages Akna-Slatina and Butino; syntypes:<br />
[LU] 219; junior primary homonym <strong>of</strong> Cobitis<br />
montanus Jerdon, 1849: 332; adjective, -us, -a, -um)<br />
Cobitis aurata balcanica natio radnensis Jaszfalusi, 1951:<br />
116, pl. 1 fig. 1 (infrasubspecific, name not available;<br />
locality: Romania: Zebrak River, a tributary <strong>of</strong> Szamos<br />
River [Somesul ?] / Maros River [Mures] near Gödemesterhaza<br />
and creek Göde)<br />
Cobitis aurata balcanica morpha elongata Oliva, Balon &<br />
Franck, 1952: 295 (infrasubspecific, name not available;<br />
locality: Czechia: Moravia: Lipník on Bečva River)<br />
Cobitis aurata balcanica morpha elata Oliva, Balon &<br />
Franck, 1952: 295 (infrasubspecific, name not available;<br />
locality: Czechia: Moravia: Lipník on Bečva River)<br />
Cobitis aurata radnensis Bănărescu, Müller & Nalbant,<br />
1960: 124 (type locality: Romania: upper Mures and Tirnava<br />
Rivers; syntypes: LU; adjective, -is, -is, -e)<br />
Cobitis aurata bosniaca Karaman, 1963: 629, fig. 1 (type<br />
locality: Bosnia-Herzegovina: creeks Saturlija and Siroka<br />
near Banja Luka, Vrba River drainage; syntypes [144]:<br />
MMNHS, lost; adjective, -us, -a, -um)<br />
? Sabanejewia aurata doiranica Economidis, 1995: 211<br />
(nomen nudum)<br />
? Sabanejewia aurata thrakica Economidis, 1995: 211<br />
(nomen nudum)<br />
? Sabanejewia aurata doiranica Economidis & Nalbant,
4.20.2 Sabanejewia balcanica, CMK 17419, 74.4 mm SL; FYR<br />
<strong>of</strong> Macedonia: Vardar drainage.<br />
4.21.1 Theriodes sandakanensis, ZRC 37645, 25.3 mm SL; Malaysia:<br />
Borneo: Sabah: Kinabatangan drainage. (Photograph by<br />
Tan Heok Hui).<br />
1997: 318, figs. 14, 16d–f (type locality: Greece: Macedonia:<br />
Lake Doirani; holotype: DZAUT 1989-116; adjective,<br />
-us, -a, -um)<br />
? Sabanejewia aurata thrakica Economidis & Nalbant, 1997:<br />
320, figs. 15, 15g–i (type locality: Greece: Thrace: Erythropotamos<br />
stream near village Mikro Dereio, Evros River<br />
drainage; DZAUT 1986-54; adjective, -us, -a, -um)<br />
4.20.3 Sabanejewia baltica Witkowski, 1994<br />
Sabanejewia aurata baltica Witkowski, 1994: 44, fig. 10<br />
(type locality: Poland: Widawa River near Kielczopwek<br />
village, 4 km from Wroclaw, Oder drainage, 51°18'N<br />
17°12'E; holotype: NMHW 638; adjective, -us, -a, -um)<br />
4.20.4 Sabanejewia bulgarica (Drensky, 1928)<br />
Cobitis bulgarica Drensky, 1928: 171, fig. 5-2 (type locality:<br />
Bulgaria: small tributaries <strong>of</strong> Danube in Widdin [Vidin]<br />
and Danube mainstream at Krai Panaira, 3 km downriver<br />
<strong>of</strong> Widdin [Vidin]; lectotype: NRM 10432 [1 <strong>of</strong> 2],<br />
designated by Kottelat & Freyh<strong>of</strong>, 2009: 81, fig. 2; adjective,<br />
-us, -a, -um)<br />
Cobitis albicoloris Chichk<strong>of</strong>f, 1932: 368, pl. 1 fig. 3 (type<br />
locality: Bulgaria: small tributaries <strong>of</strong> Danube in Widdin<br />
[Vidin] and Danube mainstream at Krai Panaira, 3 km<br />
downriver <strong>of</strong> Widdin [Vidin] [original type locality: Bulgaria:<br />
creek Provadiiska Rieca]; neotype: NRM 10432<br />
[1 <strong>of</strong> 2], designated by Kottelat & Freyh<strong>of</strong>, 2009: 81,<br />
fig. 2; objective junior synonym <strong>of</strong> Cobitis bulgarica<br />
Drensky, 1928: 171; adjective in genitive [not nominative,<br />
Code art. 31.2], treated as noun in apposition, indeclinable)<br />
Cobitis taenia tesselatus Pietschmann, 1937: 29 (type locality:<br />
Bulgaria: Widin [Vidin] on lower Danube River; syntypes:<br />
NMW 48624 [2], EAWAG 996 [1], Kottelat, 1997:<br />
92; adjective, -us, -a, -um)<br />
4.20.5 Sabanejewia caspia (Eichwald, 1838)<br />
Cobitis caspia Eichwald, 1838: 133 (type locality: Azerbaidjan:<br />
Lenkoran [Lankaran, 38°45'N 48°51'E], Caspian Sea<br />
basin; types: NT; adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
42<br />
4.20.6 Sabanejewia caucasica (Berg, 1906)<br />
Cobitis caucasica Berg, 1906b: 37, fig. (type locality: Russia:<br />
Saagdan, Great Laba River, upper Kuban drainage;<br />
lectotype: ZISP 8560, designated by Eschmeyer, 1998:<br />
341; adjective, -us, -a, -um)<br />
Remarks. Description based on 7 specimens from Terek<br />
drainage (Caspian Sea basin) and one from Kuban drainage<br />
(Black Sea basin). Eschmeyer (1998: 341) designated the<br />
specimen from Kuban as lectotype and commented that the<br />
figure in the original description shows the lectotype. Berg<br />
explicitly stated that the figure is based on one <strong>of</strong> the 6 specimens<br />
from Terek in ZMMU earlier described as Cobitis<br />
aurata by Berg (1899: 73).<br />
4.20.7 Sabanejewia kubanica Vasil'eva & Vasil'ev, 1988<br />
Sabanejewia aurata kubanica Vasil'eva & Vasil'ev, 1988:<br />
210 [p. 33 <strong>of</strong> translation], fig. 2 (type locality: Russia:<br />
Nevinka River, Kuban River drainage; holotype; ZMMU<br />
P-16384; adjective, -us, -a, -um)<br />
4.20.8 Sabanejewia larvata (Filippi, 1859)<br />
Cobitis larvata Filippi, 1859: 50 (type locality: Italy: creeks<br />
near Settimo Torinese [NE <strong>of</strong> Torino]; syntypes: MZUT<br />
510 [4], MZUP [1], ZMUC 439–441 [3], MZUF 5575<br />
[1], NMW 48558 [1], ZMB 5014 [2], Tortonese, 1940:<br />
141, Cantoni 1882: 366, Nielsen, 1974: 49, Eschmeyer<br />
& Fricke, 2010; adjective, -us, -a, -um)<br />
Cobitis taenia var. conspersa Cantoni, 1882: 364, pl. 1 fig. 3<br />
(type locality: based on a specimen <strong>of</strong> unstated locality<br />
and reference to 2 specimens from Italy: Pavia, identified<br />
by B. Crivelli as C. larvata, cited by Pavesi, 1877:<br />
499 [as p. 87]; syntypes [total 3]: MZUP [2], not listed<br />
by Cantoni, 1882; adjective, -us, -a, -um)<br />
4.20.9 Sabanejewia romanica (Bacescu, 1943)<br />
Cobitis caspia romanica Băcescu, 1943: 137, figs. 4–6, 7G<br />
(type locality: Romania: Buzau County: Bratia River, a<br />
tributary <strong>of</strong> Arges, at Vladesti, Muscel; lectotype: MGAB<br />
49916, designated by Kottelat, 1997: 93; adjective, -us,<br />
-a, -um)<br />
4.20.10 Sabanejewia vallachica (Nalbant, 1957)<br />
Cobitis aurata vallachica Nalbant, 1957: 209, figs. 1–3 (type<br />
locality: Romania: north-east Vallachia: Ialomitza River<br />
near Crivina; holotype: SMF 4087; adjective, -us, -a,<br />
-um)<br />
4.21 Theriodes Kottelat, 2012<br />
Theriodes Kottelat, 2012: 137 [appendix <strong>of</strong> present work]<br />
(type species: Acanthophthalmus sandakanensis Inger<br />
& Chin, 1962: 120, by original designation). Gender<br />
masculine.<br />
4.21.1 Theriodes sandakanensis (Inger & Chin, 1962)<br />
Acanthophthalmus sandakanensis Inger & Chin, 1962: 120,<br />
fig. 54E (type locality: Malaysia: Borneo: Sabah: Sandakan<br />
District: Sepilok Forest Reserve; holotype: FMNH<br />
68158; adjective, -is, -is, -e)
Family ELLOPOSTOMATIDAE<br />
5 Ellopostomatidae Bohlen & Šlechtová, 2009<br />
Ellopostomatinae Nalbant, 2002: pl. 7 (not available, no<br />
explicitly indicated as intentionally new, Code art. 16.1,<br />
no type genus cited, art. 16.2)<br />
Ellopostomatidae Bohlen & Šlechtová, 2009 [Aug.]: 161<br />
(type genus: Ellopostoma Vaillant, 1902: 145)<br />
Ellopostomatidae Chen, Lheknim & Mayden, 2009 [Dec.]:<br />
2204 (type genus: Ellopostoma Vaillant, 1902: 145; junior<br />
homonym <strong>of</strong> Ellopostomatidae Bohlen & Šlechtová,<br />
2009: 161)<br />
5.1 Ellopostoma Vaillant, 1902<br />
Ellopostoma Vaillant, 1902: 145 (type species: Aperioptus<br />
megalomycter Vaillant, 1902: 145, by monotypy). Gender<br />
neuter.<br />
5.1.1 Ellopostoma megalomycter (Vaillant, 1902)<br />
Aperioptus megalomycter Vaillant, 1902: 145, figs. 42–45<br />
(type locality: Indonesia: Borneo: Kalimantan Barat:<br />
Kapuas River; lectotype: RMNH 7777, designated by<br />
Roberts, 1972: 3, fig. 1; compound noun, indeclinable)<br />
5.1.2 Ellopostoma mystax Tan & Lim, 2002<br />
Ellopostoma mystax Tan & Lim, 2002: 454, fig. 3 (type locality:<br />
Thailand: Surat Thani Province: Tapi drainage: Khlong<br />
Sok on highway 410, 5 km west <strong>of</strong> Phanom; holotype:<br />
CAS 96698; noun in apposition, indeclinable)<br />
Family BARBUCCIDAE<br />
6 Barbuccidae Kottelat, 2012<br />
Barbuccidae Kottelat, 2012: 140 [appendix <strong>of</strong> present work]<br />
(type genus: Barbucca Roberts, 1989: 100)<br />
6.1 Barbucca Roberts, 1989<br />
Barbucca Roberts, 1989: 100 (type species: Barbucca diabolica<br />
Roberts, 1989: 100, by original designation). Gender<br />
feminine.<br />
6.1.1 Barbucca diabolica Roberts, 1989<br />
Barbuca diabolica Roberts, 1989: 100, fig. 75 (type locality:<br />
Indonesia: Borneo: Kalimantan Barat: small forested<br />
stream where it flows into Sungai Mandai 2–3 km upstream<br />
from confluence with Kapuas mainstream, 17 km<br />
west-southwest <strong>of</strong> Putussibau; holotype: MZB 3536; adjective,<br />
-us, -a, -um)<br />
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5.1.2 Ellopostoma mystax, CMK 5166, 46.1 mm SL; Thailand:<br />
Tapi drainage.<br />
5.1.2 Ellopostoma mystax, CAS 96698, 55.3 mm SL, holotype.<br />
(Photograph by Tan Heok Hui).<br />
6.1.1 Barbucca diabolica, ZRC uncat., 36.0 mm SL; Indonesia:<br />
Borneo: Kahayan drainage. (Photograph by Tan Heok Hui).<br />
6.1.1 Barbucca diabolica, ZRC 47747, 29.0 mm SL. (Photograph<br />
by Tan Heok Hui).
7 Balitoridae Swainson, 1839<br />
Balitorinae Swainson, 1839: 190 (type genus: Balitora Gray,<br />
1830: vol. 1, pl. 88)<br />
Homalopteraeformes Bleeker, 1860a: 422 (type genus:<br />
Homaloptera van Hasselt, 1823: 133)<br />
Sinohomalopterini Chen, 1980b: 208 (type genus: Sinohomaloptera<br />
Fang, 1930a: 26)<br />
Taxonomic notes. Chen (1980b) recognised two tribes within<br />
his subfamily Homalopterinae, which he called Homalopterini<br />
and Sinohomalopterini. I cannot read the Chinese text<br />
and therefore cannot comment on details. It is not clear to<br />
which non-Chinese species he had access. Judging from the<br />
list <strong>of</strong> species in his tree (p. 206) the coverage was limited.<br />
I am not able to recognise these two lineages as he defined<br />
them.<br />
I have earlier hypothesized that, within Balitoridae, a lineage<br />
can be defined by the morphology <strong>of</strong> the snout and the<br />
mouth (Kottelat & Chu, 1988c: 182). Within this lineage,<br />
Balitora and Hemimyzon are distinguished only by having<br />
two simple pelvic rays in Balitora (vs. 3–7 in Hemimyzon),<br />
a more slender body and pelvic fins widely separated (vs.<br />
more or less widened body, with pelvic-fin bases from widely<br />
separated posteriorly to fused). In the field, juveniles <strong>of</strong><br />
some sympatric species are almost indistinguishable (e.g.<br />
B. annamitica and H. papilio). Authors have placed species<br />
in one or the other genus only on the basis <strong>of</strong> the presence <strong>of</strong><br />
one (vs. two) barbels at the angle <strong>of</strong> the mouth, but the second<br />
barbel seems to be only an elongated papilla and is certainly<br />
not a character on which to distinguish genera. Sinohomaloptera<br />
was similarly distinguished from Balitora only<br />
by the presence <strong>of</strong> two barbels at the angle <strong>of</strong> the mouth;<br />
here too, the second barbel is an elongated papilla, but in<br />
most material I have examined, this papilla is not even elongated<br />
and therefore Sinohomaloptera cannot be distinguished<br />
from Balitora. In Chen's (1980b) tree, Balitora and Sinohomaloptera<br />
are in widely separated branches, in different<br />
tribes.<br />
The lineage comprising Balitora, Hemimyzon, Lepturichthys,<br />
Jinshaia, Metahomaloptera and Sinogastromyzon<br />
seems monophyletic and it could be tempting to call it a subfamily<br />
and place all other genera in a second subfamily. This,<br />
however, would be premature. To recognise two subfamilies<br />
implies that these remaining genera also constitute a monophyletic<br />
lineage, which is not demonstrated. Especially, there<br />
is no usable information to determine the position <strong>of</strong> the species<br />
described as 'Homaloptera' from Peninsular India.<br />
Nomen dubium<br />
Platyptera sinensis Bleeker, 1872: 127, 152 (type locality:<br />
China; holotype: specimen on which painting is based;<br />
adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Family BALITORIDAE<br />
44<br />
7.1 Balitora Gray, 1830<br />
Balitora Gray, 1830: vol. 1, pl. 88 (type species: Balitora<br />
brucei Gray, 1830: pl. 88, by subsequent designation by<br />
Jordan, 1919: 178). Gender feminine.<br />
Sinohomaloptera Fang, 1930a: 26 (as subgenus <strong>of</strong> Homaloptera<br />
van Hasselt, 1823: 133; type species: Homaloptera<br />
kwangsiensis Fang, 1930a: 27, by original designation).<br />
Gender feminine.<br />
Species inquirenda et incertae sedis<br />
Bhavania arunachalensis Nath, Dam, Bhutia, Dey & Das,<br />
2007: 72, pls. 1–2 (type locality: India: Arunachal<br />
Pradesh: Lohit District: Noadhing River drainage near<br />
Namsai, about 30 km from Tezu; holotype: ZSI/APFS<br />
P-488; adjective, -is, -is, -e)<br />
7.1.1 Balitora annamitica Kottelat, 1988<br />
Balitora annamitica Kottelat, 1988b: 498, fig. 5 (type locality:<br />
Cambodia: Boun Long [13°42'N 107°00'E], Grande<br />
Cascade; holotype: MNHN 1993.253; adjective, -us, -a,<br />
-um)<br />
7.1.2 Balitora brucei Gray, 1830<br />
Balitora Brucei Gray, 1830: vol. 1, pl. 88 fig. 1 (type locality:<br />
India: Assam: Priang River near Cherrapunji; neotype:<br />
RMNH 11924, designated by Kottelat, 1988b: 491;<br />
noun in genitive, indeclinable)<br />
Balitora maculata Gray, 1830: vol.1, pl. 88 fig. 2 (type locality:<br />
India: Assam: Priang River near Cherrapunji, Assam,<br />
India; neotype: RMNH 11924, designated by Kottelat,<br />
1988b: 491; simultaneous objective synonym <strong>of</strong> Balitora<br />
brucei Gray, 1830: vol. 1, pl. 88 fig. 1, first reviser<br />
[Kottelat, 1988b: 493] gave precedence to B. brucei;<br />
adjective, -us, -a, -um)<br />
Platycara anisura M'Clelland & Griffith, in M'Clelland,<br />
1842: 587, pl. 18 fig. 1 (type locality: India: Kassyah<br />
Hills; syntypes [2]: LU; compound noun, indeclinable)<br />
7.1.3 Balitora burmanica Hora, 1932<br />
Balitora brucei var. burmanicus Hora, 1932a: 291, pl. 11<br />
fig. 6 (type locality: Burma: Meekalan, Salween drainage;<br />
lectotype: MCSNG 15171, designated by Kottelat,<br />
1988b: 491; adjective, -us, -a, -um)<br />
Balitora brucei var. melanosoma Hora, 1932a: 291, pl. 10<br />
fig. 6 (type locality: Burma: stream Megla, Thaugyin<br />
River, on Thai-Burmese border; holotype: BMNH<br />
1920.9.8.1; could be treated as infrasubspecific and unavailable,<br />
but use as a subspecies [e.g. Hora, 1950: 52]<br />
before 1985 makes it available [Code art. 45.6.4.1]; simultaneous<br />
subjective synonym <strong>of</strong> Balitora brucei burmanicus<br />
Hora, 1932: 291; first reviser [Kottelat, 1988b:<br />
498] gave precedence to B. b. burmanica; compound<br />
noun, indeclinable)
Nomenclatural notes. Silas (1953: 207) listed specimen ZSI<br />
F 11034/1 as holotype <strong>of</strong> B. burmanica. As Hora (1932a)<br />
had not designated a holotype, all the specimens he examined<br />
were syntypes (see Kottelat, 1988a: 491).<br />
7.1.4 Balitora eddsi Conway & Mayden, 2010<br />
Balitora eddsi Conway & Mayden, 2010: 1467, fig. 1 (type<br />
locality: Nepal: Bardiya District: from a 25 km stretch <strong>of</strong><br />
Gerwa River, Karnali River drainage, between Chisapani<br />
and Kothiaghat, 28°34'00"N 81°13'59.88"E to<br />
28°23'60"N 81°11'59.99"E; holotype: KU 40276; noun<br />
in genitive, indeclinable)<br />
7.1.5 Balitora kwangsiensis (Fang, 1930)<br />
Homaloptera Kwangsiensis Fang, 1930a: 27, pl. 1 figs. 1–2<br />
(type locality: China: Guangxi: Lin-yueng-shien; holotype:<br />
MMNHN 899; adjective, -is, -is, -e)<br />
Sinohomaloptera h<strong>of</strong>fmanni Herre, 1938: 429, fig. 1 (type<br />
locality: China: Hainan: Cheung Kon Ts'uen; holotype:<br />
CAS-SU 33002, Hora, 1950: 54; noun in genitive, indeclinable)<br />
Balitora heteroura Pan, Liu & Zheng, 1983: 105, fig. 1 (type<br />
locality: China: Guangdong: Beijiang River; holotype (?):<br />
SCNU 8012; compound noun, indeclinable)<br />
? Balitora nigrocorpa Nguyen, 2005: 594, fig. 29 (type locality:<br />
Vietnam: Ha Giang Province: Lo River; holotype:<br />
NCNTTSI; as Sinohomaloptera in key and figure caption;<br />
also spelt nigrocopa p. 594 fig. 29, an obvious inadvertent<br />
error, thus incorrect original spelling [Code art.<br />
32.5.1]; corpa is not a Latin word, indeclinable)<br />
? Balitora vanlani Nguyen, 2005: 597, fig. 31 (type locality:<br />
Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; as Sinohomaloptera<br />
in key and figure caption; etymology not explained, a<br />
noun in genitive or in apposition, both indeclinable)<br />
? Balitora haithanhi Nguyen, 2005: 599, fig. 32 (type locality:<br />
Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; as Sinohomaloptera<br />
in key and figure caption; etymology not explained, a<br />
noun in genitive or in apposition, both indeclinable)<br />
? Hemimyzon songamensis Nguyen, 2005: 601, fig. 33 (type<br />
locality: Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; spelt sorgamensis<br />
p. 302, songamensis p. 602 fig. 33, 691, songamenis<br />
p. 601; as first reviser I select songamensis as the correct<br />
original spelling [in fact, none <strong>of</strong> these spellings is correct<br />
for a name based on Song Gam, but this cannot be<br />
corrected, Code art. 32.3]; adjective, -is, -is, -e)<br />
7.1.6 Balitora lancangjiangensis (Zheng, 1980)<br />
Sinohomaloptera lancangjiangensis Zheng, 1980: 110,<br />
figs. 1–2 (type locality: China: Yunnan: Menghai Xian;<br />
syntypes: IHB 78IV0565–569 [4], DBJU 78IV0432–435<br />
[4]; adjective, -is, -is, -e)<br />
? Balitora vanlongi Nguyen, 2005: 596, fig. 30 (type locality:<br />
Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; as Sinohomaloptera<br />
in key and figure caption; treated as a noun in genitive,<br />
indeclinable)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
45<br />
7.1.7 Balitora laticauda Bhoite, Jadhav & Dahanukar, 2012<br />
? Balitora shimogensis Silas & Kalawar, in Kalawar &<br />
Kelkar, 1958: 675 (nomen nudum; localities: India: Karnataka:<br />
Shimoga / Maharasthra: Kolhapur)<br />
Balitora laticauda Bhoite, Jadhav & Dahanukar, 2012: 3039,<br />
figs. 1–2 (type locality: India: Maharasthra: Satara District:<br />
Krishna drainage, stream at Venegaon village near<br />
Krishna River bridge, 17°29.94'N 74°07.08'E, 590 masl;<br />
holotype: ZSI/WRC P/2848; compound noun, indeclinable)<br />
7.1.8 Balitora ludongensis Liu & Chen, in Liu, Zhu, Wei<br />
& Chen, 2012<br />
Balitora ludongensis Liu & Chen, in Liu, Zhu, Wei & Chen,<br />
2012: 370, figs. 1–3 (type locality: China: Guangxi: Jingxi<br />
County: Ludong Town: Qilong stream in Pingjiang Village,<br />
Xijiang drainage; holotype: KIZ 2008008640; adjective,<br />
-is, -is, -e)<br />
7.1.9 Balitora longibarbata (Chen, in Zheng, Chen &<br />
Huang, 1982)<br />
Sinohomaloptera longibarbatus Chen, in Zheng, Chen &<br />
Huang, 1982: 394, fig. 1 (type locality: China: Yunnan:<br />
Yilian County: Nanpang-jiang drainage; 24°50'N<br />
103°10'E; syntypes: KIZ 774170–173, 175–179, 181<br />
[10]; adjective, -us, -a, -um)<br />
7.1.10 Balitora meridionalis Kottelat, 1988<br />
Balitora meridionalis Kottelat, 1988b: 498, fig. 4 (type locality:<br />
Thailand: Chantaburi Prov: Kao Soi Dao, Chan River<br />
headwaters; holotype: NIFI uncat.; adjective, -is, -is, -e)<br />
7.1.11 Balitora mysorensis Hora, 1941<br />
Balitora brucei var. mysorensis Hora, 1941b: 232 (type locality:<br />
India: Karnataka: Mysore: Sivasamudram, Cauvery<br />
drainage; holotype: ZSI F 13512/1, Bhoite et al., 2012:<br />
3039, figs. 3b, 4c; adjective, -is, -is, -e)<br />
7.1.12 ? Balitora nantingensis Chen, Cui & Yang, 2005<br />
Balitora nantingensis Chen, Cui & Yang, 2005: 22, fig. 2<br />
(type locality: China: Yunnan: Lincang Prefecture:<br />
Yongde County: Daxueshan township: Mangjiu River<br />
(23°58'55"N 99°41'17"E), a tributary <strong>of</strong> Nanting River<br />
[Salween drainage]; holotype: KIZ 20026475; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. Differences between B. nantingensis and<br />
B. burmanica (whose type locality is in the same drainage)<br />
seem slight, especially considering the variability observed<br />
in large series <strong>of</strong> B. burmanica from the middle Salween.<br />
The length <strong>of</strong> the caudal peduncle seems the less ambiguous<br />
character to distinguish the two species.<br />
7.2 Balitoropsis Smith, 1945<br />
Balitoropsis Smith, 1945: 278 (type species: Balitoropsis<br />
bartschi Smith, 1945: 279, by original designation). Gender<br />
feminine.<br />
Pseudohomaloptera Silas, 1953: 204 (type species: Homaloptera<br />
tatereganii Popta, 1905: 180, by original designation).<br />
Gender feminine.
7.1.3 Balitora burmanica, CMK 14679, 77.0 mm SL; Thailand:<br />
Salween drainage: Mae Nam Moei.<br />
7.1.1 Balitora cf. annamitica, CMK 22912, 65.4 mm SL.<br />
7.2.1 Balitoropsis batek (Tan, 2009)<br />
Homaloptera batek Tan, 2009: 49, figs. 1–4 (type locality:<br />
Indonesia: Borneo: Central Kalimantan: Katingan basin,<br />
Mendawai sub-basin, Sungei Baha'e, km 64 logging road<br />
at buffer zone <strong>of</strong> Bukit Raya-Bukit Baka National Park,<br />
0°47.593'S 112°19.220'E; holotype: MZB 10990; noun<br />
in apposition, indeclinable)<br />
7.2.2 Balitoropsis leonardi (Hora, 1941)<br />
Homaloptera leonardi Hora, 1941a: 61, pl. 5 figs. 5–6 (type<br />
locality: Malaysia: Pahang: King George V National Park<br />
[now Taman Negara National Park], Kuala Tahan; holotype:<br />
ZSI F 13213/1, Menon & Yazdani, 1968: 118; noun<br />
in genitive, indeclinable)<br />
Taxonomic notes. Tentatively placed in Balitoropsis.<br />
7.2.3 Balitoropsis ophiolepis (Bleeker, 1853)<br />
Homaloptera ophiolepis Bleeker, 1853a: 160 (type locality:<br />
Indonesia: Java: Bandong [Bandung]; lectotype: RMNH<br />
4986, designated by Alfred, 1961: 35; compound noun,<br />
indeclinable)<br />
7.2.4 Balitoropsis sexmaculata (Fowler, 1934)<br />
Homaloptera sexmaculata Fowler, 1934: 98, figs. 47–48<br />
(type locality: Thailand: Chiang Mai; holotype: ANSP<br />
56374; adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
46<br />
7.2.8 Balitoropsis yunnanensis, CMK 22214, 56.3 mm SL; Laos:<br />
Mekong drainage: Nam Theun.<br />
7.2.8 Balitoropsis yunnanensis, CMK 22214, 56.3 mm SL.<br />
Homaloptera septemmaculata Fowler, 1934: 99, figs. 49–<br />
50 (type locality: Thailand: Chiang Mai; holotype: ANSP<br />
56402; simultaneous subjective synonym <strong>of</strong> Homaloptera<br />
sexmaculata Fowler, 1934: 98, first reviser [Hora, 1950:<br />
51] gave precedence to H. sexmaculata; adjective, -us,<br />
-a, -um)<br />
7.2.5 Balitoropsis stephensoni (Hora, 1932)<br />
Homaloptera stephensoni Hora, 1932a: 281, pl. 11 fig. 1<br />
(type locality: Indonesia: Borneo: Kalimantan Timur:<br />
Upper Mahakam River; holotype: RMNH 7633, Tan,<br />
2009: 61, fig. 8; noun in genitive, indeclinable)<br />
? Homaloptera weberi Hora, 1932a: 284, pl. 11 fig. 2 (type<br />
locality: Malaysia: Borneo: Sarawak: Akar River; syntypes:<br />
BMNH 1895.7.2.81 [1], ZMA 100.990 [1], ZSI<br />
F 11292/1 [1], Nijssen et al., 1982: 29, Silas, 1953: 193,<br />
Menon & Yazdani, 1968: 119, Eschmeyer & Fricke, 2010;<br />
noun in genitive, indeclinable)<br />
7.2.6 Balitoropsis tatereganii (Popta, 1905)<br />
Homaloptera Tate Reganii Popta, 1905: 180 (type locality:<br />
Indonesia: Borneo: Bo River; holotype: RMNH 7632,<br />
Tan, 2009: 57, fig. 7; also in Popta, 1906: 182, pl. 10<br />
fig. 40; must be emended in tatereganii, Code art.
7.3.1 Bhavania australis, CMK 9312, 76.5 mm SL; India: Kerala:<br />
Chalakudy River. (Photograph by Tan Heok Hui).<br />
7.3.1 Bhavania australis, ZRC 34512, 82.0 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
32.5.2.2; noun in genitive, indeclinable)<br />
Taxonomic notes. Tentatively placed in Balitoropsis.<br />
7.2.7 Balitoropsis vulgaris (Kottelat & Chu, 1988)<br />
Homaloptera vulgaris Kottelat & Chu, 1988b: 103, fig. 1<br />
(type locality: China: Yunnan: Xishuangbanna: Mong Han<br />
County, 21°50'N 101°E; holotype: KIZ 788229; adjective,<br />
-is, -is, -e)<br />
7.2.8 Balitoropsis yunnanensis Chen, 1978<br />
Balitoropsis yunnanensis Chen, 1978: 334, figs. 1–2 (type<br />
locality: China: Yunnan: Lancang River [Mekong] in<br />
Yongping Xian [25°28'01"N 99°32'24"E]; holotype: IHB<br />
60VII012; adjective, -is, -is, -e)<br />
7.2.9 Balitoropsis yuwonoi (Kottelat, 1998)<br />
Homaloptera yuwonoi Kottelat, 1998b: 267, fig. 1 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: vicinity<br />
<strong>of</strong> Danau Sentarum Wildlife Reserve: Sungai Hulu Leboyan<br />
at Keluwin; 1°08'51"N 112°15'32"E; holotype:<br />
MZB 5938; noun in genitive, indeclinable)<br />
Taxonomic notes. Tentatively placed in Balitoropsis.<br />
7.2.10 Balitoropsis zollingeri (Bleeker, 1853)<br />
Homaloptera Javanica van Hasselt, 1823: 133, 1824: 377<br />
(nomen nudum, Kottelat, 1987b: 373)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
47<br />
7.4.1 Cryptotora thamicola, AMS I.25987-001, 28.4 mm SL,<br />
holotype; Thailand: Mae Hong Son Province: Tham Susa cave.<br />
Homaloptera Zollingeri Bleeker, 1853a: 159 (type locality:<br />
Indonesia: Java: Batavia [Jakarta] and Bandong [Bandung];<br />
syntypes: lost, Bleeker, 1860c: 89; noun in genitive,<br />
indeclinable)<br />
Homaloptera javanica Bleeker, 1860c: 89 (unnecessary replacement<br />
name for Homaloptera zollingeri Bleeker,<br />
1853a: 159; adjective, -us, -a, -um)<br />
Balitoropsis bartschi Smith, 1945: 279, fig. 56 (type locality:<br />
Thailand: Trang Province: waterfall stream on Kao<br />
Chong; holotype: USNM 107963; noun in genitive, indeclinable)<br />
Homaloptera nigra Alfred, 1969: 217, pl. 1 figs. 1–2 (type<br />
locality: Malaysia: Pahang: King George V National Park<br />
[now Taman Negara], Chegar Sireh, Tahan River; holotype:<br />
ZRC 2009; adjective, -er, -ra, -rum)<br />
7.3 Bhavania Hora, 1920<br />
Bhavania Hora, 1920: 202 (type species: Platycara australis<br />
Jerdon, 1849: 333, by subsequent designation by Silas,<br />
1953: 183). Gender feminine.<br />
7.3.1 Bhavania australis (Jerdon, 1849)<br />
Platycara Australis Jerdon, 1849: 333 (type locality: India:<br />
Nilgiris: "the small mountain stream that passes close by<br />
the bungalow in the Walliar jungle"; holotype: LU; adjective,<br />
-is, -is, -e)<br />
Bhavania annandalei Hora, 1920: 203, pl. 10 fig. 1 (type<br />
locality: India: Travancore: Tenmalai; holotype: ZSI<br />
F 2554/1; noun in genitive, indeclinable)<br />
7.4 Cryptotora Kottelat, 1998<br />
Cryptotora Kottelat, 1998b: 270 (type species: Homaloptera<br />
thamicola Kottelat, 1988c: 288, by original designation).<br />
Gender feminine.<br />
7.4.1 Cryptotora thamicola (Kottelat, 1988)<br />
Homaloptera thamicola Kottelat, 1988c: 228, fig. 4 (type<br />
locality: Thailand: Mae Hong Son Province: Tham Susa;<br />
19°28'N 98°08'E; holotype: AMS I.25987-001; noun in<br />
apposition, indeclinable)<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
'inhabitant <strong>of</strong>' are nouns and indeclinable.
7.5 Hemimyzon Regan, 1911<br />
Hemimyzon Regan, 1911: 32 (type species: Homaloptera<br />
formosana Boulenger, 1894b: 463, by original designation).<br />
Gender masculine.<br />
Dienbienia Nguyen & Nguyen, 2002: 9 (type species: Dienbienia<br />
namnuaensis Nguyen & Nguyen, 2002: 10, by<br />
original designation). Gender feminine.<br />
7.5.1 Hemimyzon confluens Kottelat, 2000<br />
Hemimyzon confluens Kottelat, 2000: 51, fig. 20 (type locality:<br />
Laos: Xiangkhouang Province: Nam Ngum, rapids<br />
downstream <strong>of</strong> Ban Latbouak; 19°36'28"N 103°14'23"E;<br />
holotype: ZRC 45317; participle, indeclinable, proposed<br />
as a noun in apposition)<br />
7.5.2 Hemimyzon ecdyonuroides Freyh<strong>of</strong> & Herder, 2002<br />
Hemimyzon ecdyonuroides Freyh<strong>of</strong> & Herder, 2002a: 54,<br />
figs. 1–2 (type locality: Vietnam [Mekong drainage, Sesan<br />
system]: Kontum Province: Pako River about 50 km<br />
north <strong>of</strong> Kontum; 14°39.60'N 107°46.98'E; holotype:<br />
ZFMK 22136; adjective, indeclinable)<br />
7.5.3 Hemimyzon elongatus (Chen & Li, in Li & Chen, 1985)<br />
Balitora elongata Chen & Li, in Li & Chen, 1985: 169, fig. 1<br />
(type locality: China: Yunnan: Yangbi River; holotype:<br />
KIZ 839072; adjective, -us, -a, -um)<br />
7.5.4 Hemimyzon formosanus (Boulenger, 1984)<br />
Homaloptera formosana Boulenger, 1894b: 463 (type locality:<br />
Central Taiwan; holotype: BMNH 1894.11.14.11,<br />
Eschmeyer & Fricke, 2010; adjective, -us, -a, -um)<br />
Homaloptera taiwanica Kishinouye, 1905: 177 (type locality:<br />
Taiwan; types: LU; adjective, -us, -a, -um)<br />
7.5.5 Hemimyzon khonensis Kottelat, 2000<br />
Hemimyzon khonensis Kottelat, 2000: 52, fig. 21 (type locality:<br />
Laos: Champasak Province: Mekong mainstream at<br />
Ban Hang Khone, below Khone Falls; holotype: ZRC<br />
45320; adjective, -is, -is, -e)<br />
7.5.6 Hemimyzon macropterus Zheng, in Zheng &<br />
Huang, 1982<br />
Hemimyzon macroptera Zheng, in Zheng, Chen & Huang,<br />
1982: 398, fig. 5 (type locality: China: Yunnan: Yiliang<br />
Xian [approx. 25°N 103°E]; syntypes: KIZ 773437 [1],<br />
IHB [773435–436 [2], DBJU 773434, 438 [2]; compound<br />
adjective, -us, -a, -um)<br />
7.5.7 Hemimyzon megalopseos Li & Chen, 1985<br />
Hemimyzon megalopseos Li & Chen, 1985: 170, fig. 2 (type<br />
locality: China: Yunnan: Yiliang; holotype: KIZ 774193;<br />
noun in apposition, indeclinable)<br />
7.5.8 Hemimyzon nanensis Doi & Kottelat, 1998<br />
Hemimyzon nanensis Doi & Kottelat, 1998: 8, fig. 1 (type<br />
locality: Thailand: Nan Province: Mae Nam Wa at Ban<br />
Nam Wa, Mae Nam Nan system, Chao Phraya drainage;<br />
holotype: NSMT P 36130; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
48<br />
7.5.9 Hemimyzon nujiangensis (Zhang & Zheng, in Zheng<br />
& Zhang, 1983)<br />
Balitora nujiangensis Zhang & Zheng, in Zheng & Zhang,<br />
1983: 66, fig. 1 (type locality: China: Yunnan: Nujiang<br />
[Salween River]: Liu-ku; holotype: IHB 81X4327; adjective,<br />
-is, -is, -e)<br />
7.5.10 Hemimyzon papilio Kottelat, 1998<br />
Hemimyzon papilio Kottelat, 1998a: 61, fig. 96 (type locality:<br />
Laos: Khammouan Province: Nam Theun, waterfall<br />
at 18°01'40"N 104°58'54"E; holotype: ZRC 41788; noun<br />
in apposition, indeclinable)<br />
7.5.11 Hemimyzon pengi (Huang, in Zheng, Chen &<br />
Huang, 1982)<br />
Balitora pengi Huang, in Zheng, Chen & Huang, 1982: 395,<br />
fig. 2 (type locality: China: Yunnan: Xishuangbanna:<br />
Menghai County; syntypes: KIZ 73046–53 [8]; noun in<br />
genitive, indeclinable)<br />
Dienbienia namnuaensis Nguyen & Nguyen, 2002: 10, fig. 1<br />
(type locality: Vietnam: Lai Chau Province: Dien Bien<br />
district: Nam Nua at Noongluong; holotype: NCNTTSI<br />
H.04.10.01.01; also spelt nammuaensis p. 14, an obvious<br />
inadvertent error, thus incorrect original spelling<br />
[Code art. 32.5.1]; adjective, -is, -is, -e)<br />
7.5.12 Hemimyzon pumilicorpora Zheng & Zhang, 1987<br />
Hemimyzon pumilicorpora Zheng & Zhang, 1987: 81, fig.<br />
(type locality: China: Guizhou: Anshun Prefecture: Huagguoshu<br />
[falls at 25°59'31"N 105°39'58"E; on Beihe River,<br />
Zhujiang drainage]; syntypes: IHB 83IV0046, 48, 54<br />
[3], DBJU 83IV0050–52 [3]; treated as a noun in apposition,<br />
indeclinable [correct spelling should have been<br />
pumilicorpus])<br />
7.5.13 Hemimyzon sheni Chen & Fang, 2009<br />
Hemimyzon sheni Chen & Fang, 2009: 186, fig. 2 (type locality:<br />
Taiwan: Taitung County: small tributary (3 km south<br />
to Yi-Ting mountain) in upper reaches <strong>of</strong> Tar-Ju River,<br />
Tar-Ren village; holotype: NTOU P-2007-07-077; noun<br />
in genitive, indeclinable)<br />
7.5.14 Hemimyzon taitungensis Tzeng & Shen, 1982<br />
Hemimyzon taitungensis Tzeng & Shen, 1982: 166, figs. 5–<br />
6 (type locality: Taiwan: Taitung County: Haiduan Township<br />
[23°06"N 121°11"E]: Shinwu-leu River, Lee-daou;<br />
holotype: NTUM 04941; adjective, -is, -is, -e)<br />
7.5.15 Hemimyzon tchangi (Zheng, in Zheng, Chen &<br />
Huang, 1982)<br />
Balitora tchangi Zheng, in Zheng, Chen & Huang, 1982:<br />
396, fig. 3 (type locality: China: Yunnan: Xishuangbanna:<br />
Jinghong Xian; holotype: IHB 78III0432; noun in<br />
genitive, indeclinable)<br />
7.5.16 Hemimyzon yaotanensis (Fang, 1931)<br />
? Gobius Tsin-ting-yu Dabry de Thiersant, 1872: 179, pl. 37<br />
fig. 9 (name not available, not binominal; locality: China:<br />
Sichuan: Leang-chan-hien)<br />
Sinohomaloptera yaotanensis Fang, 1931a: 137, fig. 1 (type
7.5.1 Hemimyzon confluens, CMK 22714, 52.0 mm SL; Laos:<br />
Mekong drainage: Nam Ngum.<br />
7.5.1 Hemimyzon confluens, CMK 22714, 52.0 mm SL.<br />
locality: China: Sichuan: Yao-tan, Wa-chang, Luchow;<br />
holotype: MMNHN 1888; adjective, -is, -is, -e)<br />
Sinohomaloptera yaotanensis acuticauda Fang, 1931a: 143,<br />
fig. 5 (type locality: China: Sichuan: Yao-tan, Wa-chang,<br />
Luchow; holotype: MMNHN 2249; compound noun, indeclinable)<br />
Taxonomic notes. Authors have considered Gobius tsin-tingyu<br />
as the fish later described as Psilorhynchus sinensis [now<br />
Jinshaia sinensis]. This seems unlikely as Dabry de Thiersant's<br />
figure does not show the typical body shape, the long<br />
caudal peduncle, the deeply forked caudal fin with longer<br />
lower lobe, and the large pectoral fins. Among the fishes<br />
now reported from Sichuan (Ding, 1994) it shows the greatest<br />
similarity to H. yaotanensis.<br />
7.6 Homaloptera van Hasselt, 1823<br />
Homaloptera van Hasselt, 1823: 133 (type species: Homaloptera<br />
ocellata van der Hoeven, 1830: 211, by subsequent<br />
monotypy in van der Hoeven, 1830: 211; also spelt<br />
Homalophra p. 132, first reviser [Kottelat, 1987b: 373]<br />
gave precedence to Homaloptera). Gender feminine.<br />
Helgia Vinciguerra, 1890: 328 (type species: Helgia bilineata<br />
Blyth, 1860: 172, by subsequent designation by<br />
Jordan, 1920: 448). Gender feminine.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
49<br />
7.6.7 Homaloptera orthogoniata, CMK 21835, 56.0 mm SL; Indonesia:<br />
Borneo: Mahakam drainage.<br />
7.6.5 Homaloptera ocellata, ZRC 43899, 67.3mm SL. (Photograph<br />
by Tan Heok Hui).<br />
7.6.1 Homaloptera bilineata Blyth, 1860<br />
Homoloptera [sic] bilineata Blyth, 1860: 172 (type locality:<br />
Burma: Tenasserim provinces; syntypes: ? ZSI, ? AMS<br />
[? possibly the syntypes <strong>of</strong> Nemacheilus serpentarius<br />
Day, 1870a: 551]; adjective, -us, -a, -um)<br />
Nemacheilus serpentarius Day, 1870a: 551 (type locality:<br />
unknown [Burma]; syntypes [3]: ZSI A.955 [2, lost],<br />
? AMS [lost ?], Whitehead & Talwar, 1976: 157, Ferraris<br />
et al., 2000: 301; adjective, -us, -a, -um)<br />
7.6.2 Homaloptera confuzona Kottelat, 2000<br />
Homaloptera confuzona Kottelat, 2000: 53, fig. 22 (type<br />
locality: Thailand: Trat Province: Khlong Fit at Ban<br />
Kraduk Chang, road 3157 from Trat to Borai, about<br />
2–3 km after junction with road 3271; 12°28'N 102°38'E;<br />
holotype: ZRC 45319; compound noun, indeclinable)<br />
7.6.3 'Homaloptera' menoni Shaji & Easa, 1995<br />
Homaloptera menoni Shaji & Easa, 1995: 395, fig. 1 (type<br />
locality: India: Kerala: Palghat District: Muthikulam<br />
Forest, Indekkuthodu in Siruvani, a tributary <strong>of</strong> Bhavani;<br />
holotype: KFRI 85; noun in genitive, indeclinable)<br />
7.6.4 'Homaloptera' montana Herre, 1945<br />
Homaloptera montana Herre, 1945: 400 (type locality: India:<br />
Tamil Nadu: brook on Puthutotam Estate in Anamallai<br />
Hills at about 3600 ft., Valparai Post Office; holotype:<br />
CAS-SU 39871, Böhlke, 1953: 40; adjective, -us,<br />
-a, -um)
7.7.4 Homalopteroides smithi, CMK 22142, 36.1 mm SL; Thailand:<br />
Phangnga.<br />
7.7.4 Homalopteroides smithi, CMK 22142, 35.3 mm SL.<br />
7.6.5 Homaloptera ocellata van der Hoeven, 1830<br />
Homaloptera ocellata van den Hoeven, 1830: 211, 1833:<br />
pl. 12 fig. 13 (type locality: Indonesia: Java; holotype:<br />
RMNH 2723, Roberts, 1993: 24; adjective, -us, -a, -um)<br />
Balitora erythrorhina Valenciennes, in Cuvier & Valenciennes,<br />
1846: 93, pl. 524 (type locality: Java: Buitenzorg<br />
[Bogor]; syntypes: MNHN 3121 [3], Bertin & Estève,<br />
1948: 100; compound adjective, -us, -a, -um)<br />
Balitora pavonina Valenciennes, in Cuvier & Valenciennes,<br />
1846: 97 (type locality: Java [probably from vicinity <strong>of</strong><br />
Bogor]; holotype: MNHN 3123, Bertin & Estève, 1948:<br />
100; adjective, -us, -a, -um)<br />
Homaloptera salusur Bleeker, 1853a: 161 (type locality:<br />
Indonesia: Java: Batavia [Jakarta] and Tjampea [Ciampea];<br />
lectotype: RMNH 5075, designated by Alfred, 1961:<br />
35; noun in apposition, indeclinable)<br />
Homaloptera polylepis Bleeker, 1853a: 162 (type locality:<br />
Indonesia: Java: Buitenzorg [Bogor] and Tjipannas<br />
[Cipanas]; syntypes [2]: among RMNH 7049 [13], Alfred,<br />
1961: 35; compound noun, indeclinable)<br />
7.6.6 Homaloptera ogilviei Alfred, 1967<br />
Homaloptera ogilviei Alfred, 1967: 587, fig. 1 (type locality:<br />
Malaysia: Negri Sembilan: Jelai River at 14 th mile<br />
along road from Tampin to Kuala Pilah; holotype: ZRC<br />
1555; noun in genitive, indeclinable)<br />
7.6.7 Homaloptera orthogoniata Vaillant, 1902<br />
Homaloptera orthogoniata Vaillant, 1902: 122, figs. 33–35<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
50<br />
(type locality: Indonesia: Borneo: Kalimantan Timur:<br />
Bloeoe River [Bluu, 0°42'N 114°24'E]; lectotype: RMNH<br />
7790A, designated by Tan & Ng, 2005a: 1 [abstract];<br />
adjective, -us, -a, -um)<br />
7.6.8 Homaloptera parclitella Tan & Ng, 2005<br />
Homaloptera parclitella Tan & Ng, 2005a: 7, figs. 5–6 (type<br />
locality: Malaysia: Terengganu basin: Sekayu waterfalls,<br />
rock pools about 5 minutes walk upstream <strong>of</strong> chalets [chalets:<br />
04°57.80'N 102°57.21'E]; holotype: ZRC 49257;<br />
compound noun, indeclinable)<br />
7.6.9 'Homaloptera' pillaii Indra & Remadevi, 1981<br />
Homaloptera pillaii Indra & Remadevi, 1981: 67, pl. 2 fig. 1<br />
(type locality: India: Kerala: Silent Valley: Kunthi River;<br />
ZSI/SRS 462, Menon, 1987: 228; noun in genitive,<br />
indeclinable)<br />
7.6.10 'Homaloptera' santhamparaiensis Arunachalam,<br />
Johnson & Rema Devi, 2002<br />
Homaloptera santhamparaiensis Arunachalam, Johnson &<br />
Rema Devi, 2002: 32, figs. 1–2 (type locality: India:<br />
Kerala: Idukki District; Panniyar stream, a tributary <strong>of</strong><br />
Periyar River, above Ponmudi Reservoir at Santhampari<br />
Hills (4 km from Pooparai Village), 9°82'N [sic] 77°15'E;<br />
holotype: ZSI/SRS F.5322; adjective, -is, -is, -e)<br />
7.6.11 'Homaloptera' silasi Kurup & Radhakrishnan, 2011<br />
Homaloptera silasi Kurup & Radhakrishnan, 2011: 224,<br />
fig. 1 (type locality: India: Kerala: Periyar Tiger Reserve:<br />
Kattamadithode, a small stream tributary <strong>of</strong> Periyar River,<br />
at Chokkanpetty; holotype: ZSI/WGR CLT/V/F13118;<br />
noun in genitive, indeclinable)<br />
7.7 Homalopteroides Fowler, 1905<br />
Homalopteroides Fowler, 1905a: 476 (type species: Homaloptera<br />
wassinkii Bleeker, 1853a: 163, by original designation).<br />
Gender masculine.<br />
Chopraia Prashad & Mukerji, 1929: 188 (type species: Chopraia<br />
rupicola Prashad & Mukerji, 1929: 188, by original<br />
designation). Gender feminine.<br />
7.7.1 Homalopteroides modestus (Vinciguerra, 1890)<br />
Helgia modesta Vinciguerra, 1890: 330, pl. 11 fig. 12 (type<br />
locality: Burma: Meekalan, Salween drainage; lectotype:<br />
MCSNG 15173-A, designated by Tortonese, 1961: 188;<br />
adjective, -us, -a, -um)<br />
7.7.2 Homalopteroides nebulosus (Alfred, 1969)<br />
Homaloptera nebulosa Alfred, 1969a: 227, pl. 1 figs. 3–4<br />
(type locality: Malaysia: Kelantan: Sok River, Kampong<br />
Sok; holotype: ZRC 2020; adjective, -us, -a, -um)<br />
7.7.3 Homalopteroides rupicola (Prashad & Mukerji, 1929)<br />
Chopraia rupicola Prashad & Mukerji, 1929: 188, pl. 8 fig. 3<br />
(type locality: Burma: Myitkyina District: small streams<br />
around Kamaing; holotype: ZSI F 10879/1, Menon &<br />
Yazdani, 1968: 118; noun in apposition, indeclinable)<br />
? Homaloptera manipurensis Arunkumar, 1999: 176, fig. 1
(type locality: India: Manipur: Chindwin drainage,<br />
Lokchao River near Moreh, 110 km from Imphal City;<br />
holotype: MUMF 3333/1A; adjective, -is, -is, -e)<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
'inhabitant <strong>of</strong>' are nouns and indeclinable.<br />
7.7.4 Homalopteroides smithi (Hora, 1932)<br />
Homaloptera smithi Hora, 1932a: 286, pl. 11 fig. 3 (type<br />
locality: Thailand: Nakon Sritamarat Province: Tadi<br />
Stream and Klong Pong at Ban Kiriwong; syntypes:<br />
KUMF 165 [3], USNM 107941 [1], 109821 [5],<br />
119459 [1], ZSI F 11293–11294/1 [4], 11295/1 [2], Menon<br />
& Yazdani, 1968: 118, Eschmeyer & Fricke, 2010;<br />
noun in genitive, indeclinable)<br />
? Homaloptera maxinae Fowler, 1937: 152, figs. 52–53 (type<br />
locality: Thailand: Tachin [Tha Chin, Samut Sakhon;<br />
13°32'22"N 100°15'20"E]; holotype: ANSP 68004; noun<br />
in genitive, indeclinable)<br />
Homaloptera lineata Smith, 1945: 277, fig. 55 (type locality:<br />
Thailand: Mekong at Chiangsen Kao; holotype:<br />
USNM 199488; adjective, -us, -a, -um)<br />
? Homaloptera indochinensis Silas, 1953: 192, fig. 2 (type<br />
locality: "Indo-China (? Tonkin)" [Vietnam: Kontum; see<br />
below]; holotype: BMNH 1933.8.19.50; adjective, -is,<br />
-is, -e)<br />
Taxonomic notes. Homaloptera indochinensis is based on<br />
a single specimen without precise locality. From Silas's description,<br />
it appears to fall within the variation <strong>of</strong> H. smithi,<br />
a very common species in the Mekong drainage. From information<br />
by E. Trewavas to S. L. Hora (in Silas, 1951: 11),<br />
the specimen had no locality label but Delacour & Lowe<br />
had collected in "Pekh, N. Tonkin [northern Vietnam; ? Pakha];<br />
Xien Khonang or Chin chuang, Laos [Xieng Khouang]<br />
and Kontum and Dak-to in Annam [Vietnam: Kontum and<br />
Dak To]". This material is registered as BMNH 1933.8.19.1–<br />
100. In fact this material had already been catalogued earlier<br />
as BMNH 1933.8.4.1–100. In this earlier list this specimen<br />
appears as H. wassinki, BMNH 1933.8.4.51, and the<br />
locality is listed as Kontum.<br />
Delacour conducted six expeditions to Laos, Cambodia<br />
and Vietnam between 1923 and 1939; Lowe accompanied<br />
him in five <strong>of</strong> these. There is no mention <strong>of</strong> fishes in the<br />
surviving Delacour notebooks (A. Hennache, pers. comm.),<br />
most <strong>of</strong> which disappeared in 1939. Hennache & Dickinson<br />
(2000) provided details <strong>of</strong> Delacour's expeditions, and from<br />
them it appears that Delacour and Lowe collected only once<br />
in Xieng Khouang, Kontum and Dakto before 1936. They<br />
collected at these localities during the second expedition in<br />
1925–1926 (see also Delacour et al., 1927). Additional fish<br />
material collected by Delacour and Lowe is in MNHN but<br />
were collected during the third (1926–1927) and fourth<br />
(1927–1929) expeditions, except for the types <strong>of</strong> Oreoglanis<br />
delacouri (MNHN 1936.31; see Ng & Kottelat, 1999:<br />
379), which must have come from the second expedition.<br />
No species <strong>of</strong> Homaloptera is known from the coastal<br />
streams on the eastern slope <strong>of</strong> the Annam Range and in the<br />
Red River drainage and this seems to eliminate Tonkin from<br />
the list <strong>of</strong> potential localities. Hennache & Dickinson (2000)<br />
mentioned two localities at which Delacour collected while<br />
in Xieng Khouang: Muongsoui [Muang Sui, nowaday Phu<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
51<br />
Kut, about 19°40'N 103°E in Nam Ngum drainage, a Mekong<br />
tributary] and Phu Ké, which is not on my maps, apparently<br />
Phu Keng, a hill about 18 km west-northwest <strong>of</strong> Phonsavan<br />
and also in Nam Ngum drainage. I conducted surveys in<br />
Xieng Khouang and the upper Nam Ngum in 1999 and 2012<br />
and have not seen H. smithi or any H. indochinensis-like<br />
fish. I also have not observed this species in northern Laos<br />
in surveys conducted in 1997, 1999 and 2007.<br />
Hennache & Dickinson (2000) did not provide details<br />
on the Kontum and Dakto localities. Both places are in the<br />
Sesan drainage, a tributary <strong>of</strong> the Mekong. The Sesan joins<br />
the Xe Kong shortly before entering the Mekong in Cambodia.<br />
Homaloptera smithi is a common species in the Xe Kong<br />
(pers. obs.). This makes Kontum or Dakto a likely type locality.<br />
7.7.5 Homalopteroides tweediei (Herre, 1940)<br />
Homaloptera tweediei Herre, 1940: 7, pl. 1 (type locality:<br />
Malaysia: Johore: Mawai District, shallow rapid creek,<br />
about 40 miles north <strong>of</strong> Singapore; holotype: CAS-SU<br />
33102, Böhlke, 1953: 40; noun in genitive, indeclinable)<br />
7.7.6 Homalopteroides wassinkii (Bleeker, 1853)<br />
Homaloptera fasciata van Hasselt, 1823: 133, 1824: 377<br />
(nomen nudum; Kottelat, 1987b: 373)<br />
Balitora ocellata Valenciennes, in Cuvier & Valenciennes,<br />
1846: 96 (type locality: Indonesia: Java: Buitenzorg<br />
[Bogor]; syntypes: MNHN 3122 [1] and specimen illustrated<br />
by Kuhl and van Hasselt [reproduced in Roberts,<br />
1993: fig. 27]; secondary junior homonym <strong>of</strong> Homaloptera<br />
ocellata van der Hoeven, 1830: 211 when placed<br />
in Homaloptera by Bleeker, 1860c: 95; adjective, -us,<br />
-a, -um)<br />
Homaloptera Wassinkii Bleeker, 1853a: 163 (type locality:<br />
Indonesia: Java: Tjampea [Ciampea] and Buitenzorg<br />
[Bogor]; lectotype: RMNH 4987, designated by Alfred,<br />
1961: 36; noun in genitive, indeclinable)<br />
Homaloptera valenciennesi Bleeker, 1860c: 95 (replacement<br />
name for Homaloptera ocellata Valenciennes, in Cuvier<br />
& Valenciennes, 1846: 96; noun in genitive, indeclinable)<br />
Homaloptera fasciata Bleeker, 1860c: 96 (unnecessary replacement<br />
name for H. wassinkii Bleeker, 1853a: 163;<br />
adjective, -us, -a, -um)<br />
Nomenclatural notes. Roberts (1993: 25) considered specimen<br />
MNHN 3122 as the holotype <strong>of</strong> Balitora ocellata. Valenciennes<br />
based his description on a single specimen and<br />
on the drawing sent by Kuhl and van Hasselt, as is obvious<br />
from the mention <strong>of</strong> the colour in life (p. 97) and explicit in<br />
the comment at the end <strong>of</strong> the description <strong>of</strong> B. pavonina (p.<br />
99). Unless it can be demonstrated that the specimen examined<br />
by Valenciennes is the same as figured by Kuhl and van<br />
Hasselt, there is no holotype but a series <strong>of</strong> syntypes.<br />
7.8 Homalopterula Fowler, 1940<br />
Homalopterula Fowler, 1940: 379 (type species: Homalopterula<br />
ripleyi Fowler, 1940: 379, by original designation).<br />
Gender feminine.
7.8.5 Homalopterula ripleyi, ZRC uncat., 42.9 mm SL; Indonesia:<br />
Sumatra: Aceh: Alas drainage. (Photograph by Tan Heok Hui).<br />
7.8.5 Homalopterula ripleyi, ZRC uncat., 42.9 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
7.8.1 Homalopterula amphisquamata (Weber & de Beaufort,<br />
1916)<br />
Homaloptera amphisquamata Weber & de Beaufort, 1916:<br />
12 (type locality: Indonesia: Sumatra: Sumatera Utara:<br />
Lau Borus [stream], discharging river <strong>of</strong> Lake Kawar,<br />
Karo Tableland Deli; syntypes [58]: ZMA 100.998 [1],<br />
100.994 [54], Nijssen et al., 1993: 214 [Weber & de<br />
Beaufort stated "type <strong>of</strong> the species in" ZMA; this is not<br />
a holotype designation as Code art. 73.1.1 requires that<br />
the authors state "that one specimen […] is the type"; the<br />
authors stated where the type is but not which <strong>of</strong> their<br />
specimens is the type, so this is not a holotype designation;<br />
therefore, all specimens are syntypes]; adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Tentatively placed in Homalopterula.<br />
7.8.2 Homalopterula gymnogaster (Bleeker, 1853)<br />
Homaloptera gymnogaster Bleeker, 1853a: 163 (type locality:<br />
Indonesia: Sumatra: Lake Meninju [Maninjau]; holotype:<br />
BMNH 1866.5.2.49, Alfred, 1961: 35; compound<br />
noun, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
52<br />
Homaloptera lepidogaster Weber & de Beaufort, 1916: 14<br />
(type locality: Sumatra: Sumatera Barat: Padang Highlands:<br />
Matur; syntypes: ZMA 100.256 [3], Nijssen et al.,<br />
1982: 29 [Weber & de Beaufort stated "type <strong>of</strong> the species<br />
in" ZMA; this is not a holotype designation as Code<br />
art. 73.1.1 requires that the authors state "that one specimen<br />
[…] is the type"; the authors stated where the type<br />
is, but not which <strong>of</strong> their specimens is the type, so this is<br />
not a holotype designation; in addition no specimen has<br />
been separated and can now be recognised as the holotype;<br />
therefore, all specimens are syntypes]; compound<br />
noun, indeclinable)<br />
Taxonomic notes. Tentatively placed in Homalopterula.<br />
7.8.3 Homalopterula heterolepis (Weber & de Beaufort,<br />
1916)<br />
Homaloptera heterolepis Weber & de Beaufort, 1916: 12<br />
(type locality: Indonesia: Sumatra: Aceh: Lake Tawar;<br />
syntypes [5]: ZMA 100.999 [3], ? AMNH 9263 [1], ? ZSI<br />
F 11035/1 [2], Nijssen et al., 1993: 214, Hora, 1950: 47,<br />
Silas, 1953: 198 [Weber & de Beaufort stated "type <strong>of</strong><br />
the species in" ZMA; this is not a holotype designation<br />
as Code art. 73.1.1 requires that the authors state "that<br />
one specimen […] is the type"; the authors stated where<br />
the type is but not which <strong>of</strong> their specimens is the type,<br />
so this is not a holotype designation; in addition no specimen<br />
has been separated and can now be recognised as<br />
the holotype; therefore, all specimens are syntypes]; compound<br />
noun, indeclinable)<br />
Taxonomic notes. Tentatively placed in Homalopterula.<br />
7.8.4 Homalopterula modiglianii (Perugia, 1893)<br />
Homaloptera Modiglianii Perugia, 1893a: 245 (type locality:<br />
Indonesia: Sumatra: Si Rambé; syntypes: MCSNG<br />
9249 [5], BMNH 1931.10.29.1–2 [2], ZSI F 11296/1 [1],<br />
Tortonese, 1961: 189, Silas, 1953: 196, Eschmeyer &<br />
Fricke, 2010; noun in genitive, indeclinable)<br />
Taxonomic notes. Tentatively placed in Homalopterula.<br />
7.8.5 Homalopterula ripleyi Fowler, 1940<br />
Homalopterula ripleyi Fowler, 1940: 379, figs. 5–7 (type<br />
locality: Indonesia: Sumatra: Aceh: Goempang River at<br />
Meloewak, 1640 feet; holotype: ANSP 68713, Ott, 2010:<br />
74, fig. 1; noun in genitive, indeclinable)<br />
7.8.6 Homalopterula vanderbilti (Fowler, 1940)<br />
Homaloptera vanderbilti Fowler, 1940: 375, figs. 1–2 (type<br />
locality: Indonesia: Sumatra: Aceh: Blangnanga in Tripa<br />
River, 3600 feet; holotype: ANSP 68688; noun in genitive,<br />
indeclinable)<br />
Homaloptera ulmeri Fowler, 1940: 377, figs. 3–4 (type locality:<br />
Indonesia: Sumatra: Aceh: Goempang River at Meloewak,<br />
1640 feet; holotype: ANSP 68700; simultaneous<br />
subjective synonym <strong>of</strong> Homaloptera vanderbilti Fowler,<br />
1940: 375, first reviser [Kottelat et al., 1993: 54] gave<br />
precedence to H. vanderbilti; noun in genitive, indeclinable)<br />
Taxonomic notes. Tentatively placed in Homalopterula.
7.9.3 Jinshaia sinensis, CMK 13067, 85.6 mm SL; China: Sichuan:<br />
Yangtze drainage: Yalong.<br />
7.9.3 Jinshaia sihnensis, CMK 13067, 85.6 mm SL.<br />
7.9 Jinshaia Kottelat & Chu, 1988<br />
Jinshaia Kottelat & Chu, 1988c: 191 (type species: Psilorhynchus<br />
sinensis Sauvage & Dabry de Thiersant, 1874:<br />
14, by original designation). Gender feminine.<br />
7.9.1 Jinshaia abbreviata (Günther, 1892)<br />
Homaloptera abbreviata Günther, 1892: 248, pl. 3 fig. B<br />
(type locality: China: Sichuan: a mountain stream running<br />
into Min River; holotype: BMNH 1891.6.13.40,<br />
Eschmeyer & Fricke, 2010; adjective, -us, -a, -um)<br />
7.9.2 ? Jinshaia niulanjiangensis Li, Mao & Lu, in Li,<br />
Mao, Lu, Sun & Lu, 1998<br />
Jinshaia niulanjiangensis Li, Mao & Lu, in Li, Mao, Lu,<br />
Sun & Lu, 1998: 1, fig. 1 (type locality: China: Yunnan:<br />
Zhanyi County: Niulan Jiang, tributary <strong>of</strong> Jinsha Jiang<br />
[Yangtze], at Deze Xiang [25°59'N 103°36'E]; holotype:<br />
HRAS or FACQR 9207001; also spelt niulanjingensis<br />
p. 3, an obvious inadvertent error, thus incorrect original<br />
spelling [Code art. 32.5.1]; adjective, -is, -is, -e)<br />
7.9.3 Jinshaia sinensis (Sauvage & Dabry de Thiersant,<br />
1874)<br />
Psilorhynchus sinensis Sauvage & Dabry de Thiersant, 1874:<br />
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7.10.2 Lepturichthys fimbriatus, CMK 13068, 104.2 mm SL;<br />
China: Sichuan: Yangtze drainage: Yalong.<br />
7.10.2 Lepturichthys fimbriatus, CMK 13068, 104.2 mm SL.<br />
14 (type locality: China: Sichuan: Teang-chan-hien; holotype:<br />
MNHN 7892, Bertin & Estève, 1948: 101; adjective,<br />
-is, -is, -e)<br />
7.10 Lepturichthys Regan, 1911<br />
Lepturichthys Regan, 1911: 31 (type species: Homaloptera<br />
fimbriata Günther, 1888: 433, by original designation).<br />
Gender masculine.<br />
7.10.1 Lepturichthys dolichopterus Dai, 1985<br />
Lepturichthys dolichopterus Dai, 1985: 221, figs. 1–2 (type<br />
locality: China: Fujian: Nanping [26°35'N 118°15'E],<br />
upper Min River; holotype: ASIZB 770428; compound<br />
adjective, -us, -a, -um)<br />
7.10.2 Lepturichthys fimbriatus (Günther, 1888)<br />
Homaloptera fimbriata Günther, 1888: 433 (type locality:<br />
China: Yangtsze-Kiang at Ichang [Hubei: Yangtze at<br />
Ychang, 30°43'N 111°17'N]; holotype: BMNH<br />
1888.5.15.40, Eschmeyer & Fricke, 2010; also in<br />
Günther, 1892: 248, pl. 3 fig. A; adjective, -us, -a, -um)<br />
Lepturichthys güntheri Hora, 1932a: 295, pl. 10 fig. 7 (type<br />
locality: China: Sichuan: mountain streams running into<br />
Min River; syntypes: BMNH 1891.6.13.41 [2]; spelling<br />
must be corrected as guentheri, Code art. 32.5.2.1; noun<br />
in genitive, indeclinable)<br />
Lepturichthys nicholsi Hora, 1932a: 297, pl. 10 fig. 8, pl. 12<br />
fig. 3 (type locality: China: Hunan: Tungting Lake; holotype:<br />
ZSI 11098/1; noun in genitive, indeclinable)
7.11.3 Metahomaloptera omeiensis, KIZ 795676, 54.0 mm SL;<br />
China: Sichuan: Yahan.<br />
7.11.3 Metahomaloptera omeiensis, KIZ 795676, 54.0 mm SL.<br />
7.11 Metahomaloptera Chang, 1944<br />
Metahomaloptera Chang, 1944: 54 (type species: Metahomaloptera<br />
omeiensis Chang, 1944: 54, by monotypy).<br />
Gender feminine.<br />
7.11.1 Metahomaloptera hangshuiensis Xie, Yang &<br />
Gong, 1984<br />
Metalohomaloptera [sic] omeiensis hangshuiensis Xie, Yang<br />
& Gong, 1984: 63, figs. 3–4 (type locality: China: Hubei:<br />
Yangriwan River [upper Hanjiang River], Mount Shengnongjia;<br />
syntypes: HACW 817022–33, 824268–285,<br />
825058–62 [35]; adjective, -is, -is, -e)<br />
7.11.2 Metahomaloptera longicauda Yang, Chen & Yang,<br />
2007<br />
Metahomaloptera longicauda Yang, Chen & Yang, 2007:<br />
64, fig. 1 (type locality: China: Yunnan: Zhanyi County:<br />
Yangtze drainage, Deze town, Chuanhedong village,<br />
25°54'09.4"N 103°34'49.5"E; holotype: KIZ 20060304;<br />
compound noun, indeclinable)<br />
7.11.3 Metahomaloptera omeiensis Chang, 1944<br />
Metahomaloptera omeiensis Chang, 1944: 54 (type locality:<br />
China: Sichuan: Loshan and Omei; syntypes: [LU]<br />
1701, 1702, 2552–2556, 2558, 2560, 2561 [10]; adjective,<br />
-is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
54<br />
7.12.1 Neohomaloptera johorensis, ZRC uncat., 21.3 mm SL; Indonesia:<br />
Borneo: Kahayan drainage. (Photograph by Tan Heok Hui).<br />
7.12 Neohomaloptera Herre, 1944<br />
Neohomaloptera Herre, 1944: 50 (subgenus <strong>of</strong> Homaloptera<br />
van Hasselt, 1823: 133; type species: Homaloptera<br />
johorensis Herre, 1944: 51, by original designation). Gender<br />
feminine.<br />
7.12.1 Neohomaloptera johorensis (Herre, 1944)<br />
Homaloptera johorensis Herre, 1944: 51 (type locality:<br />
Malaysia: Johor: brook near Simpang Rengam; holotype:<br />
CAS-SU 39840, Böhlke, 1953: 40; adjective, -is, -is, -e)<br />
7.13 Sinogastromyzon Fang, 1930<br />
Sinogastromyzon Fang, 1930a: 35 (type species: Sinogastromyzon<br />
wui Fang, 1930a: 36, by original designation).<br />
Gender masculine.<br />
Taxonomic notes. The validitidy <strong>of</strong> the many names created<br />
in recent years in the Vietnamese literature cannot be evaluated.<br />
The descriptions are <strong>of</strong> little use and the quality <strong>of</strong> the<br />
illustrations <strong>of</strong> most species does not allow to guess their<br />
identity. As most species have very restricted ranges, it is<br />
expected that those from widely distant localities might in<br />
fact be valid, while the many species described from exactly<br />
the same locality may be found to represent single species.<br />
7.13.1 Sinogastromyzon chapaensis Mai, 1978<br />
Sinogastromyzon chapaensis Mai, 1978: 220, fig. 102 (type<br />
locality: Vietnam [Lao Cai Province: Sapa; Kottelat,<br />
2001a: 101]; syntypes: DVZUT; adjective, -is, -is, -e)<br />
? Sinogastromyzon hexaocellum Nguyen, 2005: 612, fig. 39<br />
(type locality: Vietnam: Lai Chau Province: Phong Tho<br />
district: Nam So stream at Muong So [Song Da drainage];<br />
holotype: NCNTTSI; author indicated as "Hao &<br />
Duc", pp. 307, 612 fig. 39; treated as a noun in apposition,<br />
indeclinable)<br />
7.13.2 ? Sinogastromyzon daon Nguyen, 2005<br />
Sinogastromyzon daon Nguyen, 2005: 605, fig. 35 (type
locality: Vietnam: Lai Chau Province: Phong Tho district:<br />
Nam So stream [Song Da drainage]; holotype:<br />
NCNTTSI; author indicated as "Hao & Duc", pp. 306,<br />
605 fig. 35; treated as noun in apposition, indeclinable)<br />
Taxonomic notes. Liu et al. (2010: 35) considered this species<br />
as valid (in their key) but without discussion and without<br />
access to material.<br />
7.13.3 Sinogastromyzon dezeensis Li, Mao & Lu, in Li,<br />
Sun, Lu & Mao, 1999<br />
Sinogastromyzon dezeensis Li, Mao & Lu, in Li, Sun, Lu &<br />
Mao, 1999: 45, fig. 1 (type locality: China: Yunnan:<br />
Qujing prefecture: Zhanyi County: Niulan Jiang River at<br />
Deze Xiang [Deze village; 25°59'N 103°36'E] [Yangtze<br />
drainage]; holotype: HRAS or FACQR 9204011; adjective,<br />
-is, -is, -e)<br />
7.13.4 ? Sinogastromyzon hagiangensis Nguyen, 2005<br />
Sinogastromyzon hagiangensis Nguyen, 2005: 608, fig. 37<br />
(type locality: Vietnam: Ha Giang Province: Ha Giang town,<br />
Lo River; holotype: NCNTTSI; adjective, -is, -is, -e)<br />
Taxonomic notes. Liu et al. (2010: 35) considered this species<br />
as valid (in their key) but without discussion and without<br />
access to material.<br />
7.13.5 Sinogastromyzon hsiashiensis Fang, 1931<br />
Sinogastromyzon hsiashiensis Fang, 1931b: 48, fig. 3 (type<br />
locality: China: Kweichow [Guizhou]: Hsia-shih, Ma-hahsien;<br />
holotype: MMNHN 2998; adjective, -is, -is, -e)<br />
7.13.6 ? Sinogastromyzon hypercorpus Nguyen, 2005<br />
Sinogastromyzon hypercorpus Nguyen, 2005: 603, fig. 34<br />
(type locality: Vietnam: Lai Chau Province: Phong Tho<br />
district: Muong So, Nam So stream [Song Da drainage];<br />
holotype: NCNTTSI; also spelt hypercorfus p. 606, an<br />
obvious inadvertent error, thus incorrect original spelling<br />
[Code art. 32.5.1]; compound noun, indeclinable)<br />
Taxonomic notes. Liu et al. (2010: 35) considered this species<br />
as valid (in their key) but without discussion and without<br />
access to material.<br />
7.13.7 Sinogastromyzon lixianjiangensis Liu, Chen &<br />
Yang, 2010<br />
Sinogastromyzon lixianjiangensis Liu, Chen & Yang, 2010:<br />
26, fig. 1 (type locality: China: Yunnan: Mojiang County:<br />
Sinanjiang River (tributary <strong>of</strong> Lixianjiang, tributary<br />
<strong>of</strong> Red River); 23°07'38.13"N 101°47'44.32"E; holotype:<br />
KIZ 200401799; adjective, -is, -is, -e)<br />
7.13.8 Sinogastromyzon macrostoma Liu, Chen & Yang,<br />
2010<br />
Sinogastromyzon macrostoma Liu, Chen & Yang, 2010: 31<br />
(type locality: China: Yunnan: Mojiang County: Amojiang<br />
River, 23°02'47.44"N 101°46'49.48"E; holotype: KIZ<br />
200401820; compound noun, indeclinable)<br />
7.13.9 ? Sinogastromyzon maon Nguyen & Nguyen, in<br />
Nguyen, 2005<br />
Sinogastromyzon maon Nguyen [V. H.] & Nguyen [H. D.],<br />
in Nguyen [V. H.], 2005: 607, fig. 36 (type locality: Viet-<br />
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55<br />
7.13.15 Sinogastromyzon puliensis, CMK 17471, 69.4 mm SL;<br />
Taiwan: Wu drainage.<br />
7.13.15 Sinogastromyzon puliensis, CMK 17471, 69.4 mm SL.<br />
nam: Song La Province: Song Ma district: Ma River;<br />
holotype: NCNTTSI; treated as noun in apposition, indeclinable)<br />
Taxonomic notes. Liu et al. (2010: 35) considered this species<br />
as valid (in their key) but without discussion and without<br />
access to material.<br />
7.13.10 ? Sinogastromyzon minutus Mai, 1978<br />
Sinogastromyzon minutum Mai, 1978: 222, fig. 104 (type<br />
locality: Vietnam: Lai Chau Province: Muong Muon, Nam<br />
Muc; syntypes: DVZUT; adjective, -us, -a, -um)<br />
Taxonomic notes. Liu et al. (2010: 35) considered this species<br />
as valid (in their key) but without discussion and without<br />
access to material.<br />
7.13.11 Sinogastromyzon multiocellum Nguyen, 2005<br />
Sinogastromyzon multiocellum Nguyen, 2005: 614, fig. 40<br />
(type locality: Vietnam: Lai Chau Province: Phong Tho<br />
district: Nam So stream at Muong So [Song Da drainage];<br />
holotype: NCNTTSI; author indicated as "Hao &<br />
Duc" pp. 307, 614 fig. 40; also spelt multiocelum p. 613,<br />
an obvious inadvertent error, thus incorrect original spelling<br />
[Code art. 32.5.1]; treated as a noun in apposition,<br />
indeclinable)
7.14.1 Travancoria elongata, ZRC 34507, 98.8 mm SL, holotype;<br />
India: Kerala: Chalakudy River.<br />
7.14.1 Travancoria elongata, ZRC 34508, 79.3 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
Taxonomic notes. Considered valid following Liu et al.<br />
(2010: 35).<br />
7.13.12 Sinogastromyzon namnaensis Nguyen, 2005<br />
Sinogastromyzon namnaensis Nguyen, 2005: 610, fig. 38<br />
(type locality: Vietnam: Lai Chau Province: Lai Chau<br />
town, Nam Na River; holotype: NCNTTSI; adjective,<br />
-is, -is, -e)<br />
7.13.13 Sinogastromyzon nanpanjiangensis Li, 1987<br />
Sinogastromyzon nanpanjiangensis Li, 1987: 101, fig. 1<br />
(type locality: China: Yunnan: Nanpanjiang River in Lunan<br />
County [24°42'N 103°10'E]; holotype: KIZ 753010;<br />
adjective, -is, -is, -e)<br />
7.13.14 Sinogastromyzon nantaiensis Chen, Han & Fang,<br />
2002<br />
Sinogastromyzon nantaiensis Chen, Han & Fang, 2002: 240,<br />
fig. 1 (type locality: Taiwan: Pingtung County: Liukuei,<br />
Darjin, Kaoping River drainage, Laonon River; holotype:<br />
NMMB P 00465; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
56<br />
7.13.15 Sinogastromyzon puliensis Liang, 1974<br />
Sinogastromyzon puliensis Liang, 1974: 153, fig. 13 (type<br />
locality: Taiwan: Ta-tu-chi, Puli; holotype: NTUM 50280;<br />
adjective, -is, -is, -e)<br />
7.13.16 Sinogastromyzon rugocauda Mai, 1978<br />
Sinogastromyzon rugocauda Mai, 1978: 221, fig. 103 (type<br />
locality: Vietnam: Song La Province: Song Ma district:<br />
Nam Cong stream; syntypes: DVZUT; compound noun,<br />
indeclinable)<br />
7.13.17 Sinogastromyzon sichangensis Chang, 1944<br />
Sinogastromyzon sichangensis Chang, 1944: 53 (type locality:<br />
China: Sichuan: Anning River near Taihochang,<br />
Sichang; syntypes: [LU] 2086, 2094, 2095 [3]; adjective,<br />
-is, -is, -e)<br />
7.13.18 Sinogastromyzon szechuanensis Fang, 1930<br />
Sinogastromyzon szechuanensis Fang, 1930b: 99, fig. 1 (type<br />
locality: China: Szechuan [Sichuan]; holotype: SSCN<br />
6052; adjective, -is, -is, -e)<br />
7.13.19 Sinogastromyzon tonkinensis Pellegrin & Chevey,<br />
1935<br />
Sinogastromyzon tonkinensis Pellegrin & Chevey, 1935: 232,<br />
fig. 1 (type locality: Vietnam: Tonkin: Lai-Chau; holotype:<br />
MNHN 1935.41; adjective, -is, -is, -e)<br />
7.13.20 Sinogastromyzon wui Fang, 1930<br />
Sinogastromyzon wui Fang, 1930a: 36, pl. 2 figs. 8–9 (type<br />
locality: China: Guangxi: San-fang, Lo-ching-shien [Luocheng];<br />
MMNHN 515; noun in genitive, indeclinable)<br />
Sinogastromyzon intermedius Fang, 1931b: 54, fig. 7 (type<br />
locality: China: southwestern Guangxi: Tungkwei, Lungchow;<br />
holotype: MMNHN 1543; adjective, -us, -a, -um)<br />
Sinogastromyzon sanhoensis Fang, 1931b: 56, fig. 9 (type<br />
locality: China: South Kweichow [Guizhou]: San-hoshien;<br />
holotype: MMNHN 3481; adjective, -is, -is, -e)<br />
7.14 Travancoria Hora, 1941<br />
Travancoria Hora, 1941b: 228 (type species: Travancoria<br />
jonesi Hora, 1941b: 230, by original designation). Gender<br />
feminine.<br />
7.14.1 Travancoria elongata Pethiyagoda & Kottelat, 1994<br />
Travancoria elongata Pethiyagoda & Kottelat, 1994: 104,<br />
figs. 11–12 (type locality: India: Kerala: Chalakudy River,<br />
26 km upstream <strong>of</strong> Chalakudy town, near Vetilapara;<br />
holotype: ZRC 34507; adjective, -us, -a, -um)<br />
7.14.2 Travancoria jonesi Hora, 1941<br />
Travancoria jonesi Hora, 1941b: 230, pl. 8 figs. 5–6 (type<br />
locality: India: Travancore: streams within a radius <strong>of</strong> 5<br />
miles <strong>of</strong> Pampadampara, Peerumedu Taluq; holotype: ZSI<br />
F 13507/1, Menon & Yazdani, 1968: 119; noun in genitive,<br />
indeclinable)
8 Gastromyzontidae Fowler, 1905<br />
Gastromyzoninae Fowler, 1905: 477 (type genus: Gastromyzon<br />
Günther, 1874: 454; correct stem is Gastromyzont–<br />
and correct spelling is Gastromyzontinae)<br />
Lepidoglanidae Jordan, 1923: 149 (type genus: Lepidoglanis<br />
Vaillant, 1890: 82; correct stem is Lepidoglanid– and<br />
correct spelling is Lepidoglanididae)<br />
Crossostominae Silas, 1953: 219 (type genus: Crossostoma<br />
Sauvage, 1878a: 88; invalid because <strong>of</strong> junior homonymy<br />
<strong>of</strong> type genus, Code art. 39; correct stem is Crossostomat–<br />
and correct spelling is Crossostomatinae)<br />
Glaniopsini Silas, 1953: 259 (type genus: Glaniopsis Boulenger,<br />
1899a: 228)<br />
Pseudogastromyzoni Silas, 1953: 299 (type genus Pseudogastromyzon<br />
Nichols, 1925e: 1; correct stem is Pseudogastromyzont–<br />
and correct spelling is Pseudogastromyzontini)<br />
Beaufortini Chen, 1980b: 207 (type genus: Beaufortia Hora,<br />
1932a: 318; correct stem is Beauforti– and the correct<br />
spelling is Beaufortiini)<br />
Parhomalopterini Chen, 1980b: 207 (type genus: Parhomaloptera<br />
Vaillant, 1902: 129)<br />
Taxonomic notes. Authors have recognised and named various<br />
subdivisions within Gastromyzontidae. For example,<br />
Silas (1953: 219) recognised two subfamilies: Gastromyzontinae<br />
and Crossostomatinae. Chen (1980b: 207)<br />
recognised four tribes ("groups"): Parhomalopterini, Crossostomatini,<br />
Beaufortiini and Gastromyzontini. Gastromyzontini<br />
(the genera from Borneo) probably corresponds<br />
to a monophyletic lineage. Crossostomatini and<br />
Beaufortiini too, possibly correspond to monophyletic lineages.<br />
Parhomalopterini, with one genus in Borneo and the<br />
remaining ones in mainland Southeast Asia, is apparently<br />
an heterogeneous assemblage. It seems premature to recognise<br />
subfamilies or other divisions within Gastromyzontidae.<br />
8.1 Annamia Hora, 1932<br />
Annamia Hora, 1932a: 306 (type species: Parhomaloptera<br />
normani Hora, 1930: 584, by original designation). Gender<br />
feminine.<br />
Species inquirenda<br />
Annamia thuathienensis Nguyen [H. D.] & Nguyen [V. H.],<br />
in Nguyen [V. H.], 2005: 579, fig. 21 (type locality: Vietnam:<br />
Thua Thien Hue Province: Bo River [a small coastal<br />
drainage]; holotype: HNUE; adjective, -is, -is, -e)<br />
8.1.1 Annamia normani (Hora, 1930)<br />
Parhomaloptera normani Hora, 1930: 584, pl. 15 (type locality:<br />
Vietnam: Annam: Kontum [14°23'N 107°59'E; Se-<br />
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Family GASTROMYZONTIDAE<br />
57<br />
san drainage, Mekong basin]; holotype: BMNH<br />
1930.5.5.1, Eschmeyer & Fricke, 2010; noun in genitive,<br />
indeclinable)<br />
8.2 Beaufortia Hora, 1932<br />
Beaufortia Hora, 1932: 318 (type species: Gastromyzon leveretti<br />
Nichols & Pope, 1927: 340, by original designation).<br />
Gender feminine.<br />
Taxonomic notes. The validitidy <strong>of</strong> the many names created<br />
in recent years in the Vietnamese literature cannot be evaluated.<br />
The descriptions are <strong>of</strong> little use and the quality <strong>of</strong> the<br />
illustrations <strong>of</strong> most species does not allow to guess their<br />
identity. As most species have very restricted ranges, it is<br />
expected that those from widely distant localities might in<br />
fact be valid, while the many species described from exactly<br />
the same locality may be found to represent single species.<br />
For some species, however, there is not even informative<br />
locality data.<br />
8.2.1 ? Beaufortia buas (Mai, 1978)<br />
Gastromyzon buas Mai, 1978: 215, fig. 99 (type locality:<br />
northern Vietnam [Bua River, Song La and Phu Tho provinces;<br />
Kottelat, 2001a: 98]; syntypes: DVZUT; treated<br />
as noun in apposition, indeclinable)<br />
8.2.2 Beaufortia cyclica Chen, 1980<br />
Beaufortia cyclica Chen, 1980a: 115, fig. 6 (type locality:<br />
China: Guangxi: West River in Longzhow Xian; holotype:<br />
IHB 75IV1417; adjective, -us, -a, -um)<br />
Beaufortia polliciformis Dai, in Zheng, 1981: 172, fig. 142<br />
(type locality: China: Guangxi: Longzhou County: Dong<br />
Gui River; holotype: ASIZB 751842; adjective, -is, -is, -e)<br />
Taxonomic notes. Possibly a junior synonym <strong>of</strong> B. elongata.<br />
8.2.3 Beaufortia daon (Mai, 1978)<br />
Gastromyzon daon Mai, 1978: 216, fig. 100 (type locality:<br />
Vietnam: Lai Chau Province: Phong Tho and Nam Na<br />
streams [Song Da drainage]; syntypes: DVZUT; treated<br />
as noun in apposition, indeclinable)<br />
8.2.4 Beaufortia elongata (Mai, 1978)<br />
Gastromyzon elongatus Mai, 1978: 213, fig. 98 (type locality:<br />
northern Vietnam; syntypes: DVZUT; adjective, -us,<br />
-a, -um)<br />
Taxonomic notes. No usable locality information. Apparently<br />
close to or a senior synonym <strong>of</strong> B. cyclica, known from<br />
the Pearl River drainage in Guangxi, China. Nguyen (2005:<br />
284) identified as B. elongata material from the Song Con<br />
River in Thai Nguyen Province, in the Red River drainage.<br />
The identity <strong>of</strong> this species and its range remain to be determined.
8.1.1 Annamia normani, CMK 22291, 79.1 mm SL; Laos: Mekong<br />
drainage: Xe Kon g.<br />
8.1.1 Annamia normani, CMK 15802, 69.5 mm SL.<br />
8.2.5 Beaufortia huangguoshuensis Zheng & Zhang, 1987<br />
Beaufortia huangguoshuensis Zheng & Zhang, 1987: 80<br />
(type locality: China: Guizhou: Anshun prefecture: Shenning<br />
County: Dabang River, at upper part <strong>of</strong> Huagguoshu<br />
waterfall [25°59'31"N 105°39'58"E; on Baihe River],<br />
Zhujiang drainage; syntypes: IHB 83IV0049 [1],<br />
DBJU 83IV0053 [1]; adjective, -is, -is, -e)<br />
Nomenclatural notes. The description <strong>of</strong> B. huangguoshuensis<br />
first appears in Zheng & Zhang (1987: 80) who<br />
refer to the then 'in press' original description in 'The fishes<br />
<strong>of</strong> the Pearl River'. This description actually appeared later<br />
(Zheng, 1989: 259, fig. 198).<br />
8.2.6 Beaufortia intermedia Tang & Wang, in Tang, Wang<br />
& Yu, 1997<br />
Beaufortia intermedia Tang & Wang, in Tang, Wang & Yu,<br />
1997: 19, fig. 1 (type locality: China: Guizhou: Sandu<br />
County; 107°74'N [sic] 25°58'E; holotype: IHB 87IV512;<br />
adjective, -us, -a, -um)<br />
8.2.7 Beaufortia kweichowensis (Fang, 1931)<br />
Gastromyzon leveretti kweichowensis Fang, 1931b: 41, fig. 1<br />
(type locality: China: Kweichow [Guizhou]: San-ho<br />
Hsien; holotype: MMNHN 3539; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
58<br />
8.2.3 Beaufortia daon, CMK 15354, 45.9 mm SL; Laos: Houaphan<br />
Province: Nam Ma drainage.<br />
8.2.3 Beaufortia daon, CMK 15354, 45.9 mm SL.<br />
Beaufortia kweichowensis gracilicauda Chen & Zheng,<br />
1980: 97, fig. 8 (type locality: China: Guangdong: East<br />
River in Xinfeng Xian and North River in Shaoguan Shi;<br />
syntypes: IHB 76IV5880, 5881, 6986, 6987, 7736 [5],<br />
DBJU 76IV7742–7747 [6]; compound noun, indeclinable)<br />
8.2.8 Beaufortia leveretti (Nichols & Pope, 1927)<br />
Gastromyzon leveretti Nichols & Pope, 1927: 340, fig. 12<br />
(type locality: China: Hainan: Nodoa; holotype: AMNH<br />
8366; noun in genitive, indeclinable)<br />
8.2.9 Beaufortia liui Chang, 1944<br />
Beaufortia liui Chang, 1944: 55 (type locality: China: Sichuan:<br />
Loshan and Yaan; syntypes: [LU] 2247, 2446 [2];<br />
noun in genitive, indeclinable)<br />
8.2.10 ? Beaufortia loos (Mai, 1978)<br />
Gastromyzon loos Mai, 1978: 217, fig. 101 (type locality:<br />
Vietnam [Song Lo basin; Kottelat, 2001a: 99]; syntypes:<br />
DVZUT; treated as noun in apposition, indeclinable)<br />
8.2.11 Beaufortia niulanensis Chen, Huang & Yang, 2009<br />
Beaufortia niulanensis Chen, Huang & Yang, 2009: 639,
8.3.1 'Erromyzon' pachychilus, KIZ 200241775, 46.9 mm SL;<br />
China: Guangxi: Pearl River drainage: Dayaoshan Mountain. (Photograph<br />
by Yang Jian).<br />
8.3.1 'Erromyzon' pachychilus, ZRC 53444, about 20 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
figs. 1–2 (type locality: China: Yunnan: Zhanyi County:<br />
Deze town [25°59'N 103°36'E], middle Niulan Jiang<br />
River, Yangtze drainage, 26°01'N 103°37'E; holotype:<br />
KIZ 2006004660; adjective, -is, -is, -e)<br />
8.2.12 Beaufortia pingi (Fang, 1930)<br />
Gastromyzon pingi Fang, 1930a: 31, pl. 1 figs. 3–5 (type<br />
locality: China: Guangxi: stream in Lin-yueng-shien;<br />
holotype: MMNHN 909; noun in genitive, indeclinable)<br />
8.2.13 Beaufortia polylepis Chen, in Zheng, Chen & Huang,<br />
1982<br />
Beaufortia polylepis Chen, in Zheng, Chen & Huang, 1982:<br />
397, fig. 4 (type locality: China: Yunnan: Yiliang Xian<br />
[approx. 25°N 103°E]; syntypes: KIZ 774311, 315, 323,<br />
324, 326, 330, 338, 342, 343, 347 [10]; compound noun,<br />
indeclinable)<br />
8.2.14 Beaufortia szechuanensis (Fang, 1930)<br />
Gastromyzon szechuanensis Fang, 1930a: 36, fig. 7 (type<br />
locality: China: Sichuan: Omei-shien; holotype: SSCN<br />
2468; adjective, -is, -is, -e)<br />
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59<br />
8.3.2 Erromyzon compactus, ZRC 49636, 31.8 mm SL, holotype;<br />
Vietnam: Quang Ninh Province: Ba Che River.<br />
8.3.2 Erromyzon compactus, ZRC 49636, 31.8 mm SL.<br />
8.2.15 ? Beaufortia yunnanensis (Li, Lu & Mao, in Li,<br />
Mao, Lu, Sun & Lu, 1988)<br />
Paraprotomyzon yunnanensis Li, Lu & Mao, in Li, Mao,<br />
Lu, Sun & Lu, 1998: 4, fig. 21 (type locality: China: Yunnan:<br />
Luoping County: Huangni He (tributary <strong>of</strong> Nanpanjiang)<br />
at Neigei Cun; 24°20'N 104°30'E; holotype: HRAS<br />
or FACQR 9204001; adjective, -is, -is, -e)<br />
8.2.16 Beaufortia zebroida (Fang, 1930)<br />
Gastromyzon pingi zebroidus Fang, 1930a: 35, pl. 2 figs. 6–7<br />
(type locality: China: Guangxi: Tung-kwei [Donggui],<br />
Lung-chow [Longzhou] ["at the boundary between Kwangsi<br />
and Annam"; between Guangxi and northern Vietnam];<br />
holotype: MMNHN 1544; adjective, -us, -a, -um)<br />
? Beaufortia fasciolata Nguyen, 2005: 588, fig. 26 (type locality:<br />
Vietnam: Cao Bang Province: Bang Giang River at<br />
Cao Bang; holotype: NCNTTSI; adjective, -us, -a, -um)<br />
? Beaufortia triocellata Nguyen, 2005: 590, fig. 27 (type<br />
locality: Vietnam: Cao Bang Province: Bang Giang River<br />
at Cao Bang; holotype: NCNTTSI; adjective, -us, -a, -um)<br />
? Beaufortia multiocellata Nguyen, 2005: 592, fig. 28 (type<br />
locality: Vietnam: Cao Bang Province: Bang Giang Riv-
er at Cao Bang; holotype: NCNTTSI; also spelt multiocellatun<br />
p. 592, multicellata p. 690, obvious inadvertent<br />
errors, thus incorrect original spellings [Code art.<br />
32.5.1]; adjective, -us, -a, -um)<br />
8.3 Erromyzon Kottelat, 2004<br />
Erromyzon Kottelat, 2004c: 306 (type species: Protomyzon<br />
sinensis Chen, 1980a: 106, by original designation).<br />
Gender masculine.<br />
Species incertae sedis<br />
8.3.1 'Erromyzon' pachychilus (Chen, 1980)<br />
Protomyzon pachychilus Chen, 1980a: 106, fig. 3 (type locality:<br />
China: Guangxi: a mountain stream <strong>of</strong> West River;<br />
syntypes: IHB 7–9, 12–14, 18 [7]; compound adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Not congeneric with other species <strong>of</strong><br />
Erromyzon (Yang Jian, pers. comm.). Six syntypes seen in<br />
IHB. The "lip lamina" <strong>of</strong> Chen (1980a) (see Kottelat, 2004c)<br />
is apparently a flattened lateral, roundish expansion <strong>of</strong> the<br />
lower lip immediately in front <strong>of</strong> the maxillary barbel.<br />
8.3.2 Erromyzon compactus Kottelat, 2004<br />
Erromyzon compactus Kottelat, 2004c: 307, fig. 8 (type locality:<br />
Vietnam: Quang Ninh Province: Ba Che district: Ba<br />
Che River 4 km upstream <strong>of</strong> Ba Che city; 21°16'34"N 107°<br />
14'54"E; holotype: ZRC 49636; adjective, -us, -a, -um)<br />
8.3.3 Erromyzon sinensis (Chen, 1980)<br />
Protomyzon sinensis Chen, 1980a: 106, fig. 4 (type locality:<br />
China: Guangxi: West River in Lonsheng, Lipu and Jinxiu<br />
counties; syntypes [15]: IHB 75-IV-1801–1805, 2572–<br />
2574, 2579, 2581, 2804, 3217–3220 [15]; adjective, -is,<br />
-is, -e)<br />
8.3.4 Erromyzon yangi Neely, Conway & Mayden, 2007<br />
Erromyzon yangi Neely, Conway & Mayden, 2007: 98, fig. 1<br />
(type locality: China Guangxi: Jinciu County: Xi Jiang,<br />
tributary <strong>of</strong> Zhu Jiang (Pearl River), near Meicun,<br />
24°03.508'N 110°06.611'E; holotype: KIZ 200304423;<br />
noun in genitive, indeclinable)<br />
8.4 Formosania Oshima, 1919<br />
Crossostoma Sauvage, 1878a: 88 (type species: Crossostoma<br />
davidi Sauvage, 1878a: 89, by monotypy; junior homonym<br />
<strong>of</strong> Crossostoma Morris & Lycett, 1851: 72 in Gastropoda,<br />
Crossostoma Gosse, 1855: 310 in Polychaeta,<br />
and Crossostoma Agassiz, 1862: 154 in Medusae). Gender<br />
neuter.<br />
Formosania Oshima, 1919: 194 (type species: Formosania<br />
gilberti Oshima, 1919: 194, by original designation).<br />
Gender feminine.<br />
8.4.1 Formosania chenyiyui (Zheng, 1991)<br />
Crossostoma chenyiyui Zheng, 1991: 79, fig. 1 (type locality:<br />
China: Fujian: Chang Xian; syntypes: IHB 74VI1260–<br />
65 [6]; noun in genitive, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
60<br />
8.4.2 Formosania davidi (Sauvage, 1878)<br />
Crossostoma Davidi Sauvage, 1878a: 89 (type locality: China:<br />
western Fukien [Fujian] at 1200 masl [Fujian: Kuatun,<br />
27°51'N 117°48'E; see below]; syntypes: MNHN<br />
9817 [4], BMNH 1908.4.9.1 [1], Bertin & Estève, 1948:<br />
102, Eschmeyer & Fricke, 2010; noun in genitive, indeclinable)<br />
Crossostoma fascicauda foochowensis Tchang, 1932b: 123,<br />
fig. 3 (type locality: China: Fukien [Fujian]: Foochow;<br />
holotype: ZMFIB 6480; adjective, -is, -is, -e)<br />
Taxonomic notes. The locality <strong>of</strong> the syntypes <strong>of</strong> C. davidi<br />
is given in original description as "western Fukien" and collected<br />
by David in 1874. David (1875: map) was only once<br />
in Fukien [now Fujian] and only in a very restricted area<br />
near the village <strong>of</strong> Koaten along the border with Jiangsi.<br />
David (1875: 261) explicitly mentioned collecting fish that<br />
obviously are balitorids. The locality data in the MNHN<br />
database indicates an altitude <strong>of</strong> 3000 masl (vs. 1200 in the<br />
original description). David (1875: 284, 285) reported that<br />
Koaten is at 1200 masl, on the southern slope <strong>of</strong> a hill whose<br />
altitude he estimated to be 2200 masl based on his barometer,<br />
and that mountains around did not appear to exceed 3000<br />
masl. Koaten is identified as Kuatun and has been visited<br />
later by several naturalists (e.g.: La Touche, 1892: 402 (map),<br />
Pope, 1929: 339, 343, pl. 19, 1932: 490, pl. 111, 1935: 10,<br />
496; see also Rickett et al., 1896: 489; Slater, 1897: 169;<br />
Thomas, 1898: 769; Boulenger, 1899b: 159). Fooden (1983:<br />
16) gives the coordinates as 27°51'N 117°48'E, which is<br />
possibly for the summit <strong>of</strong> "Kuatun Mountain" (Wuyi Shan,<br />
27°51'34"N 117°47'00"E, 2158 masl [which, incidentally,<br />
shows the accuracy <strong>of</strong> David's mensurations]).<br />
8.4.3 Formosania fascicauda (Nichols, 1926)<br />
Crossostoma fascicauda Nichols, 1926: 2, fig. 2 (type locality:<br />
China: Fukien [Fujian]: Fuching Hsien; holotype:<br />
AMNH 8475; compound noun, indeclinable)<br />
8.4.4 Formosania fasciolata (Wang, Fan & Chen, 2006)<br />
Crossostoma fasciolatus Wang, Fan & Chen, 2006: 902, figs.<br />
1–2 (type locality: China: Zhejiang: Tiashun County<br />
[Taishun, 27°33'54"N 119°43'00"E]: rivulet in Wuyanling<br />
Nature Reserve, Liguan stream, Feiyun River system;<br />
holotype: ZMNH 8380338; adjective, -us, -a, -um)<br />
8.4.5 Formosania galericula (Zhang, in Zhang & Wang,<br />
2011)<br />
Crossostoma galericula Zhang, in Zhang & Wang, 2011: 85,<br />
fig. 1 (type locality: China: Zhejiang: Qingyuan County:<br />
Hehu township, Nanlu (mountain) stream (27°43'N<br />
19°16'E) in Baishanzu National Natural Protection Zone,<br />
in Oujiang drainage; holotype: Xiaoshan Institute <strong>of</strong><br />
Technology <strong>of</strong> Zhejiang Ocean University, Xiaoshan<br />
ZH 070616; treated as a noun in apposition, indeclinable)<br />
Nomenclatural notes. Although the paper is authored by<br />
Zhang & Wang and contains only the description <strong>of</strong> C. galericula,<br />
authorship <strong>of</strong> the new species is attributed to Zhang<br />
alone. Locality data are extracted from Chinese text and differ<br />
from English abstract. The etymology <strong>of</strong> galericula is<br />
not stated; the word exists in Latin and means wigs (plural),
it should have been in the singular (galericulum) or as an<br />
adjective (galericulatus, -a, -um).<br />
8.4.6 Formosania lacustris (Steindachner, 1908)<br />
Homaloptera formosanum Steindachner, 1908a: 86 (type<br />
locality: Taiwan: Lake Candidius; syntypes: NMW 47138<br />
[3], 48680 [3], 48682 [3], 48683 [3], 48690 [4]; junior<br />
primary homonym <strong>of</strong> Homaloptera formosana Boulenger,<br />
1894b: 463; adjective, -us, -a, -um)<br />
Crossostoma lacustre Steindachner, 1908b: 110 (replacement<br />
name for Homaloptera formosanum Steindachner, 1908a:<br />
86; adjective, -is, -is, -e)<br />
Formosania gilberti Oshima, 1919: 194, pl. 49 (type locality:<br />
Taiwan: Tamusui River near Shinten; lectotype:<br />
FMNH 59085, designated by Eschmeyer, 1998: 647, Ibarra<br />
& Stewart, 1987: 36; noun in genitive, indeclinable)<br />
8.4.7 Formosania paucisquama (Zheng, 1981)<br />
Crossostoma paucisquama Zheng, 1981: 57, fig. 1 (type<br />
locality: China: Guangdong: Puning County: mountain<br />
stream on Danan Shan [mountain], in Lianjiang drainage;<br />
syntypes: IHB 80VIII0037, 112, 114 [3], DBJU<br />
80VIII0035–36, 38–40, 111, 113, 115–116 [9]; compound<br />
noun, indeclinable)<br />
8.4.8 Formosania stigmata (Nichols, 1926)<br />
Crossostoma stigmata Nichols, 1926: 4, fig. 3 (type locality:<br />
China: Fukien [Fujian]; holotype: AMNH 8476; adjective,<br />
-us, -a, -um)<br />
8.4.9 Formosania tengi (Watanabe, 1983)<br />
Crossostoma tengi Watanabe, 1983: 111, fig. 5 (type locality:<br />
Taiwan: Kaoshiung County: Kaochung; holotype:<br />
WIRI 20; noun in genitive, indeclinable)<br />
8.4.10 Formosania tinkhami (Herre, 1934)<br />
Crossostoma tinkhami Herre, 1934: 286 (type locality: China:<br />
Guangdong: Loh Fau Shan, 3500 ft.; holotype:<br />
LNHSM; noun in genitive, indeclinable)<br />
8.5 Gastromyzon Günther, 1874<br />
Gastromyzon Günther, 1874: 454 (type species: Gastromyzon<br />
borneensis Günther, 1874: 454, by monotypy). Gender<br />
masculine.<br />
Lepidoglanis Vaillant, 1890: 82 (type species: Lepidoglanis<br />
monticola Vaillant, 1890: 82, by monotypy). Gender<br />
masculine.<br />
Nomenclatural notes. Vaillant (1890) did not indicate the<br />
gender <strong>of</strong> Lepidoglanis. He also did not explain the etymology<br />
and it is not known if he used the Greek glanis (feminine)<br />
or the Latin glanis (masculine). Because lepis is a Greek<br />
word, I consider Lepidoglanis as formed on the Greek glanis<br />
and feminine. The stem for a family group named formed<br />
on Lepidoglanis therefore is Lepidoglanid- (e.g. Lepidoglanididae<br />
Jordan, 1923: 149).<br />
8.5.1 Gastromyzon aequabilis Tan, 2006<br />
Gastromyzon aequabilis Tan, 2006: 147, fig. 91 (type locality:<br />
Malaysia: Borneo: Sabah: Lahad Datu, Danum Val-<br />
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61<br />
ley, Kuamut, unnamed stream at km 111 on main line<br />
west after turn<strong>of</strong>f to Borneo Rainforest Lodge,<br />
5°01'06.0"N 117°32'38.4"E; holotype: MUS uncat.; adjective,<br />
-is, -is, -e)<br />
8.5.2 Gastromyzon aeroides Tan & Sulaiman, 2006<br />
Gastromyzon aeroides Tan & Sulaiman, 2006: 9, figs. 3–4<br />
(type locality: Malaysia: Borneo: Sabah: Mengalong River<br />
drainage: Sipitang: Sungai Malamum, ca. 9 km into<br />
track, tributary to Mengalong River; 4°59.120'N<br />
115°37.581'E; holotype: MUS uncat.; noun in apposition,<br />
indeclinable)<br />
8.5.3 Gastromyzon auronigrus Tan, 2006<br />
Gastromyzon auronigrus Tan, 2006: 177, pl. 17B, fig. 114<br />
(type locality: Malaysia: Borneo: Sabah: Kota Marudu:<br />
Marak Parak: Sungai Kinarom, Kampong Loguhang, ca.<br />
6 km downstream <strong>of</strong> Serinsim Station, 6°19.733'N<br />
116°44.403'E; holotype: ZRC 47121; originally stated<br />
as an adjective [should have been auroniger], -us, -a, -um)<br />
8.5.4 Gastromyzon bario Tan, 2006<br />
Gastromyzon bario Tan, 2006: 76, figs. 34–36, pl. 11A (type<br />
locality: Malaysia: Borneo: Sarawak: Baram River drainage,<br />
Arur Dalan, headwater <strong>of</strong> Sungai Padapur, near Bario;<br />
holotype: IRSNB 824; noun in apposition, indeclinable)<br />
8.5.5 Gastromyzon borneensis Günther, 1874<br />
Gastromyzon borneensis Günther, 1874: 454 (type locality:<br />
Malaysia: Borneo: Sabah: Beaufort, sources <strong>of</strong> Mingalong<br />
River [near Labuan]; lectotype: BMNH<br />
1874.11.24.1, designated by Roberts, 1982: 502, Tan,<br />
2006: 56, fig. 12; adjective, -is, -is, -e)<br />
8.5.6 Gastromyzon contractus Roberts, 1982<br />
Gastromyzon contractus Roberts, 1982: 504, fig. 6 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Sungai<br />
Tebelian where it flows into Sungai Pinoh, 19 km upstream<br />
from Nangapinoh, 0°30'S 111°45'E; holotype:<br />
MZB 3447; adjective, -us, -a, -um)<br />
8.5.7 Gastromyzon cornusaccus Tan, 2006<br />
Gastromyzon cornusaccus Tan: 2006: 68, fig. 26, pl. 10B<br />
(type locality: Malaysia: Borneo: Sabah: Kota Marudu,<br />
Sungai Kinarom; holotype: UMSB 02233; noun in apposition,<br />
indeclinable)<br />
8.5.8 Gastromyzon cranbrooki Tan & Sulaiman, 2006<br />
Gastromyzon cranbrooki Tan & Sulaiman, 2006: 3, figs. 1–<br />
2 (type locality: Brunei Darussalam: Temburon District:<br />
Temburong River drainage, Sungai Belalong, in front and<br />
near Kuala Belalong Field Studies Centre, 4°32'50.4"N<br />
115°09'27.6"E; holotype: UBD uncat.; noun in genitive,<br />
indeclinable)<br />
8.5.9 Gastromyzon crenastus Tan & Leh, 2006<br />
Gastromyzon crenastus Tan & Leh, 2006: 9, figs. 3–4 (type<br />
locality: Malaysia: Borneo: Sarawak: Serian, Sungai<br />
Kubas, 6.9 km from Tebelu Tebakang turn<strong>of</strong>f, 5.8 km
8.4.6 Formosania lacustris, CMK 17450, 86.1 mm SL; Taiwan:<br />
Wu River.<br />
8.4.6 Formosania lacustris, CMK 17450, 86.1 mm SL.<br />
inside right side road, 1°09°10.0"N 110°29'22.7"E; holotype:<br />
SMK uncat.; adjective, -us, -a, -um)<br />
8.5.10 Gastromyzon ctenocephalus Roberts, 1982<br />
Gastromyzon ctenocephalus Roberts, 1982: 505, fig. 7 (type<br />
locality: Malaysia: Borneo: Sarawak: Senah; holotype:<br />
BMNH 1893.3.6.269; compound adjective, -us, -a, -um)<br />
8.5.11 Gastromyzon danumensis Chin & Inger, 1989<br />
Gastromyzon danumensis Chin & Inger, 1989: 54, fig. 2 (type<br />
locality: Malaysia: Borneo: Sabah: Lahad Datu District:<br />
Sungei Palum Tambun, a tributary <strong>of</strong> Segama River in<br />
Danum Valley Conservation Area; holotype: FMNH<br />
98126 [not 98125], Tan, 2006: 145, fig. 88; adjective,<br />
-is, -is, -e)<br />
8.5.12 Gastromyzon embalohensis Rachmatika, 1998<br />
Gastromyzon embalohensis Rachmatika, 1998: 652, fig. 1<br />
(type locality: Indonesia: Borneo: Kalimantan Barat:<br />
unnamed stream near its confluence with Sungai Tekelan,<br />
tributary <strong>of</strong> Embaloh River, "ca. 126'646"N<br />
11228'196"E" [? 1°26.646'N 112°28.196'E]; holotype:<br />
MZB 9205; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
62<br />
8.5.17 Gastromyzon introrsus, ZRC uncat., 49.6 mm SL; Malaysia:<br />
Borneo: Sabah: Ranau: Labuk. (Photograph by Tan Heok Hui).<br />
8.5.17 Gastromyzon introrsus, ZRC 47002, 45.7 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
8.5.13 Gastromyzon extrorsus Tan, 2006<br />
Gastromyzon extrorsus Tan, 2006: 70, figs. 28–29, pl. 10C<br />
(type locality: Malaysia: Borneo: Sabah: Penampang:<br />
Petagas drainage, Penampang River, Sungai Moyog,<br />
Kampong Kibunut, 05°53.301'N 116°14.102'E; holotype:<br />
MUS uncat.; treated as a noun in apposition, indeclinable)<br />
8.5.14 Gastromyzon farragus Tan & Leh, 2006<br />
Gastromyzon farragus Tan & Leh, 2006: 15, figs. 5–6 (type<br />
locality: Malaysia: Borneo: Sarawak: Serian, Sungai<br />
Kubas, 6.9 km from Tebelu Tebakang turn<strong>of</strong>f, 5.8 km<br />
inside right side road, 1°09°10.0'N 110°29°22.7'E; holotype:<br />
SMK uncat.; adjective, -us, -a, -um)<br />
8.5.15 Gastromyzon fasciatus Inger & Chin, 1961<br />
Gastromyzon fasciatus Inger & Chin, 1961: 173 (type locality:<br />
Malaysia: Borneo: Sarawak: Third Division: Sungai<br />
Dapu, tributary <strong>of</strong> Baleh River near mouth <strong>of</strong> Sungai<br />
Putai, 1°48'N 113°45'E; holotype: FMNH 68119, Tan,<br />
2006: 94, fig. 48; adjective, -us, -a, -um)<br />
8.5.16 Gastromyzon ingeri Tan, 2006<br />
Gastromyzon ingeri Tan, 2006: 149, fig. 94 (type locality:<br />
Malaysia: Borneo: Sabah: Tawau: Tawau Hill Parl, Tawau
River; holotype: FMNH [ex 108270]; noun in genitive,<br />
indeclinable)<br />
8.5.17 Gastromyzon introrsus Tan, 2006<br />
Gastromyzon introrsus Tan, 2006: 72, figs. 31–32, pl. 10D<br />
(type locality: Malaysia: Borneo: Sabah: Keningau: Sungai<br />
Agudon, mile 1 from Keningau to Crocker Range<br />
National Park HQ, Padas drainage, 5°21.206'N<br />
116°07.532'E; holotype: MUS uncat.; treated as a noun<br />
in apposition, indeclinable)<br />
8.5.18 Gastromyzon katibasensis Leh & Chai, 2003<br />
Gastromyzon katibasensis Leh & Chai, 2003: 277, fig. 1 (type<br />
locality: Malaysia: Borneo: Sarawak: Katibas, Song:<br />
Menyarin Camp [1°39.212'N 112°13.568'E] at confluence<br />
<strong>of</strong> Menyarin stream with Katibas headwaters; holotype:<br />
SMK FE.KTB.01–03; adjective, -is, -is, -e)<br />
8.5.19 Gastromyzon lepidogaster Roberts, 1982<br />
Gastromyzon lepidogaster Roberts, 1982: 509, figs. 9–10<br />
(type locality: Malaysia: Borneo: Sabah: Beaufort District:<br />
sources <strong>of</strong> Mengalong; holotype: BMNH<br />
1874.11.24.5; compound noun, indeclinable)<br />
8.5.20 Gastromyzon megalepis Roberts, 1982<br />
Gastromyzon megalepis Roberts, 1982: 510, fig. 11 (type<br />
locality: Malaysia: Borneo: Sarawak: Third District: tributary<br />
<strong>of</strong> Baleh River between Sungai Entunau and Sungai<br />
Putai, Rajang drainage; holotype: FMNH 68126;<br />
compound noun, indeclinable)<br />
8.5.21 Gastromyzon monticola (Vaillant, 1890)<br />
Lepidoglanis monticola Vaillant, 1890: 82 (type locality:<br />
Malaysia: Borneo: Sabah: Mt. Kinabalu [headwaters <strong>of</strong><br />
Tampassuk River; Tan, 2006: 58]; lectotype: MNHN<br />
1889-84, designated by Tan, 2006: 61, fig. 17; also in<br />
Vaillant, 1893: pl. 1 fig. 3; noun in apposition, indeclinable)<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
'inhabitant <strong>of</strong>' are nouns and indeclinable.<br />
8.5.22 Gastromyzon ocellatus Tan & Ng, 2004<br />
Gastromyzon ocellatus Tan & Ng, 2004: 269, figs. 1, 5 (type<br />
locality: Malaysia: Borneo: Sarawak: Bau, Serikin area,<br />
Sungai Petiak, 1°21.25'N 110°06.81'E; holotype: SBC<br />
uncat.; adjective, -us, -a, -um)<br />
8.5.23 Gastromyzon ornaticauda Tan & Martin-Smith, 1998<br />
Gastromyzon ornaticauda Tan & Martin-Smith, 1998: 367,<br />
figs. 3–4 (type locality: Malaysia: Borneo: Sabah: Lahad<br />
Datu: Kuamut, unnamed stream at km 113 on Main<br />
Line West logging track, 5°00'40"N 117°31'40"E; holotype:<br />
MUS uncat.; compound noun, indeclinable)<br />
8.5.24 Gastromyzon pariclavis Tan & Martin-Smith, 1998<br />
Gastromyzon pariclavis Tan & Martin-Smith, 1998: 362,<br />
fig. 1 (type locality: Malaysia: Borneo: Sabah: Lahad<br />
Datu: Kuamut, unnamed stream at km 111 on Main Line<br />
West logging track, 5°01'05"N 117°32'40"E; holotype:<br />
MUS uncat.; compound noun, indeclinable)<br />
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8.6.3 Glaniopsis cf. hanitschi, ZRC uncat., 55.5 mm SL; Malaysia:<br />
Borneo: Sabah: Ranau: Labuk. (Photograph by Tan Heok Hui).<br />
8.6.3. Glaniopsis hanitschi, ZRC uncat., 40.4 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
8.5.25 Gastromyzon praestans Tan, 2006<br />
Gastromyzon praestans Tan, 2006: 96, figs. 51–52 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: mainstream<br />
<strong>of</strong> Sungai Pinoh 20–60 km upstream from Nangapinoh,<br />
0°27.5'–0°41.5'S 111°39–111°45.5'E; holotype:<br />
MZB 3450; treated as a noun in apposition [but in fact<br />
an adjective], indeclinable)<br />
8.5.26 Gastromyzon psiloetron Tan, 2006<br />
Gastromyzon psiloetron Tan, 2006: 126, pl. 14A, fig. 73 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Kayan<br />
drainage, Bahau: Lalut Birai River next to Lalut Birai<br />
field station, tributary to Enggeng Bio, itself draining to<br />
Bahau River, 2°52.58'N 115°49.19'E; holotype: MZB<br />
9348; noun in apposition, indeclinable)<br />
8.5.27 Gastromyzon punctulatus Inger & Chin, 1961<br />
Gastromyzon punctulatus Inger & Chin, 1961: 173 (type<br />
locality: Malaysia: Borneo: Sarawak: Third Division:<br />
Sungai Dapu, tributary <strong>of</strong> Baleh River near mouth <strong>of</strong><br />
Sungai Putai, 1°48'N 113°45'E; holotype: FMNH 68116,<br />
Tan, 2006: 84, fig. 40; adjective, -us, -a, -um)
8.5.28 Gastromyzon ridens Roberts, 1982<br />
Gastromyzon ridens Roberts, 1982: 515, fig. 14 (type locality:<br />
Indonesia: Borneo: Kalimantan Barat: Sungai Pinoh<br />
20–60 km upstream from Nangapinoh; holotype: MZB<br />
3455; participle, indeclinable)<br />
8.5.29 Gastromyzon russulus Tan, 2006<br />
Gastromyzon russulus Tan, 2006: 163, pl. 16B, fig. 105 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Kayan<br />
drainage, Bahau: Enggeng Bio River, draining to Bahau<br />
River, up to 6 riffles upstream <strong>of</strong> field station, 2°52.58'N<br />
115°49.19'E; holotype: MZB 10702; adjective, -us, -a,<br />
-um)<br />
8.5.30 Gastromyzon scitulus Tan & Leh, 2006<br />
Gastromyzon scitulus Tan & Leh, 2006: 3, figs. 1–2 (type<br />
locality: Malaysia: Borneo: Sarawak: Serian, Sungai<br />
Kubas, 6.9 km from Tebelu Tebakang turn<strong>of</strong>f, 5.8 km<br />
inside right side road, 1°09'10.0"N 110°29'22.7"E; holotype:<br />
SMK uncat.; adjective, -us, -a, -um)<br />
8.5.31 Gastromyzon spectabilis Tan, 2006<br />
Gastromyzon spectabilis Tan, 2006: 161, pl. 16A, fig. 102<br />
(type locality: Malaysia: Borneo: Sabah: Lahad Datu,<br />
Sungai Danum camp II; holotype: MUS 0366; adjective,<br />
-is, -is, -e)<br />
8.5.32 Gastromyzon stellatus Tan, 2006<br />
Gastromyzon stellatus Tan, 2006: 136, pl. 14C, figs. 82 (type<br />
locality: Malaysia: Borneo: Sarawak: Bau, Serikin area,<br />
Sungai Petiak, 1°21.25'N 110°06.81'E; holotype: SMK<br />
uncat.; adjective, -us, -a, -um)<br />
8.5.33 Gastromyzon umbrus Tan, 2006<br />
Gastromyzon umbrus Tan, 2006: 106, figs. 60–61 (type locality:<br />
Indonesia: Borneo: Kalimantan Timur: Sungai Sebuku<br />
drainage: Sungai Bantul at Bantul logging camp,<br />
draining to Sungai Tulit, 4°08'54"N 116°48'18"E; holotype:<br />
MZB 16471; used as an adjective, -us, -a, -um)<br />
8.5.34 Gastromyzon venustus Tan & Sulaiman, 2006<br />
Gastromyzon venustus Tan & Sulaiman, 2006: 13, figs. 5–6<br />
(type locality: Brunei Darussalam: Temburong District:<br />
Temburong River drainage, Sungai Belalong, in front and<br />
near Kuala Belalong Field Studies Centre, 4°32'50.4"N<br />
115°09'27.6"E; holotype: UBD uncat.; adjective, -us,<br />
-a, -um)<br />
8.5.35 Gastromyzon viriosus Tan, 2006<br />
Gastromyzon viriosus Tan, 2006: 165, fig. 107 (type locality:<br />
Malaysia: Borneo: Sarawak: Bintulu, Tatau District,<br />
Sungai Sawi; holotype: ZRC 39709; adjective, -us, -a,<br />
-um)<br />
8.5.36 Gastromyzon zebrinus Tan, 2006<br />
Gastromyzon zebrinus Tan, 2006: 139, pl. 15A, fig. 84 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Sambas<br />
drainage: Bengkayan, about 60–70 km from Sarawak<br />
border; holotype: MZB 9350; adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
64<br />
8.6 Glaniopsis Boulenger, 1899<br />
Glaniopsis Boulenger, 1899a: 228 (type species: Glaniopsis<br />
hanitschi Boulenger, 1899a: 228, by monotypy). Gender<br />
feminine.<br />
8.6.1 Glaniopsis denudata Roberts, 1982<br />
Glaniopsis denudata Roberts, 1982: 517, fig. 17 (type locality:<br />
Malaysia: Borneo: Sabah: Sungai Kidikarok; holotype:<br />
BMNH 1957.2.27.1; adjective, -us, -a, -um)<br />
8.6.2 Glaniopsis gossei Roberts, 1982<br />
Glaniopsis gossei Roberts, 1982: 518, fig. 18 (type locality:<br />
Malaysia: Borneo: Sarawak: Arur Dalan, a torrential<br />
headwater <strong>of</strong> Sungai Padapur, Baram drainage, near Bario;<br />
holotype: IRNSB 621; noun in genitive, indeclinable)<br />
8.6.3 Glaniopsis hanitschi Boulenger, 1899<br />
Glaniopsis Hanitschi Boulenger, 1899a: 228 (type locality:<br />
Malaysia: Borneo: Sabah: Kadamaian River on Mount<br />
Kinabalu; syntypes: BMNH 1898.8.19.17–18 [2], ZRC<br />
1753 [1], Alfred, 1970: 69, Eschmeyer & Fricke, 2010;<br />
noun in genitive, indeclinable)<br />
8.6.4 Glaniopsis multiradiata Roberts, 1982<br />
Glaniopsis multiradiata Roberts, 1982: 521, fig. 20 (type<br />
locality: Malaysia: Borneo: Sarawak: Arur Dalan, a torrential<br />
headwater <strong>of</strong> Sungai Padapur, Baram drainage,<br />
near Bario; holotype: IRNSB 619; adjective, -us, -a,<br />
-um)<br />
8.7 Hypergastromyzon Roberts, 1989<br />
Hypergastromyzon Roberts, 1989: 91 (type species: Hypergastromyzon<br />
humilis Roberts, 1989: 92, by original designation;<br />
also spelt Hypogastromyzon p. 83, an inadvertent<br />
error as evidenced by etymology p. 92, thus incorrect<br />
original spelling [Code art. 32.5.1]). Gender masculine.<br />
8.7.1 Hypergastromyzon eubranchus Roberts, 1991<br />
Hypergastromyzon eubranchus Roberts, 1991: 334, fig. 1<br />
(type locality: Malaysia: Borneo: Sarawak: Batang Ai<br />
near Wong Mepal; holotype: BMNH 1984.11.15.1; compound<br />
noun, indeclinable)<br />
8.7.2 Hypergastromyzon humilis Roberts, 1989<br />
Hypergastromyzon humilis Roberts, 1989: 92, fig. 72 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Sungei<br />
Tamang, a tributary <strong>of</strong> Sungei Pinoh entering it opposite<br />
mouth <strong>of</strong> Sungei Kelawai; holotype: MZB 3480; adjective,<br />
-is, -is, -e)<br />
8.8 Katibasia Kottelat, 2004<br />
Katibasia Kottelat, 2004c: 302 (type species: Katibasia insidiosa<br />
Kottelat, 2004c: 303, by original designation).<br />
Gender feminine.<br />
8.8.1 Katibasia insidiosa Kottelat, 2004<br />
Katibasia insidiosa Kottelat, 2004c: 303, fig. 2 (type locality:<br />
Malaysia: Borneo: Sarawak: Song District: Sungai
8.7.2 Hypergastromyzon humilis, CMK 10574, 30.4 mm SL; Indonesia:<br />
Borneo: Kapuas drainage.<br />
8.7.2 Hypergastromyzon humilis, CMK 10574, 31.5 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
Melinau, Ulu Katibas, Rajang drainage; holotype: ZRC<br />
49631; adjective, -us, -a, -um)<br />
8.9 Liniparhomaloptera Fang, 1935<br />
Liniparhomaloptera Fang, 1935a: 93 (type species: Parhomaloptera<br />
disparis Lin, 1934: 225, by original designation).<br />
Gender feminine.<br />
8.9.1 Liniparhomaloptera disparis (Lin, 1934)<br />
Parhomaloptera disparis Lin, 1934: 225, figs. 1–3 (type<br />
locality: China: Guangdong: Poh-lo County: Loh Fau<br />
Shan; holotype: FESC H 15; adjective, -ar, -aris, -are)<br />
8.9.2 ? Liniparhomaloptera monoloba (Mai, 1978)<br />
Homaloptera monoloba Mai, 1978: 207, fig. 94 (type locality:<br />
Vietnam: Bac Thai Province: Ky Phu stream; syntypes:<br />
DVZUT; compound noun, indeclinable)<br />
8.9.3 Liniparhomaloptera obtusirostris Zheng & Chen,<br />
1980<br />
Liniparhomaloptera obtusirostris Zheng & Chen, 1980: 92,<br />
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8.8.1 Katibasia insidiosa, ZRC 49631, 31.2 mm SL, holotype;<br />
Malaysia: Borneo: Sarawak: Rajang drainage. (Photograph by Tan<br />
Heok Hui).<br />
8.8.1 Katibasia insidiosa, ZRC 49631, 31.2 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
fig. 3 (type locality: China: Guangdong: Xinyi County: a<br />
mountain stream <strong>of</strong> West River; syntypes: IHB 77VII80,<br />
83–84, 86–87 [5], DBJU 77VIII88, 106, 111 [3]; compound<br />
noun, indeclinable)<br />
8.9.4 Liniparhomaloptera qiongzhongensis Zheng &<br />
Chen, 1980<br />
Liniparhomaloptera disparis qiongzhongensis Zheng &<br />
Chen, 1980: 91, fig. 2 (type locality: China: Hainan:<br />
Wanquan River in Quiongzhong Xian; syntypes: IHB<br />
76V9621–9624, 9626 [5], DBJU 76V9629, 9630, 9632,<br />
9634 [4]; also spelt qiongzhengensis p. 89, an obvious<br />
inadvertent error, thus incorrect original spelling [Code<br />
art. 32.5.1]; adjective, -is, -is, -e)<br />
8.10 Neogastromyzon Popta, 1905<br />
Neogastromyzon Popta, 1905: 180 (type species: Neogastromyzon<br />
nieuwenhuisii Popta, 1905: 181, by monotypy).<br />
Gender masculine.<br />
8.10.1 Neogastromyzon brunei Tan, 2006<br />
Neogastromyzon brunei Tan, 2006: 194, fig. 127, pl. 18C<br />
(type locality: Brunei Darussalam: Temburong District:<br />
Belalong drainage: Sungai Enkabang, about 15 minutes<br />
upstream <strong>of</strong> Kuala Belalong Field Studies Centre,<br />
4°32'13.5"N 115°09'35.0"E; holotype: UBD uncat.; noun<br />
in apposition, indeclinable)
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
8.9.4 Liniparhomaloptera cf. qiongzhongensis, CMK 14888, 72.1<br />
mm SL; Vietnam: Mong Cai. 8.10.4 Neogastromyzon kottelati, CMK 11706, 35.2 mm SL; Indonesia:<br />
Borneo: Kapuas drainage.<br />
8.9.4 Liniparhomaloptera cf. qiongzhongensis, CMK 14888, 72.1<br />
mm SL.<br />
8.10.2 Neogastromyzon chini Tan, 2006<br />
Neogastromyzon chini Tan, 2006: 189, figs. 122–123, pl.<br />
18A–B (type locality: Malaysia: Borneo: Sarawak: Third<br />
Division: Rajang drainage, tributary Baleh River, between<br />
Sungai Entunau and Sungai Putai; holotype: FMNH<br />
97439; noun in genitive, indeclinable)<br />
8.10.3 Neogastromyzon crassiobex Tan, 2006<br />
Neogastromyzon crassiobex Tan, 2006: 187, fig. 120, pl. 17C<br />
(type locality: Malaysia: Borneo: Sabah: Lahad Datu:<br />
Kuamut drainage, unnamed hill stream, 5°00'20"N<br />
117°30'10"E; holotype: ZRC 47396; noun in apposition,<br />
indeclinable)<br />
8.10.4 Neogastromyzon kottelati Tan, 2006<br />
Neogastromyzon kottelati Tan, 2006: 192, fig. 125 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Danau<br />
Sentarum area, Sungai Hulu Leboyan at Keluwin; holotype:<br />
MZB 10704; noun in genitive, indeclinable)<br />
8.10.5 Neogastromyzon nieuwenhuisii Popta, 1905<br />
Neogastromyzon Nieuwenhuisii Popta, 1905: 181 (type locality:<br />
Indonesia: Borneo: Kalimantan Timur: Howong River<br />
[about 0°15'N 115°30'E]; holotype: RMNH 7640; also<br />
66<br />
8.10.5 Neogastromyzon nieuwenhuisii, ZRC 47394, 55.2 mm SL;<br />
Indonesia: Borneo: Kayan drainage. (Photograph by Tan Heok<br />
Hui).<br />
in Popta, 1906: 192, pl. 10 fig. 41; noun in genitive, indeclinable)<br />
8.10.6 Neogastromyzon pauciradiatus (Inger & Chin,<br />
1961)<br />
Gastromyzon pauciradiatus Inger & Chin, 1961: 174 (type<br />
locality: Malaysia: Borneo: Sarawak: Third Division:<br />
unnamed tributary <strong>of</strong> Baleh River opposite Sungai Laie;<br />
holotype: FMNH 68121, Tan, 2006: 185, fig. 118; adjective,<br />
-us, -a, -um)<br />
8.11 Paraprotomyzon Pellegrin & Fang, 1935<br />
Paraprotomyzon Pellegrin & Fang, 1935: 99 (type species:<br />
Paraprotomyzon multifasciatus Pellegrin & Fang, 1935:<br />
103, by original designation). Gender masculine.<br />
8.11.1 Paraprotomyzon bamaensis Tang, 1997<br />
Paraprotomyzon bamaensis Tang, 1997: 108, fig. 1 (type<br />
locality: China: Guangxi: Bama County, in Hongshuihe<br />
River drainage; holotype: IHB 86VIII0001; adjective, -<br />
is, -is, -e)
8.11.3 Paraprotomyzon multifasciatus, KIZ 2011001333, 33.9 mm<br />
SL; China: Guangxi: Pearl River drainage. (Photograph by Chen<br />
Xiao-Yong).<br />
8.11.3 Paraprotomyzon multifasciatus, ZSI F669/2, syntype, size<br />
unknown. (From Hora & Jayaram, 1950: 63).<br />
8.11.2 Paraprotomyzon lungkowensis Xie, Yang & Gong,<br />
1984<br />
Paraprotomyzon lungkowensis Xie, Yang & Gong, 1984:<br />
62, figs. 1–2 (type locality: China: Hubei: Lungkow<br />
stream, Mount Shengnongjia; syntypes: HACW 826001,<br />
835274–278, 294–278 [sic], IHB 835279–281 [total 9];<br />
adjective, -is, -is, -e)<br />
8.11.3 Paraprotomyzon multifasciatus Pellegrin & Fang,<br />
1935<br />
Paraprotomyzon Multifasciatus Pellegrin & Fang, 1935: 103,<br />
fig. 2 (type locality: China: Sichuan: Kwai-chow; syntypes:<br />
MNHN 1935-36–39 [4], ZSI F 669/2 [1], Bertin & Estève,<br />
1948: 102, Silas, 1953: 231; adjective, -us, -a, -um)<br />
8.11.4 Paraprotomyzon niulanjiangensis Lu, Lu & Mao,<br />
2005<br />
Paraprotomyzon niulanjiangensis Lu, Lu & Mao, 2005: 202,<br />
fig. 1 (type locality: China: Yunnan: Zhanyi County: Niulan<br />
Jiang, tributary <strong>of</strong> Jinsha Jiang [Yangtze], at Deze<br />
Xiang, 25°59'N 103°36'E; holotype: FACQR 200306005;<br />
adjective, -is, -is, -e)<br />
8.12 Parhomaloptera Vaillant, 1902<br />
Parhomaloptera Vaillant, 1902: 129 (type species: Parhomaloptera<br />
obscura Vaillant, 1902: 130, by monotypy). Gender<br />
feminine.<br />
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8.12.1 Parhomaloptera microstoma, ZRC uncat., 63.4 mm SL; Indonesia:<br />
Borneo: Kayan drainage. (Photograph by Tan Heok Hui).<br />
8.12.1 Parhomaloptera microstoma, ZRC uncat., 56.6 mm SL.<br />
(Photograph by Tan Heok Hui).<br />
8.12.1 Parhomaloptera microstoma (Boulenger, 1899)<br />
Homaloptera microstoma Boulenger, 1899a: 228 (type locality:<br />
Malaysia: Borneo: Sarawak: Akar River; holotype:<br />
BMNH 1895.7.2.49, Eschmeyer & Fricke, 2010; compound<br />
noun, indeclinable)<br />
Parhomaloptera obscura Vaillant, 1902: 130, figs. 36–38<br />
(type locality: Indonesia: Borneo: Kalimantan Timur:<br />
Bloeoe River [Bluu, 0°42'N 114°24'E]; syntypes [3]:<br />
RMNH, MNHN 1903-196 [1], Bertin & Estève, 1948:<br />
103; adjective, -us, -a, -um)<br />
8.13 Plesiomyzon Zheng & Chen, 1980<br />
Plesiomyzon Zheng & Chen, 1980: 90 (type species: Plesiomyzon<br />
baotingensis Zheng & Chen, 1980: 90, by original<br />
designation). Gender masculine.<br />
8.13.1 Plesiomyzon baotingensis Zheng & Chen, 1980<br />
Plesiomyzon baotingensis Zheng & Chen, 1980: 90, fig. 1<br />
(type locality: China: Hainan: Baoting Xian; syntypes:<br />
IHB 76VI6077, 6079 [2], DBJU 76VI6078 [1]; adjective,<br />
-is, -is, -e)<br />
8.14 Protomyzon Hora, 1932<br />
Protomyzon Hora, 1932a: 306 (type species: Homaloptera<br />
whiteheadi Vaillant, 1893: 92, by original designation).<br />
Gender masculine.<br />
Progastromyzon Hora & Jayaram, 1952: 191 (type species:
8.13.1 Plesiomyzon baotingensis, ZRC uncat., 39.4 mm SL; China:<br />
Hainan Island. (Photograph by Tan Heok Hui).<br />
8.13.1 Plesiomyzon baotingensis, ZRC uncat., 39.4 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
Progastromyzon griswoldi Hora & Jayaram, 1952: 192,<br />
by original designation). Gender masculine.<br />
8.14.1 Protomyzon aphelocheilus Inger & Chin, 1962<br />
Protomyzon aphelocheilus Inger & Chin, 1962: 110, fig. 53<br />
(type locality: Malaysia: Borneo: Sabah: Tambunan District:<br />
Sungei Kaingeran; holotype: FMNH 68166; noun<br />
in apposition, indeclinable)<br />
8.14.2 Protomyzon borneensis Hora & Jayaram, 1952<br />
Protomyzon borneensis Hora & Jayaram, 1952: 193, fig. 2<br />
(type locality: Malaysia: Borneo: Sabah: Mount Kinabalu:<br />
Bohanan River between Jinompan and Ranan, Kampong<br />
Kundasang; holotype: MCZ 37207 [not 34801];<br />
adjective, -is, -is, -e)<br />
8.14.3 Protomyzon griswoldi (Hora & Jayaram, 1952)<br />
Progastromyzon griswoldi Hora & Jayaram, 1952: 192, fig. 1<br />
(type locality: Malaysia: Borneo: Sabah: Mount Kinabalu:<br />
Kaddamayaa River at Kiau; holotype: MCZ 37206<br />
[not 34806]; noun in genitive, indeclinable)<br />
8.14.4 Protomyzon whiteheadi (Vaillant, 1893)<br />
Homaloptera Whiteheadi Vaillant, 1893: 92, pl. 1 fig. 2 (type<br />
locality: Malaysia: Borneo: Sabah: Kinabalu; syntypes:<br />
MNHN 1889-86 [1], 1889-87 [1], 1889-88 [5]; noun in<br />
genitive, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
68<br />
8.14.2 Protomyzon borneensis, ZRC uncat., 32.4 mm SL; Malaysia:<br />
Borneo: Sabah: Ranau: Labuk. (Photograph by Tan Heok Hui).<br />
8.14.2 Protomyzon borneensis, ZRC uncat., 40.5 mm SL. (Photograph<br />
by Tan Heok Hui).<br />
8.15 Pseudogastromyzon Nichols, 1925<br />
Pseudogastromyzon Nichols, 1925e: 1 (subgenus <strong>of</strong><br />
Hemimyzon Regan, 1911: 32; type species: Hemimyzon<br />
zebroidus Nichols, 1925e: 1, by monotypy). Gender masculine.<br />
Labigastromyzon Tang & Chen, 1996: 234 (subgenus <strong>of</strong><br />
Pseudogastromyzon Nichols, 1925e: 1; type species:<br />
Crossostoma fangi Nichols, 1931a: 63, by original designation;<br />
spellt Labigastromyzo p. 234, an obvious inadvertent<br />
error, thus incorrect original spelling [Code art.<br />
32.5.1]). Gender masculine.<br />
8.15.1 Pseudogastromyzon changtingensis Liang, 1942<br />
Pseudogastromyzon fasciatus changtingensis Liang, 1942:<br />
1, fig. 1 (type locality: China: western Fukien [Fujian]:<br />
Chang ting-hsien; holotype: Institute <strong>of</strong> Zoology and<br />
Botany, Provincial Academy <strong>of</strong> Fukien 9; adjective, -is,<br />
-is, -e)<br />
? Pseudogastromyzon tungpeiensis Chen & Liang, 1949:<br />
158, fig. 1 (type locality: China: Guangdong: Tung-peishiu,<br />
Lienhsien; holotype: Kwantung Provincial College<br />
<strong>of</strong> Letters and Science 651; adjective, -is, -is, -e)<br />
8.15.2 Pseudogastromyzon cheni Liang, 1942<br />
Pseudogastromyzon cheni Liang, 1942: 4, fig. 2 (type local-
ity: China: western Fukien [Fujian]: Changting-hsien;<br />
holotype: Institute <strong>of</strong> Zoology and Botany, Provincial<br />
Academy <strong>of</strong> Fukien 6; noun in genitive, indeclinable)<br />
8.15.3 Pseudogastromyzon fangi (Nichols, 1931)<br />
Crossostoma fangi Nichols, 1931a: 263, fig. (type locality:<br />
China: Guangdong: near Canton; holotype: AMNH 10111<br />
[Lingnan University Canton P-81], Eschmeyer & Fricke,<br />
2010; noun in genitive, indeclinable)<br />
8.15.4 Pseudogastromyzon fasciatus (Sauvage, 1878)<br />
Psilorhynchus fasciatus Sauvage, 1878a: 88 (type locality:<br />
China: western Fukien [Fujian] at 1200 masl [Fujian:<br />
Kuatun, 27°51'N 117°48'E; see Crossostoma davidi];<br />
syntypes: MNHN A.3449 [11], 9821 [3], Bertin & Estève,<br />
1948: 102, 103; adjective, -us, -a, -um)<br />
Hemimyzon zebroidus Nichols, 1925e: 1, fig. 1 (type locality:<br />
China: Fukien [Fujian]: Yenping; holotype: AMNH<br />
8392; adjective, -us, -a, -um)<br />
Pseudogastromyzon fasciatus jiulongjiangensis Chen,<br />
1980a: 110, fig. 5 (type locality: China: Fujian: Jiulong<br />
River in Longyan Xian and Nanjing Xian; syntypes: IHB<br />
74V445, 447, 448, 761, 763, 765, 766, 768–770, 776,<br />
779, 782–784 [15]; adjective, -is, -is, -e)<br />
8.15.5 Pseudogastromyzon laticeps Chen & Zheng, in<br />
Zheng & Chen, 1980<br />
Pseudogastromyzon laticeps Chen & Zheng, in Zheng &<br />
Chen, 1980: 96, fig. 7 (type locality: China: Guangdong:<br />
Haifeng Xian: a mountain stream at Lianhua Shan; syntypes:<br />
IHB 76IV8166–8170, 8196–8200 [10], DBJU<br />
76IV8171–8175, 8202–8206 [10]; compound adjective,<br />
indeclinable)<br />
8.15.6 Pseudogastromyzon lianjiangensis Zheng, 1981<br />
Pseudogastromyzon lianjiangensis Zheng, 1981: 59, fig. 2<br />
(type locality: China: Guangdong: Puning County: mountain<br />
stream on Danan Shan [mountain], in Lianjiang drainage;<br />
syntypes: IHB 80VIII0008, 24 [2], DBJU<br />
80VIII0005, 22, 27 [3]; adjective, -is, -is, -e)<br />
8.15.7 Pseudogastromyzon meihuashanensis Li, 1998<br />
Pseudogastromyzon meihuashanensis Li, 1998: 260, fig. 1<br />
(type locality: China: Fujian: Meihuashan Nature Reserve,<br />
Dutou Station, Manzhuxi [Manzhu stream], Jiulongjiang<br />
drainage, 25°25'N 116°55'E; holotype: FM<br />
00248; also spelt meihuashangensis p. 260, an obvious<br />
inadvertent error, thus incorrect original spelling [Code<br />
art. 32.5.1]; adjective, -is, -is, -e)<br />
8.15.8 Pseudogastromyzon myersi Herre, 1932<br />
Pseudogastromyzon myersi Herre, 1932: 430 (type locality:<br />
China: Hong Kong island; holotype: CAS-SU 25731,<br />
Böhlke, 1953: 40; noun in genitive, indeclinable)<br />
Pseudogastromyzon maculatum Chen & Zheng, in Zheng &<br />
Chen, 1980: 95, fig. 6 (type locality: China: Guangdong:<br />
East River in Lianping Xian and Xinfeng Xian; syntypes:<br />
IHB 76V7355, 7357, 7359, 7361, 7363, 7365, 7367,<br />
7371, 7587, 7593 [10], DBJU 76V7595, 7599, 7785,<br />
7788, 7789 [5]; adjective, -us, -a, -um)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
69<br />
8.15.2 Pseudogastromyzon cf. cheni, ZRC uncat., 39.2 mm SL;<br />
China: aquarium specimen. (Photograph by Tan Heok Hui).<br />
8.15.2 Pseudogastromyzon cf. cheni, ZRC uncat., 39.2 mm SL.<br />
(Photograph by Tan Heok Hui).<br />
8.15.9 Pseudogastromyzon peristictus Zheng & Li, 1986<br />
Pseudogastromyzon peristictus Zheng & Li, 1986: 77, fig. 1<br />
(type locality: China: Guangdong: Fengshun Xian; syntypes:<br />
DBJU 85VIII109–118, 120–121 [12], IHB<br />
85VIII107–108, 119, 122 [4]; etymology unknown, treated<br />
as noun in apposition, indeclinable)<br />
8.16 Sewellia Hora, 1932<br />
Sewellia Hora, 1932: 315 (type species: Balitora lineolata<br />
Valenciennes, in Cuvier & Valenciennes, 1846: 99, by<br />
original designation). Gender feminine.<br />
Diardichthys Roberts, 1998: 275 (subgenus <strong>of</strong> Sewellia Hora,<br />
1932a: 315; type species: Sewellia diardi Roberts, 1998:<br />
281, by original designation). Gender masculine.<br />
Parasewellia Nguyen (H. D.) & Nguyen (V. H.), in Nguyen<br />
(V. H.), 2005: 543 (type species: Parasewellia tetralobata<br />
Nguyen & Nguyen, in Nguyen, 2005: 545, by original<br />
designation). Gender feminine.<br />
Taxonomic notes. The validitidy <strong>of</strong> the many names created<br />
in recent years in the Vietnamese literature cannot be evaluated.<br />
The descriptions are <strong>of</strong> little use and the quality <strong>of</strong> the<br />
illustrations <strong>of</strong> most species does not allow to guess their<br />
identity. As most species have very restricted ranges, it is<br />
expected that those from widely distant localities might in<br />
fact be valid, while the many species described from exactly<br />
the same locality may be found to represent single species.
8.16.12 Sewellia speciosa, CMK 15551, 45.8 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
8.16.5 Sewellia elongata, CMK 15529, 63.5 mm SL.<br />
8.16.1 Sewellia albisuera Freyh<strong>of</strong>, 2003<br />
Sewellia albisuera Freyh<strong>of</strong>, 2003: 226, figs. 1–2 (type locality:<br />
Vietnam: Quang Nam Da Nang Province: Thu Bon<br />
River about 16 km west <strong>of</strong> Than My; 15°40.33'N<br />
107°48.45'E; holotype: ZFMK 32913; noun in apposition,<br />
indeclinable)<br />
Parasewellia tetralobata Nguyen [H. D.] & Nguyen [V. H.],<br />
in Nguyen [V. H.], 2005: 545, fig. 1 (type locality: Vietnam:<br />
Quang Nam Province: Thu Bon River [a coastal<br />
drainage south <strong>of</strong> Danang]; holotype: HNUE; adjective,<br />
-us, -a, -um)<br />
8.16.2 Sewellia analis Nguyen & Nguyen, in Nguyen, 2005<br />
Sewellia analis Nguyen [H. D.] & Nguyen [V. H.], in Nguyen<br />
[V. H.], 2005: 587, fig. 25 (type locality: Vietnam:<br />
Thua Thien Hue Province: A Luoi district, A Sap, Hong<br />
Van, Tarinh; holotype: HNUE; also spelt analus p. 278,<br />
587 fig. 25, 689; as first reviser I select analis as the<br />
correct original spelling; noun in apposition, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
70<br />
8.16.3 Sewellia breviventralis Freyh<strong>of</strong> & Serov, 2000<br />
Sewellia breviventralis Freyh<strong>of</strong> & Serov, 2000a: 231, figs. 9–<br />
10 (type locality: Vietnam: Kontum Province: Pako River<br />
about 50 km north <strong>of</strong> Kontum, a tributary <strong>of</strong> Sesan,<br />
Mekong drainage; 14°39'60"N 107°46'98"E; holotype:<br />
ZFMK 20955; compound noun, indeclinable)<br />
8.16.4 Sewellia diardi Roberts, 1998<br />
Sewellia diardi Roberts, 1998: 281, fig. 6 (type locality: Laos:<br />
Attapu Province.: lower Xe Nam Noi 1.5 km downstream<br />
from bridge on route 232 to Attapu, 26 km from Ban<br />
Nam Tang, 270 masl; holotype: ZRC 40364; noun in<br />
genitive, indeclinable)<br />
8.16.5 Sewellia elongata Roberts, 1998<br />
Sewellia elongata Roberts, 1998: 283, fig. 7 (type locality:<br />
Laos: Champasak Province: Bolaven plateau: Xe Nam<br />
Noi at proposed dam site for Xe Nam Noi-Xe Pian hydropower<br />
dam, 16 km southeast <strong>of</strong> Ban Nam Tang, 730<br />
masl; holotype: ZRC 40361; adjective, -us, -a, -um)<br />
8.16.6 Sewellia lineolata (Valenciennes, in Cuvier & Valenciennes,<br />
1846)<br />
Balitora lineolata Valenciennes, in Cuvier & Valenciennes,<br />
1846: 99 (type locality: Vietnam: Cochinchine; syntypes:<br />
MNHN 2906 [3], BMNH 1931.10.26.3 [1], RMNH 2011<br />
[2], ZSI F 11291/1 [1], Bertin & Estève, 1948: 104, Kottelat,<br />
1994b: 109, Silas, 1953: 230; adjective, -us, -a, -um)<br />
Sewellia songboensis Nguyen [H. D.] & Nguyen [V. H.], in<br />
Nguyen [V. H.], 2005: 581, fig. 22 (type locality: Vietnam:<br />
Thua Thien Hue Province: A Luoi district: A Sap;<br />
holotype: HNUE; adjective, -is, -is, -e)<br />
8.16.7 Sewellia marmorata Serov, 1996<br />
Sewellia marmorata Serov, 1996: 197, fig. 1 (type locality:<br />
Vietnam: Zalai-Kontum Province: "a mountain brook at<br />
the area <strong>of</strong> the forestry Kon-Khanym", 1250 masl,<br />
13°57'N 107°32'E [Gia Lai Province: stream at Kon Ha<br />
Nuong, about 70 km north <strong>of</strong> An Khe, 13°57'N 108°32'E;<br />
Freyh<strong>of</strong> & Serov, 2000a: 223]; holotype: ZMMU P<br />
19760; adjective, -us, -a, -um)<br />
8.16.8 Sewellia media Nguyen & Nguyen, in Nguyen, 2005<br />
Sewellia medius Nguyen [H. D.] & Nguyen [V. H.], in Nguyen<br />
[V. H.], 2005: 585, fig. 24 (type locality: Vietnam:<br />
Thua Thien Hue Province: A Luoi district, A Sap, Hong<br />
Van, Tarinh; holotype: HNUE; adjective, -us, -a, -um)<br />
Sewellia brevis Nguyen [H. D.] & Nguyen [V. H.], in Nguyen<br />
[V. H.], 2005: 278, 585, fig. 24 (an alternative name<br />
used simultaneously for Sewellia medius Nguyen &<br />
Nguyen, in Nguyen, 2005: 585; as first reviser I give precedence<br />
to S. medius; adjective, -is, -is, -e)<br />
8.16.9 Sewellia monolobata (Nguyen & Nguyen, in Nguyen,<br />
2005)<br />
Parasewellia monolobata Nguyen [H. D.] & Nguyen [V. H.],<br />
in Nguyen [V. H.], 2005: 546, fig. 2 (type locality: Vietnam:<br />
Quang Nam Province: Thu Bon River [a coastal<br />
drainage south <strong>of</strong> Danang]; holotype: HNUE; adjective,<br />
-us, -a, -um)
Parasewellia polylobata Nguyen [H. D.] & Nguyen [V. H.],<br />
in Nguyen [V. H.], 2005: 548, fig. 3 (type locality: Vietnam:<br />
Quang Nam Province: Thu Bon River [a coastal<br />
drainage south <strong>of</strong> Danang]; holotype: HNUE; also spelt<br />
polyloba in caption <strong>of</strong> fig. 3, as first reviser I select polylobata<br />
as the correct original spelling; subjective simultaneous<br />
synonym <strong>of</strong> Parasewellia monolobata Nguyen<br />
& Nguyen, in Nguyen, 2005: 546; as first reviser, I give<br />
precedence to P. monolobata; adjective, -us, -a, -um)<br />
8.16.10 Sewellia patella Freyh<strong>of</strong> & Serov, 2000<br />
Sewellia patella Freyh<strong>of</strong> & Serov, 2000a: 235, figs. 13–14<br />
(type locality: Vietnam: Gia Lai Province: stream Azun<br />
about 30 km east <strong>of</strong> Pleiku; 14°02'34"N 108°21'07" E;<br />
holotype: ZFMK 20962; noun in apposition, indeclinable)<br />
8.16.11 Sewellia pterolineata Roberts, 1998<br />
Sewellia pterolineata Roberts, 1998: 277, fig. 4 (type locality:<br />
Vietnam: Nghia Bin Province: Trac Khuc River; holotype:<br />
RMNH 31832; adjective, -us, -a, -um)<br />
8.16.12 Sewellia speciosa Roberts, 1998<br />
Sewellia speciosa Roberts, 1998: 279, fig. 5 (type locality:<br />
Laos: Attapu Province.: lower Xe Nam Noi 1.5 km downstream<br />
from bridge on route 232 to Attapu, 26 km from<br />
Ban Nam Tang, 270 masl; holotype: ZRC 40368; adjective,<br />
-us, -a, -um)<br />
8.16.13 Sewellia trakhucensis Nguyen & Nguyen, in<br />
Nguyen, 2005<br />
Sewellia trakhucensis Nguyen [H. D.] & Nguyen [V. H.], in<br />
Nguyen [V. H.], 2005: 583, fig. 23 (type locality: Vietnam:<br />
Quang Ngai Province, Son Ha, Tra Khuc River;<br />
holotype: HNUE; adjective, -is, -is, -e)<br />
8.17 Vanmanenia Hora, 1932<br />
Homalosoma Boulenger, 1901: 270 (name not available,<br />
incorrect subsequent spelling for Homaloptera van Hasselt,<br />
1823: 133 [Boulenger, in litt. to Hora, 1932: 309]; [if<br />
treated as available then: type species: Homalosoma stenosoma<br />
Boulenger, 1901: 270, by monotypy; junior homonym<br />
<strong>of</strong> Homalosoma Wagler, 1830: 190 in Reptilia, and<br />
Homalosoma Agassiz, 1848: 531 in Coleoptera; not junior<br />
homonym <strong>of</strong> Homalosoma Oersted, in Keferstein,<br />
1865: 436, in Vermes, which is not available because listed<br />
in synonymy (Code art. 11.6)]). Gender neuter.<br />
Vanmanenia Hora, 1932a: 309 (type species: Homalosoma<br />
stenosoma Boulenger, 1901: 270, by original designation).<br />
Gender feminine.<br />
Praeformosania Fang, 1935a: 71 (type species: Praeformosania<br />
pingchowensis Fang, 1935a: 72, by original designation).<br />
Gender feminine.<br />
Taxonomic notes. The validitidy <strong>of</strong> the many names created<br />
in recent years in the Vietnamese literature cannot be evaluated.<br />
The descriptions are <strong>of</strong> little use and the quality <strong>of</strong> the<br />
illustrations <strong>of</strong> most species does not allow to guess their<br />
identity. As most species have very restricted ranges, it is<br />
expected that those from widely distant localities might in<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
71<br />
fact be valid, while the many species described from exactly<br />
the same locality may be found to represent single species.<br />
For some species there is not even informative locality data.<br />
The colour pattern <strong>of</strong> Vanmanenia species strikingly changes<br />
with growth.<br />
Nomenclatural notes. Boulenger (1901: 270) used the name<br />
Homalosoma, without mentioning it as a new genus. Since,<br />
authors have speculated as to whether it was intended as a<br />
new genus or was a lapsus for Homaloptera and/or a confusion<br />
with the snake genus Homalosoma (Boulenger also<br />
published on reptiles). The answer to these speculations has<br />
already been given by Boulenger himself in a letter to Hora<br />
(1932: 309): "Homalosoma stenosoma is obviously a lapsus<br />
calami for Homaloptera stenosoma and only a lapsus<br />
memoriae could account for my using Homalosoma to denote<br />
a new genus, that name being in use for a well known<br />
genus <strong>of</strong> snakes". Therefore, Homalosoma is an incorrect<br />
subsequent spelling <strong>of</strong> Homaloptera. Should one consider it<br />
as available, it would be permanently invalid as a junior homonym<br />
for Homalosoma Wagler, 1830 (see above).<br />
Species inquirenda et incertae sedis<br />
8.17.1 ? Vanmanenia multiloba (Mai, 1978)<br />
Homaloptera multiloba Mai, 1978: 208, fig. 95 (type locality:<br />
northern Vietnam; syntypes: DVZUT; compound<br />
noun, indeclinable)<br />
Species inquirenda et incertae sedis<br />
8.17.2 ? Vanmanenia nahangensis Nguyen, 2005<br />
Vanmanenia nahangensis Nguyen, 2005: 572, fig. 17 (type<br />
locality: Vietnam: Tuyen Quang Province: Na Hang district:<br />
Gam River; holotype: NCNTTSI; also spelt nahangenis<br />
p. 573, an obvious inadvertent error, thus incorrect<br />
original spelling [Code art. 32.5.1]; adjective, -is,<br />
-is, -e)<br />
Species inquirenda et incertae sedis<br />
8.17.3 Vanmanenia tetraloba (Mai, 1978)<br />
Homaloptera tetraloba Mai, 1978: 210, fig. 97 (type locality:<br />
northwestern Vietnam; syntypes: DVZUT; compound<br />
noun, indeclinable)<br />
Species inquirenda et incertae sedis<br />
8.17.4 Vanmanenia ventrosquamata (Mai, 1978)<br />
Homaloptera ventrosquamata Mai, 1978: 209, fig. 96 (type<br />
locality: Vietnam: Cau River [Bac Thai, Bac Giang and<br />
Bac Ninh provinces]; syntypes: DVZUT; adjective, -us,<br />
-a, -um)<br />
8.17.5 Vanmanenia caldwelli (Nichols, 1925)<br />
Homaloptera caldwelli Nichols, 1925c: 1 (type locality:<br />
China: Fukien [Fujian]: near Yenping; holotype: AMNH<br />
8413; noun in genitive, indeclinable)<br />
8.17.6 Vanmanenia caobangensis Nguyen, 2005<br />
Vanmanenia caobangensis Nguyen, 2005: 571, fig. 16 (type<br />
locality: Vietnam: Cao Bang Province: Bang Giang River<br />
at Cao Bang, Red River drainage; holotype: NCNTT-<br />
SI; adjective, -is, -is, -e)
8.17.7 Vanmanenia crassicauda, top, CMK 15328, 23.8 mm SL;<br />
bottom, ZRC 45390, 66.2 mm SL, holotype; Laos: Houaphan Province:<br />
Nam Xam drainage.<br />
8.17.7 Vanmanenia crassicauda, CMK 15410, 56.9 mm SL.<br />
8.17.7 Vanmanenia crassicauda Kottelat, 2000<br />
Vanmanenia crassicauda Kottelat, 2000: 75, fig. 66 (type<br />
locality: Laos: Houaphan Province: Houay Keap, waterfall<br />
adjacent to road on creek entering Nam Xam at km 5<br />
on road from Xam Tai to Ban Houatangoua; 20°01'00"N<br />
104°35'44"E; holotype: ZRC 45390; compound noun,<br />
indeclinable)<br />
? Vanmanenia microlepis Nguyen, 2005: 578, fig. 20 (type<br />
locality: Vietnam: Lai Chau Province: Phong Tho district:<br />
Nam So stream [Song Da drainage]; holotype:<br />
NCNTTSI; compound noun, indeclinable)<br />
8.17.8 Vanmanenia gymnetrus Chen, 1980<br />
Vanmanenia gymnetrus Chen, 1980a: 100, fig. 1 (type locality:<br />
China: Fujian: Longyan County: Jiulong River; syntypes:<br />
IHB 74VI0786–0791 [6]; noun in apposition, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
72<br />
8.17.9 Vanmanenia hainanensis Chen & Zheng, in Zheng<br />
& Chen, 1980<br />
Vanmanenia hainanensis Chen & Zheng, in Zheng & Chen,<br />
1980: 93, fig. 4 (type locality: China: Hainan: mountain<br />
stream <strong>of</strong> Chang Jiang in Qiongzhong Xian; syntypes:<br />
IHB 76V9142, 9168, 9173–9179 [9], DBJU 76V9181,<br />
9183, 9185, 9188, 9190, 9191 [6]; adjective, -is, -is, -e)<br />
8.17.10 Vanmanenia homalocephala Zhang & Zhao, 2000<br />
Vanmanenia homalocephala Zhang & Zhao, 2000: 458, fig. 1<br />
(type locality: China: Guangxi: Yongfu County: creek in<br />
Baishou, in Luoqing Jiang system, tributary <strong>of</strong> Liu Jiang,<br />
Pear River drainage; 26°35'N 118°15'E; holotype: ASIZB<br />
70180; compound adjective, -us, -a, -um)<br />
8.17.11 Vanmanenia lineata Fang, 1935<br />
Vanmanenia lineata Fang, 1935a: 78, fig. 11 (type locality:<br />
China: north-western Guangxi: Kwang-lau, Ling-yuenhsien;<br />
holotype: NRIBAS M.737; adjective, -us, -a, -um)<br />
Formosania yaoshanensis Wu, 1939: 128, pl. 3 fig. 7 (type<br />
locality: China: Guangxi: Li-kiang at Yao-shan; syntypes:<br />
[LU] 264–268 [5]; adjective, -is, -is, -e)<br />
8.17.12 Vanmanenia pingchowensis (Fang, 1935)<br />
Gobius Pa-che-tsee-yu Dabry de Thiersant, 1872: 179, pl. 37<br />
fig. 10 (name not binominal, not available; locality: China:<br />
Sichuan)<br />
Praeformosania pingchowensis Fang, 1935a: 72, fig. 9 (type<br />
locality: China: South Kweichow [Guizhou]: Ping-chowhsien;<br />
holotype: NRIBAS 4024; adjective, -is, -is, -e)<br />
Praeformosania intermedia Fang, 1935a: 75, fig. 10 (type<br />
locality: China: South Kweichow [Guizhou]: Hu-yuenhsien<br />
[or Tu-yuen-hsien, p. 76], in mountain streams running<br />
into Tung-ting Lake; holotype: NRIBAS M.2709;<br />
simultaneous subjective synonym <strong>of</strong> Praeformosania<br />
pingchowensis Fang, 1935a: 72; first reviser [Chen,<br />
1980a: 99] gave precedence to P. pingchowensis; adjective,<br />
-us, -a, -um)<br />
Vanmanenia polylepis Pan, Liu & Zheng, 1983: 107, fig. 2<br />
(type locality: China: Guangdong: Beijiang River; holotype:<br />
SCNU 3194 [mentioned in Chinese text only]; compound<br />
noun, indeclinable)<br />
Taxonomic notes. Among the fishes presently reported from<br />
Sichuan (Ding, 1994), Gobius pa-che-tsee-yu, as illustrated<br />
by Dabry de Thiersant, shows the greatest similarity with<br />
V. pingchowensis.<br />
8.17.13 Vanmanenia serrilineata Kottelat, 2000<br />
Vanmanenia serrilineata Kottelat, 2000: 76, fig. 68 (type<br />
locality: Laos: Louangnamtha Province: Nam Tha at Ban<br />
Finho, about 14 km north <strong>of</strong> Luang Nam Tha; 21°04'44"N<br />
101°24'09"E; holotype: ZRC 45392; adjective, -us, -a,<br />
-um)<br />
? Vanmanenia mon<strong>of</strong>asciodorsalata Nguyen, 2005: 574,<br />
fig. 18 (type locality: Vietnam: Dien Bien Province: Nam<br />
Rom River; holotype: NCNTTSI; also spelt mon<strong>of</strong>asciodorsala<br />
p. 574, fig. 18, p. 577, mon<strong>of</strong>asciodossalata<br />
p. 574, mon<strong>of</strong>asciodorsalata p. 689; as first reviser I select<br />
mon<strong>of</strong>asciodorsalata as the correct original spelling;<br />
adjective, -us, -a, -um)
? Vanmanenia trifasciodorsalata Nguyen, 2005: 576, fig. 19<br />
(type locality: Vietnam: Dien Bien Province: Nam Rom<br />
River; holotype: NCNTTSI; spelt trifasciodorsala pp.<br />
268, 575, tri<strong>of</strong>asciodossalata p. 576, trifaseudorsala p.<br />
576, fig. 19, trifasciodorsalata p. 689; as first reviser I<br />
select trifasciodorsalata as the correct original spelling;<br />
adjective, -us, -a, -um)<br />
8.17.14 Vanmanenia stenosoma (Boulenger, 1901)<br />
Homalosoma stenosoma Boulenger, 1901: 270, pl. 23 fig. 3<br />
(type locality: China: Zhejiang: Ningpo [Ningbo]; holotype:<br />
BMNH 1901.3.6.16; compound noun, indeclinable)<br />
Homaloptera caldwelli chekianensis Tchang, 1932c: 83, fig.<br />
(type locality: China: Chekiang [Zhejiang]; holotype:<br />
ZMFIB 7013; spelt chekinensis p. 83, chekianensis<br />
9 Serpenticobitidae Kottelat, 2012<br />
Serpenticobitinae Nalbant, 2002: pl. 7 (not available, no<br />
explicitly indicated as intentionally new, Code art. 16.1,<br />
no type genus cited, art. 16.2)<br />
Serpenticobitidae Kottelat, 2012: 140 [appendix <strong>of</strong> present<br />
work] (type genus: Serpenticobitis Roberts, 1997: 109)<br />
9.1 Serpenticobitis Roberts, 1997<br />
Serpenticobitis Roberts, 1997: 109 (type species: Serpenticobitis<br />
octozona Roberts, 1997: 112, by original designation).<br />
Gender feminine.<br />
Bangfaia Kottelat, 1998a: 96, fig. 134 (name published by<br />
inadvertence [existed in manuscript, corrected in pro<strong>of</strong>s<br />
after receiving description <strong>of</strong> Serpenticobitis, but some<br />
mentions were overlooked], no type species designated,<br />
nomen nudum)<br />
9.1.1 Serpenticobitis cingulata Roberts, 1997<br />
Serpenticobitis cingulata Roberts, 1997: 113, fig. 6 (type<br />
locality: Thailand: Loei Province [actually Chiang Rai<br />
Province]: Huay Ngao where it enters Mekong mainstream,<br />
1 km south <strong>of</strong> Ban Chaem Pong, about 30 km<br />
south <strong>of</strong> Chiang Khong; holotype: CAS 95170; adjective,<br />
-us, -a,<br />
-um)<br />
9.1.2 Serpenticobitis octozona Roberts, 1997<br />
Serpenticobitis octozona Roberts, 1997: 112, fig. 5 (type<br />
locality: Laos: Attapu Province: Xe Nam Noi, eastern<br />
slope <strong>of</strong> Bolaven Plateau, about 3 km downstream from<br />
ferry crossing road Saravan to Attapu and 30 km from<br />
Ban Nam Tang; holotype: CAS 95165; noun in apposition,<br />
indeclinable)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
Family SERPENTICOBITIDAE<br />
73<br />
pp. 84, 85, first reviser not researched, but chekinensis<br />
apparently an error <strong>of</strong> inadvertence, thus incorrect original<br />
spelling [Code art. 32.5.1]; adjective, -is, -is, -e)<br />
8.17.15 Vanmanenia striata Chen, 1980<br />
Vanmanenia striata Chen, 1980a: 101, fig. 2 (type locality:<br />
China: Yunnan: Yuanjiang [Red River], Xiaguan Shi;<br />
syntypes: IHB 585280–285, 446, 450, 480, 482, 646483–<br />
485 [13]; adjective, -us, -a, -um)<br />
8.17.16 Vanmanenia xinyiensis Zheng & Chen, 1980<br />
Vanmanenia xinyiensis Zheng & Chen, 1980: 94, fig. 5 (type<br />
locality: China: Guangdong: a moutain stream <strong>of</strong> West<br />
River in Xinyi Xian; syntypes: IHB 77VII070, 75, 77–<br />
79 [5], DBJU 77VII081, 90, 475 [3]; adjective, -is, -is, -e)<br />
9.1.3 Serpenticobitis zonata Kottelat, 1998<br />
Serpenticobitis zonata Kottelat, 1998a: 92, fig. 135 (type<br />
locality: Laos: Khammouan Province: Xe Bangfai basin:<br />
about 3 km upriver <strong>of</strong> Ban Pakphanang; 17°24'20"N<br />
104°45'50"E; holotype: ZRC 41810; adjective, -us, -a,<br />
-um)<br />
9.1.2 Serpenticobitis octozona, CMK 15675, 47.2 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
9.1.3 Serpenticobitis zonata, ZRC uncat., 44.5 mm SL. (Photograph<br />
by Tan Heok Hui).
10 Nemacheilidae Regan, 1911<br />
Nemacheilinae Regan, 1911: 31 (type genus: Nemacheilus<br />
Bleeker, 1863a: 37)<br />
Adiposiidae Jordan, 1923: 145 (type genus: Adiposia Annandale<br />
& Hora, 1920: 182)<br />
Lefuini Prok<strong>of</strong>iev, 2010: 890 (type genus: Lefua Herzenstein,<br />
1888: 3)<br />
Yunnanilini Prok<strong>of</strong>iev, 2010: 890 (type genus: Yunnanilus<br />
Nichols, 1925b: 1)<br />
Triplophysini Prok<strong>of</strong>iev, 2010: 892 (type genus: Triplophysa<br />
Rendahl, 1933: 21 [indicated as "Triple-space<br />
Rendahl, 1933", treated as an incorrect subsequent spelling<br />
<strong>of</strong> Triplophysa])<br />
Nomina nuda<br />
Gojeb Prok<strong>of</strong>iev, 2004b: 193<br />
Noemacheilus sikkimensis Mani, 1974: 671<br />
Noemacheilus lunanensis Li et al., 1999: 4, 5, 6 (nomen<br />
nudum; author indicated as Li & Xia, but does not seem<br />
to be published)<br />
Species inquirendae et incertae sedis<br />
Nemacheilus blythii Day, 1870a: 552 (type locality: unknown;<br />
syntypes: ZSI A.960 [2, lost], Whitehead & Talwar,<br />
1976: 156; noun in genitive, indeclinable)<br />
Cobitis montanus Jerdon, 1849: 332 (type locality: India:<br />
Karnataka: a small stream in Coorg [Kodagu, 12°25'N<br />
75°45'E]; types: LU; if later treated as valid in Schistura,<br />
would become a junior secondary homonym <strong>of</strong> Schistura<br />
montana McClelland, 1838: 947; adjective, -us, -a,<br />
-um)<br />
Schistura ocellata M'Clelland, 1839: 441 (type locality: India:<br />
Upper Assam; types: LU; adjective, -us, -a, -um)<br />
10.1 Aborichthys Chaudhuri, 1913<br />
Aborichthys Chaudhuri, 1913: 244 (type species: Aborichthys<br />
kempi Chaudhuri, 1913: 245, by monotypy). Gender<br />
masculine.<br />
Nomen nudum<br />
Aborichthys bijulensis Tekriwal & Rao, 1999: 89 (nomen<br />
nudum; locality: India: Meghalaya: Garo Hills [near Tura;<br />
p. 36])<br />
10.1.1 Aborichthys elongatus Hora, 1921<br />
Aborichthys elongatus Hora, 1921c: 735, fig. (type locality:<br />
India: Reang River, Darjeeling District; holotype: ZSI<br />
F 10087/1 [largest specimen, 74 mm SL]; secondary junior<br />
homonym <strong>of</strong> Nemachilus kungessanus elongatus<br />
Herzenstein, 1888: 44, when placed in Nemacheilus by<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Family NEMACHEILIDAE<br />
74<br />
Menon, 1999: 184; never replaced and not presently treated<br />
as congeneric, therefore replacement not needed [Code<br />
art. 59.2]; adjective, -us, -a, -um)<br />
10.1.2 Aborichthys garoensis Hora, 1925<br />
Aborichthys garoensis Hora, 1925: 233, figs. 2–4 (type locality:<br />
India: Assam: Garo Hills, Tura; holotype: ZSI<br />
F 10669/1; adjective, -is, -is, -e)<br />
Nomenclatural notes. Hora (1925) based his description<br />
on three specimens, one <strong>of</strong> which was designated as type<br />
specimen, thus it is the holotype. Its catalogue number was<br />
given as ZSI F 10669/1 and it was illustrated at natural size<br />
as fig. 1; the illustrated specimen is about 93 mm SL while<br />
the three types are 89.5, 89.3 and 85.8 mm SL. Under the<br />
same catalogue number, Menon & Yazdani (1968: 119) list<br />
two specimens as syntypes. The fate <strong>of</strong> the third specimen is<br />
apparently unknown. The holotype might be recognisable<br />
by comparaison with Hora's figure.<br />
10.1.3 Aborichthys kempi Chaudhuri, 1913<br />
Aborichthys kempi Chaudhuri, 1913: 245, pl. 7 figs. 1–1b<br />
(type locality: N.E. India: Abor Hills: Egar Stream between<br />
Renging and Rotung / Dihang River near Yembung<br />
/ Sirpo River near Renging; syntypes: ZSI F 7721/<br />
1–7723/1 [3], 7725/1–7727/1 [3], 7769/1–7770/1 [2],<br />
7778/1–7779/1 & 7878/1–7879/1 & 8297/1–8298/1 [8],<br />
Menon & Yazdani, 1968: 119; noun in genitive, indeclinable)<br />
10.1.4 Aborichthys tikaderi Barman, 1985<br />
Aborichthys tikaderi Barman, 1985: 680, fig. 1 (type locality:<br />
India: Arunachal Pradesh: Namdapha Wildlife Sanctuary;<br />
holotype: ZSI FF 2135; noun in genitive, indeclinable)<br />
10.2 Acanthocobitis Peters, 1861<br />
Acanthocobitis Peters, 1861: 712 (type species: Acanthocobitis<br />
longipinnis Peters, 1861: 712, by monotypy). Gender<br />
feminine.<br />
? Paracanthocobitis Grant, 2007c: 3 [also 2007d: unnumb.<br />
p. 51] (subgenus <strong>of</strong> Acanthocobitis Peters, 1861: 712;<br />
type species: Cobitis zonalternans Blyth, 1860: 172, by<br />
original designation). Gender feminine.<br />
Taxonomic notes. Grant (2007c) distinguished Paracanthocobitis<br />
from Acanthocobitis. This was not based on examination<br />
<strong>of</strong> specimens but on literature, one live individual and<br />
photographs. The characters said to distinguish the two subgenera<br />
are variable within and between them (position, shape<br />
and size <strong>of</strong> the suborbital flap). Body shape (elongated body)<br />
is not conclusive since the ranges <strong>of</strong> body depth overlap.<br />
The number <strong>of</strong> branched dorsal-fin rays, related to elongated<br />
body, is also marginally overlapping. The position <strong>of</strong> the
anus is a real difference but partly linked to the elongated<br />
body. The shape <strong>of</strong> the caudal fin is the only clear-cut character.<br />
On the other hand, the hypothesis that A. pavonaceus<br />
(A. longipinnis) might be a lineage distinct from the other<br />
species placed in Acanthocobitis is not unreasonable, but<br />
the hypothesis should be addressed by a proper study, based<br />
on examination <strong>of</strong> specimens <strong>of</strong> all concerned species. To<br />
distinguish lineages (subgenera) assumes a phylogenetic analysis,<br />
which is missing. There is no benefit to use a name<br />
created for no taxonomically justifiable reason.<br />
Further, the identity <strong>of</strong> A. longipinnis (type species <strong>of</strong><br />
Acanthocobitis) must be clarified.<br />
Incertae sedis<br />
10.2.1 Acanthocobitis longipinnis Peters, 1861<br />
Acanthocobitis longipinnis Peters, 1861: 712 (type locality:<br />
India: Ganges; holotype: ZMB 4795, Grant, 2007c: fig. 1;<br />
noun in apposition, indeclinable)<br />
Taxonomic notes. Grant (2007c) considered A. longipinnis<br />
Peters, 1861 to be a synonym <strong>of</strong> A. pavonaceus. This was<br />
apparently based on examination <strong>of</strong> the literature, one living<br />
specimen and photographs <strong>of</strong> syntypes <strong>of</strong> Cobitis pavonacea<br />
(one figured, SMF 68, about 100 mm SL) and <strong>of</strong> the<br />
poorly preserved holotype <strong>of</strong> A. longipinnis (ZMB 4795,<br />
156 mm SL). I have not examined the mentioned specimens,<br />
but the photographs <strong>of</strong> the types published by Grant do not<br />
allow me to accept the synonymy without a demonstration.<br />
They show specimens differing in the shape <strong>of</strong> the body (body<br />
depth 19 % SL in the holotype <strong>of</strong> A. longipinnis, vs. 12 in<br />
the figured syntype <strong>of</strong> C. pavonacea), the length <strong>of</strong> the dorsal<br />
fin base (27 % SL, vs. 32; 1.6 times in the distance between<br />
the origins <strong>of</strong> the anal and pelvic fins, vs. 1.3) and the<br />
shape <strong>of</strong> the caudal peduncle (depth 2.1 times in its length,<br />
vs. 1.13). Some <strong>of</strong> these values may depend from ability to<br />
determine the position <strong>of</strong> the base <strong>of</strong> the first and last rays <strong>of</strong><br />
the dorsal and anal fins from the photograph <strong>of</strong> A. longipinnis,<br />
but the proportions <strong>of</strong> the caudal peduncle do not suffer<br />
these reservations since they depend <strong>of</strong> bony structures (the<br />
vertebral column for the length and the hypural complex for<br />
the depth). The examination <strong>of</strong> a large series <strong>of</strong> specimens<br />
(including juveniles and adults) from a single locality is needed<br />
to understand morphological variability; the examination<br />
<strong>of</strong> several series from various localities is needed to understand<br />
geographic variability. The presented data suggest that<br />
two species might be involved, <strong>of</strong> which the identity <strong>of</strong><br />
A. longipinnis is unclear.<br />
10.2.2 Acanthocobitis botia (Hamilton, 1822)<br />
Cobitis botia Hamilton, 1822: 350, 394 (type locality: India:<br />
"northeastern parts <strong>of</strong> Bengal" [Brahmaputra River<br />
at Goalpara; Hora, 1929: 318, 1935a: 49]; types: NT;<br />
Hamilton's unpublished figure reproduced in M'Clelland,<br />
1839: pl. 51 fig. 4; noun in apposition, indeclinable)<br />
Cobitis turio Hamilton, 1822: 358, 395 (type locality: India:<br />
Brahmaputra River [at Goalpara; Hora, 1935a: 49]; types:<br />
NT; Hamilton's unpublished figure reproduced in<br />
M'Clelland, 1839: pl. 52 fig. 7; simultaneous subjective<br />
synonym <strong>of</strong> Cobitis botia Hamilton, 1822: 350; first reviser<br />
[Hora, 1935a: 52] gave precedence to C. botia; noun<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
75<br />
in apposition, indeclinable)<br />
Cobitis bilturio Hamilton, 1822: 358, 395 (type locality:<br />
India: "along with the 10th species" [=C. turio; Brahmaputra<br />
at Goalpara; Hora, 1935a: 49]; types: NT; Hamilton's<br />
unpublished figure reproduced in M'Clelland, 1839:<br />
pl. 51 fig. 6; simultaneous subjective synonym <strong>of</strong> Cobitis<br />
botia Hamilton, 1822: 350; first reviser [apparently<br />
Günther, 1868: 349] gave precedence to C. botia; simultaneous<br />
subjective synonym <strong>of</strong> Cobitis turio Hamilton,<br />
1822: 358; as first reviser, I give precedence to C. turio;<br />
noun in apposition, indeclinable)<br />
Cobitis bimucronata M'Clelland, 1839: 304, 435, pl. 51 fig. 4<br />
(unnecessary replacement name for Cobitis botia Hamilton,<br />
1822: 350; adjective, -us, -a, -um)<br />
Cobitis ocellata M'Clelland, 1839: 304, 436, pl. 51 fig. 6<br />
(unnecessary replacement name for Cobitis bilturio<br />
Hamilton, 1822: 358; adjective, -us, -a, -um)<br />
Cobitis gibbosa M'Clelland, 1839: 304, 436, pl. 52 fig. 7<br />
(unnecessary replacement name for Cobitis turio Hamilton,<br />
1822: 358; adjective, -us, -a, -um)<br />
Cobitis monocera M'Clelland, 1839: 305, 438, pl. 52 fig. 2<br />
(type locality: India: Assam; types: LU; noun in apposition,<br />
indeclinable)<br />
Cobites argentata Swainson, 1839: 310 (available by indication<br />
to Hamilton, 1822: 358, No. 10 [which is Cobitis<br />
turio]; type locality: India: Brahmaputra River [at Goalpara;<br />
Hora, 1935a: 49]; types: NT; Hamilton's unpublished<br />
figure reproduced in M'Clelland, 1839: pl. 52<br />
fig. 7; adjective, -us, -a, -um)<br />
Canthophrys unispina Swainson, 1839: 311 (available by<br />
indication to Hamilton, 1822: 350, No. 1 [which is Cobitis<br />
botia]; type locality: India: "northeastern parts <strong>of</strong> Bengal"<br />
[Brahmaputra River at Goalpara; Hora, 1929: 318,<br />
1935: 49]; types: NT; Hamilton's unpublished figure reproduced<br />
in M'Clelland, 1839: pl. 51 fig. 4; compound<br />
noun, indeclinable)<br />
Cobitis arenata Valenciennes, in Jacquemont, 1839: pl. 15<br />
fig. 1 (type locality: India; holotype: MNHN 3811, Fang,<br />
1943: 404, Daget, 1984: 512; adjective, -us, -a, -um)<br />
Nemachilus mackenziei Chaudhuri, 1910b: 183 (type locality:<br />
India: Uttar Pradesh: Cheriyadang near Kathgodam /<br />
Jaulasal in Naini Tal District / Bengal: Jharai and Jamwari<br />
Nadi near Siripur, Saran / Jhil at Purnahia, P. O.<br />
Ghorasan, Champaran District; syntypes [total 12]: ZSI<br />
F 2017/1 [1], F 4170/1–4171/1 [2], F 4172/1–4173/1 [2],<br />
Menon & Yazdani, 1968: 123; noun in genitive, indeclinable)<br />
10.2.3 Acanthocobitis mandalayensis (Rendahl, 1948)<br />
Nemacheilus rubidipinnis mandalayensis Rendahl, 1948: 21,<br />
fig. 7 A, B (type locality: Burma: Mandalay; holotype:<br />
NRM 13179 [ex MAL/1935139.3179], Kottelat, 1990a:<br />
30]; adjective, -is, -is, -e)<br />
Taxonomic notes. I earlier treated A. mandalayensis as a<br />
synonym <strong>of</strong> A . botia (Kottelat, 1990a: 29), on the basis <strong>of</strong><br />
the material then available. I have since examined a number<br />
<strong>of</strong> samples and photographs <strong>of</strong> A. botia from the Brahmaputra,<br />
Ganges and Indus drainages. Acanthocobitis mandalayensis<br />
is distinguished from both A. botia and A. rubidipinnis<br />
by a unique colour pattern (Kottelat, 2012a: 50, fig. 5).
10.1.1 Aborichthys elongatus, CMK 18565, 52.3 mm SL; India:<br />
Brahmaputra drainage: Tista.<br />
10.2.5 Acanthocobitis pavonacea, CMK 5927, 60.6 mm SL; India:<br />
Brahmaputra drainage: Dibru.<br />
10.2.4 Acanthocobitis mooreh (Sykes, 1839)<br />
Cobitis Mooreh Sykes, 1839a: 162 (type locality: India:<br />
Deccan [Mota Mola River at Poona; Sykes, 1841: 366<br />
[Pune, 18°28'N 73°48'E]]; types: NT; also in Sykes,<br />
1839b: 59, 1841: 366; noun in apposition, indeclinable)<br />
Nemacheilus sinuatus Day, 1870b: 371 (type locality: India:<br />
Wynaad; syntypes: probably BMNH 1870.5.2.17–<br />
18 [2]; adjective, -us, -a, -um)<br />
Nemacheilus aureus Day, 1872: 184 (type locality: India:<br />
Jabbalpúr; syntypes: ZSI 2574 [lost], BMNH<br />
1889.2.1.1587–1590 [4], Whitehead & Talwar, 1976:<br />
156; adjective, -us, -a, -um)<br />
10.2.5 Acanthocobitis pavonacea (M'Clelland, 1839)<br />
Cobitis pavonacea M'Clelland, 1839: 305, 437, pl. 52 fig. 1<br />
(type locality: India: Assam; syntypes: ? SMF 68 [1],<br />
9070 [1], 9090 [1], 9091 [1], Grant, 2007c: fig. 5; adjective,<br />
-us, -a, -um)<br />
10.2.6 Acanthocobitis pictilis Kottelat, 2012<br />
Acanthocobitis pictilis Kottelat, 2012a: 45, fig. 1 (type locality:<br />
Myanmar: Kayin State: stream 'Chon Son' between<br />
Kyondaw and Phadaw, about 20 km northwest <strong>of</strong><br />
Payathouzu (at border with Thailand near Chedi Sam<br />
Ong, Three Pagoda Pass); approx. 15°25'N 98°15'E;<br />
holotype: MHNG 2727.066; adjective, -is, -is, -e)<br />
10.2.7 Acanthocobitis rubidipinnis (Blyth, 1860)<br />
Cobitis rubidipinnis Blyth, 1860: 170 (type locality: Burma:<br />
Rangoon [original type locality: Burma: Tenasserim<br />
provinces]; neotype: NRM 13743 [ex NRM MAL<br />
1935809.3743], designated by Kottelat, 1990a: 35; noun<br />
in apposition, indeclinable)<br />
Cobitis semizonata Blyth, 1860: 171 (type locality: Burma:<br />
Rangoon [original type locality: Burma: Tenasserim provinces];<br />
neotype: NRM 13743 [ex MAL/1935809.3743],<br />
designated by Kottelat, 1990a: 35; simultaneous objective<br />
synonym <strong>of</strong> Cobitis rubidipinnis Blyth, 1860: 170;<br />
first reviser [apparently Day, 1878a: 614] gave precedence<br />
to C. rubidipinnis; adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
76<br />
10.2.9 Acanthocobitis zonalternans, CMK 14633, 40.0 mm SL;<br />
Thailand: Salween drainage: Mae Nam Moei.<br />
10.2.6 Acanthocobitis pictilis, CMK 17509, 69.6 mm SL.<br />
10.2.8 Acanthocobitis urophthalma (Günther, 1868)<br />
Nemachilus urophthalmus Günther, 1868: 348 (type locality:<br />
Southern Ceylon; syntypes: BMNH 1864.7.11.35–<br />
36 [2]; compound adjective, -us, -a, -um)<br />
10.2.9 Acanthocobitis zonalternans (Blyth, 1860)<br />
? Cobitis chlorosoma M'Clelland, 1839: 305, 437, pl. 52<br />
fig. 3 (type locality: India: Upper Assam; types: LU; noun<br />
in apposition, indeclinable)<br />
Cobitis zonalternans Blyth, 1860: 172 (type locality: Thailand:<br />
Tak Province: Huai Mae Charno, 4 km south <strong>of</strong><br />
Amphoe Mae Ramat, 16°58'N 98°34'E [original type<br />
locality: Burma: Tenasserim provinces]; neotype: ZSM<br />
27468, designated by Kottelat, 1990a: 42; participle, indeclinable)<br />
? Noemacheilus phuketensis Klausewitz, 1957a: 195, fig. 1,<br />
pl. 18 fig. 1 (type locality: Thailand: Phuket Island; holotype:<br />
SMF 3966; adjective, -is, -is, -e)<br />
Taxonomic notes. Several species are presently confused<br />
under the name A. zonalternans.<br />
10.3 Afronemacheilus Golubtsov & Prok<strong>of</strong>iev, in Prok<strong>of</strong>iev,<br />
2009<br />
Afronemacheilus Golubtsov & Prok<strong>of</strong>iev, in Prok<strong>of</strong>iev, 2009b:<br />
881 (type species: Nemachilus abyssinicus Boulenger,<br />
1902: 437, by original designation). Gender masculine.<br />
Nomen nudum<br />
Afronemacheilus kaffa Prok<strong>of</strong>iev, 2010: 841, 844, 870<br />
(nomen nudum)<br />
10.3.1 Afronemacheilus abyssinicus (Boulenger, 1902)<br />
Nemachilus abyssinicus Boulenger, 1902: 437 (type locality:<br />
Ethiopia: Lake Tana: Behardar; holotype: BMNH<br />
1902.12.13.435; adjective, -us, -a, -um)
10.3.1 Afronemacheilus abyssinicus, 39.7 mm SL; Ethiopia: Lake<br />
Tana: Blue Nile source. (Photograph by Alexander Golubtsov).<br />
10.4 Barbatula Linck, 1790<br />
Barbatula Linck, 1790: 38 (type species: Cobitis barbatula<br />
Linnaeus, 1758: 303, by absolute tautonymy). Gender<br />
feminine.<br />
Cobites Swainson, 1839: 190, 310 (type species: Cobitis<br />
barbatula Linnaeus, 1758: 303, by monotypy). Gender<br />
feminine.<br />
Orthrias Jordan & Fowler, 1903: 769 (type species: Orthrias<br />
oreas Jordan & Fowler, 1903: 769, by original designation).<br />
Gender masculine.<br />
Nomenclatural notes. Bis repetita placent. Linck (1790)<br />
proposed a classification <strong>of</strong> fishes based on teeth. He split<br />
the Cobitis <strong>of</strong> previous authors into two genera: Cobitis and<br />
Barbatula. Linck used the binominal system and Barbatula<br />
is clearly intended as a generic name. Linck included a single<br />
species in Cobitis (C. heteroclita, p. 33) and two species<br />
in Barbatula (C. barbatula, C. taenia). Although there is no<br />
generic diagnosis, the name is available by indication [Code<br />
art. 12.2.5] as two available names are included. Linck wrote:<br />
"Barbatula. Hierher gehören Cobitis Barbatula Taenia"<br />
[Barbatula. Here belong [plural] Cobitis Barbatula Taenia].<br />
Some have argued that "Cobitis Barbatula Taenia" means<br />
that only one species was included, that this species is<br />
C. taenia and that C. taenia is type species <strong>of</strong> Barbatula.<br />
This is erroneous. Linck used "gehören" [belong], which is<br />
a plural and means that the subject <strong>of</strong> the verb is a plural,<br />
thus that he placed more than one species in Barbatula, and<br />
that these species are barbatula and taenia. If he had included<br />
a single species he would have used the singular "gehört"<br />
[belongs]. Therefore C. barbatula is type species <strong>of</strong> Barbatula<br />
by absolute tautonymy.<br />
Today, "Cobitis Barbatula Taenia" may appear as a<br />
strange combination but not in the context <strong>of</strong> the late 18 th<br />
century, when there were no nomenclature rules, when writing<br />
was less formalised, and when typographical or typesetting<br />
errors were not rare.<br />
Some authors have attributed the authorship <strong>of</strong> Barbatula<br />
to Jordan (1917: 49). As seen from the above, this is not<br />
correct. In addition, a 'Barbatula Jordan, 1917' would be a<br />
junior homonym <strong>of</strong> Barbatula Lesson, 1837: 292 (in Aves)<br />
and is not available (Some authors refer to Lesson, 1831:<br />
164 but Barbatula is not used in that work, only the vernacular<br />
name 'barbion').<br />
10.4.1 Barbatula altayensis Zhu, 1992<br />
Barbatula altayensis Zhu, 1992: 241, figs. 1–2 (type locality:<br />
China: Xinjiang: Kelang He, tributary <strong>of</strong> Ertix He [Irtysh],<br />
near Altay City, 47°52'N 88°06'E; holotype: NGI<br />
888006; adjective, -is, -is, -e)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
77<br />
10.4.2 Barbatula barbatula, CMK 16892, 56.8 mm SL; Switzerland:<br />
Jura: Rhône drainage.<br />
10.4.13 Barbatula sturanyi, CMK 19463, 45.4 mm SL.<br />
10.4.2 Barbatula barbatula (Linnaeus, 1758)<br />
Cobitis Barbatula Linnaeus, 1758: 303 (based on specimens<br />
and Artedi [1738: gen. 2 [4], syn. 2, Cobitis tota glabra<br />
...] and Linnaeus [1746: 125, n. 332, idem]; type locality:<br />
Europae, Asiae, aquis dulcibus; syntypes: NRM 68 [3],<br />
Fernholm & Wheeler, 1983: 282 [listed as non-types],<br />
Lundberg & Svanberg, 2010: 157, fig. 4; noun in apposition,<br />
indeclinable [species names with capitalized initial<br />
in Linnaeus (1758) are nouns])<br />
Cobitis Barbatula Parisiensis La Pylaie, 1835: 534 (nomen<br />
nudum; locality: France: Paris)<br />
Cobitis Barbatula Pictava La Pylaie, 1835: 534 (nomen<br />
nudum; locality: France)<br />
Cobitis furstenbergii Bonaparte, 1846: 26 (nomen nudum;<br />
locality: eastern Germany [probably now Poland])<br />
Cobitis Fürstenbergii Heckel & Kner, 1858: 301 (cited in<br />
synonymy, name not available; author indicated as "Fitzinger<br />
(Prodr. Faun. Austr.)", which apparently is Fitzinger<br />
[1832], but the name does not appear in this work)<br />
Cobitis pironae Nardo, 1866: 143 (type locality: Italy: Basso<br />
Friuli; type material: NT ?; noun in genitive, indeclinable)<br />
Cobitis variabilis Günther, 1868: 355 (on a label in collection,<br />
listed in synonymy, name not available; locality:<br />
Czechia: Moravia)<br />
Nemachilus barbatulus var. taurica Kessler, 1877: 171, 271<br />
(type locality: Ukraine: Crimea: Salgir River; syntypes:<br />
ZISP 3029 [3], 3032 [3], Eschmeyer & Fricke, 2010;<br />
adjective, -us, -a, -um)<br />
? Nemachilus barbatulus caucasicus Berg, 1899: 26, 67 (type<br />
locality: Russia: Chechnya: Argun River, a tributary <strong>of</strong><br />
Terek; types: LU; adjective, -us, -a, -um)
Nemachilus barbatulus vardarensis Karaman, 1928: 164<br />
(type locality: FYROM: Vardar River drainage; syntypes:<br />
MMNHS, lost; also in Karaman, 1929: 173; adjective,<br />
-is, -is, -e)<br />
Nemacheilus barbatulus aberr. erythrinna Berg, 1933: 550<br />
(infrasubspecific, name not available)<br />
Nemacheilus barbatulus forma anglicana Băcescu-Mester,<br />
1967: 369 (infrasubspecific, name not available; locality:<br />
England: Cambridge River)<br />
Nemacheilus barbatulus forma blackiana Băcescu-Mester,<br />
1967: 369 (infrasubspecific, name not available; locality:<br />
England: Black Beck River)<br />
Taxonomic notes. Nemachilus barbatulus caucasicus is<br />
<strong>of</strong>ten treated as a valid subspecies <strong>of</strong> B. barbatula, but up to<br />
now its status does not seem to have been critically evaluated.<br />
Nomenclatural notes. Barbatula is usually treated as an<br />
adjective but in fact it is a noun and indeclinable.<br />
The article with the description <strong>of</strong> N. b. caucasicus had<br />
been issued in two versions, in Russian and in German. The<br />
German version is sometimes dated 1898. The preface is<br />
dated May 1899, so that publication year cannot be 1898.<br />
10.4.3 ? Barbatula cobdonensis (Gundriser, 1973)<br />
Nemachilus cobdonensis Gundriser, 1973: 77 (type locality:<br />
Russia: Tuva Republic: Lakes Khintiktig-Khol<br />
[50.35°N 89.83°E] and Ak-Khol [50°15'11"N<br />
89°36'01"E] and Mogen-Buren River, basin <strong>of</strong> Kobdo<br />
River [Khovd]; syntypes [36]: LU, Prok<strong>of</strong>iev, 2003a: 700;<br />
adjective, -is, -is, -e)<br />
Taxonomic notes. Prok<strong>of</strong>iev (2003a: 700) commented that<br />
there are two species <strong>of</strong> nemacheilids in the Mogen Buren<br />
system near Mugur-Aksy [50°23'N 90°26'E], one he identified<br />
as B. toni and the other he described as B. golubtsovi.<br />
He commented that, judging from the information in the original<br />
description, the type series <strong>of</strong> N. cobdoensis seems to<br />
include two species. Because the whereabouts <strong>of</strong> the type<br />
series are not known, and because the later description and<br />
photograph in Gundriser (1979: 27–30) do not fit with his<br />
B. golubtsovi, he concluded that either N. cobdonensis is<br />
identical with his B. 'toni' or belongs to a third species. As<br />
discussed below, I consider B. toni to be restricted to the<br />
Amur drainage. The name B. cobdonensis is available for<br />
the B. 'toni' endemic to the Khovd drainage. This will be<br />
fixed later by a neotype designation.<br />
10.4.4 ? Barbatula compressirostris (Warpachowski, 1897)<br />
Nemachilus compressirostris Warpachowski, 1897: 270, pl.<br />
11 figs. 1–3 (type locality: Russia (?): Ob drainage, lakes<br />
in northwestern Mongolia; syntypes: ZISP 11298 [2];<br />
compound noun, indeclinable)<br />
Taxonomic notes. Nemachilus compressirostris was originally<br />
based on two specimens from "lakes in northwestern<br />
Mongolia" in a paper on fishes <strong>of</strong> the Ob River drainage<br />
(Warpachowski, 1897). Warpachowski also commented that<br />
the specimens came from the same lakes as Oreoleuciscus<br />
potanini. These specimens were obtained for the Nizhniy<br />
Novgorod Fisheries Exhibition in 1896. It is not clear (to<br />
me) if this was intended to mean the same lakes as the<br />
O. potanini specimens mentioned in the same paper (p. 263:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
78<br />
Russia: Altai: Kosh Agash, 50°59'30"N 88°40'00"E, in Ob<br />
drainage) or those lakes from which O. potanini was known<br />
in Mongolia at that time. I had earlier (Kottelat, 2006: 51,<br />
103) concluded that the type locality was in western Mongolia,<br />
more precisely in the Khovd basin, and that it could<br />
potentially be a senior synonym <strong>of</strong> B. golubtsovi, which now<br />
seems unlikely. This nominal species is considered to be a<br />
synonym <strong>of</strong> B. toni by, e.g. Prok<strong>of</strong>iev (2007a: 17). I consider<br />
here B. toni to be restricted to the Amur drainage. The<br />
identity <strong>of</strong> B. compressirostris can be established only when<br />
comparison material becomes available for all three drainages<br />
in that area (upper Ob, upper Yenisei and Khovd). Its<br />
identity must be tested against B. tomiana, B. markakulensis<br />
and B. cobdoensis.<br />
10.4.5 Barbatula dgebuadzei (Prok<strong>of</strong>iev, 2003)<br />
Orthrias dgebuadzei Prok<strong>of</strong>iev, 2003a: 730, fig. 2 [p. 700<br />
<strong>of</strong> translation] (type locality: Mongolia: Zag River, Baidrag-Gol<br />
drainage; holotype: ZISP 52051; noun in genitive,<br />
indeclinable)<br />
10.4.6 Barbatula golubtsovi (Prok<strong>of</strong>iev, 2003)<br />
Orthrias golubtsovi Prok<strong>of</strong>iev, 2003a: 727, fig. 1 [p. 698 <strong>of</strong><br />
translation] (type locality: Russia: Tuva Republic: Chedi-Tei<br />
River, outflow <strong>of</strong> Ak-Khol Lake [50°15'11"N<br />
89°36'01"E], Mogen-Buren drainage part <strong>of</strong> Kobdo<br />
drainage, about 60 km west <strong>of</strong> Mugur-Aksy [50°23'N<br />
90°26'E]; holotype: ZISP 52251; noun in genitive, indeclinable)<br />
Taxonomic notes. Molecular data suggests this species belong<br />
to Triplophysa (V. Šlechtová, pers. comm.).<br />
10.4.7 ? Barbatula markakulensis (Menshikov, 1939)<br />
Nemacheilus barbatulus markakulensis Menshikov, 1939:<br />
141, fig. (type locality: Kazakhstan: Lake Marka-kul, Irtysh<br />
drainage; syntypes: ZISP 26864 [6+], Eschmeyer &<br />
Fricke, 2010; adjective, -is, -is, -e)<br />
Taxonomic notes. See under B. toni.<br />
10.4.8 Barbatula nuda (Bleeker, 1864)<br />
Nemacheilus nudus Bleeker, 1864c: 12 (type locality: Mongolia<br />
[p. 13, but 'brought from China' on p. 14; apparently<br />
China: Hebei: Chongli County: Siwantse [Xiwanzi],<br />
40°58'25"N 115°16'22"E; see below]; holotype: MNHN<br />
1450, Fang, 1941: 253, Bertin & Estève, 1948: 98; adjective,<br />
-us, -a, -um)<br />
? Nemachilus pechiliensis Fowler, 1899: 181 (type locality:<br />
China: Pechili: Tan Lan Ho, tributary <strong>of</strong> Shu Lan Ho,<br />
about 30 miles northeast <strong>of</strong> Lama-Miau or Dolon-Nor<br />
[Nei Mongol: Duolun County: Duolun-Nuur, 42°11'22"N<br />
116°28'24"E] [Pechili [Zhili], name <strong>of</strong> a former province];<br />
holotype: ANSP 16394 [missing, Böhlke, 1984:<br />
63]; adjective, -is, -is, -e)<br />
? Barbatula toni fowleri Nichols, 1925b: 3 (type locality:<br />
China: Chihli Province: Eastern Tombs [now Hebei: Zunhua,<br />
40°11'N 117°58'E; Chihli [Zhili], name <strong>of</strong> a former<br />
province]; holotype: AMNH 8409; noun in genitive, indeclinable)<br />
Barbatula stoliczkai shansi Nichols, 1925b: 6 (type locality:<br />
China: Shansi [now in Nei Monggol]: Mai-tai-chao,
42 miles east <strong>of</strong> Paotow [Baotou, 40°40'N 109°50'E];<br />
holotype: AMNH 8412; noun in apposition, indeclinable)<br />
? Barbatula toni kirinensis Tchang, 1932a: 115, fig. 2 (China:<br />
Kirin [= Jilin Province]; holotype: ZMFIB 7931; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. The identity <strong>of</strong> N. nudus Bleeker (1864c)<br />
is not clear; authors have placed it as a senior synonym <strong>of</strong><br />
either B. toni (Dybowski, 1869; sensu lato) or Triplophysa<br />
stolickai (Steindachner, 1866). In the last 30 years Chinese<br />
and Korean authors (e.g. Zhu, 1989: 29; Wang et al., 2001:<br />
168; Kim, 1997: 283) have used the name B. nuda for the<br />
B. toni <strong>of</strong> earlier authors and B. nuda became the same catchall<br />
name (see under B. toni). The type locality <strong>of</strong> B. nuda is<br />
usually listed as Mongolia. Indeed, Bleeker (1864c: 13) indicated<br />
the type locality as Mongolia, which at that time could<br />
have meant present-day Mongolia, or Nei Mongol and Xinjiang<br />
provinces <strong>of</strong> China, or some other place in northern<br />
China. But on p. 14, he wrote "described from a single specimen<br />
and brought from China by the missionary David". This<br />
holotype still exists (MNHN 1450, Bertin & Estève, 1948:<br />
98); I examined it in the early 1980s, before I became familiar<br />
with the East Asian Barbatula; since then, I have not had<br />
an opportunity to compare it side by side with Chinese,<br />
Mongolian or Korean material identified as B. nuda. Nevertheless,<br />
it is not conspecific with B. toni as recognised here<br />
(restricted to upper Amur drainage).<br />
The specimen was collected by Armand David (27 September<br />
1826–10 November 1900; for a biography, see Boutan,<br />
1993). David was a catholic missionary and stayed in<br />
China from 1862 to 1874 and visited various areas <strong>of</strong> the<br />
country. He travelled in 'southern Mongolia' [now in Nei<br />
Mongol Province <strong>of</strong> China] for the first time in 1866 (David,<br />
1867–1868; Scott, 2004: 70). As Bleeker described<br />
B. nuda in 1864, the specimen had been collected earlier.<br />
But, at that time Mongolia was a complex entity. A Greater<br />
Mongolia was administered as Outer Mongolia [now more<br />
or less corresponding to Mongolia], Inner Mongolia [now<br />
more or less corresponding to Nei Mongol] and a number <strong>of</strong><br />
leagues (former Mongolian administrative units) and banners<br />
(Mongolian/Chinese military and administrative units)<br />
in various adjacent provinces <strong>of</strong> China. The provinces, leagues<br />
and banners were overlapping but the limits <strong>of</strong> their<br />
respective jurisdictions were not always identical.<br />
David's first travel, in 1862, was a brief visit <strong>of</strong> the area<br />
<strong>of</strong> Siwantze, 25 km northeast <strong>of</strong> Kalgan [Zhangjiakou,<br />
40°49'N 114°53'E, 160 km northwest <strong>of</strong> Beijing]. Although<br />
in then Zhili Province, Zhangjiakou was seat <strong>of</strong> the military<br />
commander <strong>of</strong> the Eight Banners <strong>of</strong> Chahar, part <strong>of</strong> Greater<br />
Mongolia. Siwantze is Xiwanzi [40°58'25"N 115°16'22"E],<br />
now in Chongli County in Hebei Province. In the catholic<br />
administration, Siwantze was the see <strong>of</strong> a diocese (an administrative<br />
unit), at David's time called the Apostolic Vicariate<br />
<strong>of</strong> Mongolia. It was reasonable for a catholic missionary<br />
to label material from Xiwanzi as from 'Mongolia'.<br />
In 1863, David explored the mountains bordering the west<br />
<strong>of</strong> the plain <strong>of</strong> Beijing; this cannot be called 'Mongolia'. In<br />
1864, David travelled in Jehol, north-east <strong>of</strong> Beijing (map<br />
in David, 1875), and this was too late to have material arrived<br />
in MNHN and described by Bleeker in 1864. Jehol is<br />
a former Chinese province that included part <strong>of</strong> today's He-<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
79<br />
bei, Shanxi and Nei Mongol Provinces. This excludes<br />
present-day Mongolia as the type locality and suggest that<br />
the holotype originates from the area <strong>of</strong> Xiwanzi.<br />
Bleeker (1864c) explicitly stated that his description <strong>of</strong><br />
N. nudus was based on a specimen 115 mm TL in MNHN.<br />
Fang (1941: 253) commented that MNHN has four specimens<br />
labelled "Nemacheilus nudus, Blkr.; Mongolie: Abbé<br />
David, 1863; fishes that live in the high valleys". Fang mentioned<br />
that two <strong>of</strong> these specimens are the "types" <strong>of</strong> N. nudus<br />
(MNHN 1450); the other two he described as N. bertini<br />
(then MNHN 3800, now MNHN 3800 and B.2640). The<br />
two specimens in MNHN 1450 are the holotype and a nontype<br />
specimen <strong>of</strong> N. nudus (now possibly MNHN 3813).<br />
Herzenstein (1888: 21), Rendahl (1933: 46) and Prok<strong>of</strong>iev<br />
(2003a: 703) discussed ZISP 4471 received from MNHN<br />
as N. nudus from Sichuan. Fang (1941: 253) explained that<br />
this specimen had been taken from a jar labelled "Nemacheilus<br />
nudus Blkr.; Sse-chuan Occid.; R. P. A. David, 1870–<br />
58". He described the remaining 11 specimens (MNHN 6287)<br />
as Nemacheilus angeli.<br />
I have examined a number <strong>of</strong> samples previously identified<br />
as B. nuda from northern China (see Kottelat, 2006:<br />
53). They represent several species <strong>of</strong> Barbatula and Triplophysa;<br />
at some localities two species occur in sympatry.<br />
Names are probably available for some <strong>of</strong> them (see synonymy),<br />
but others are apparently still undescribed. Without<br />
access to more material from more localities it is not possible<br />
to clarify how many species are involved and what are<br />
their diagnostic characters, so as to determine which is the<br />
real B. nuda.<br />
10.4.9 Barbatula oreas (Jordan & Fowler, 1903)<br />
Orthrias oreas Jordan & Fowler, 1903: 769, fig. 2 (type locality:<br />
Japan: Hokkaido: Chitose, in Iburi; holotype: "the<br />
museum at Sapparo"; noun in apposition, indeclinable)<br />
10.4.10 Barbatula potaninorum (Prok<strong>of</strong>iev, 2007)<br />
Orthrias potaninorum Prok<strong>of</strong>iev, 2007a: 65, fig. 18 (type<br />
locality: "China ? (exact location not given on label; according<br />
to the itinerary <strong>of</strong> the expedition - Northern China,<br />
Gan’su Province at the border with Mongolia, upper<br />
part <strong>of</strong> the Edzin-Gol River system - Fig. 23)" [China ?:<br />
Gansu: Edsin-Gol [Ejin, Ejina He, Eiin, Hei-He, Ruo-<br />
Shui; a river flowing north to endorheic Gaxun Nuur (Juyan<br />
Lake), 42°25'20"N 100°40'07"E]]; holotype: ZISP<br />
"8121" [erroneous]; noun in genitive, indeclinable)<br />
10.4.11 Barbatula quignardi (Bacescu-Mester, 1967)<br />
Noemacheilus barbatulus quignardi Băcescu-Mester, 1967:<br />
359, figs. 1, 5c–d (type locality: France: Le Lez stream<br />
near Montpellier; holotype: MGAB 77; noun in genitive,<br />
indeclinable)<br />
Nemacheilus barbatulus forma hispanica Băcescu-Mester,<br />
1967: 369, fig. 6a–b (infrasubspecific, name not available;<br />
locality: Spain: Nervion River at Durango / Tajo River)<br />
Nemacheilus barbatulus hispanica Lelek, 1987: 256 (available<br />
by indication to Băcescu-Mester, 1967; type locality:<br />
Spain: Nervion River at Durango / Tajo River; syntypes:<br />
MGAB [11, material used by Băcescu-Mester,<br />
1967: 369]; adjective, -us, -a, -um)
10.4.8 Barbatula ‘nuda’, China: Jilin: Antu. Two syntopic species.<br />
Above, with numerous papillae on body (ASIZB 132651,<br />
75.4 mm SL, possibly real B. nuda); below, with smooth body<br />
(ASIZB 132648, 101.6 mm SL; topotype <strong>of</strong> B. kirinensis). See<br />
also differences in nostrils and respective positions <strong>of</strong> dorsal and<br />
pelvic fins.<br />
10.4.8 Barbatula nuda, MNHN 1450, 115 mm TL, holotype; China:<br />
Hebei: Xiwanzi.<br />
10.4.12 Barbatula sawadai (Prok<strong>of</strong>iev, 2007)<br />
Orthrias sawadai Prok<strong>of</strong>iev, 2007a: 84, fig. 26 (type locality:<br />
Mongolia: Yenisei River drainage: Lake Baikal basin:<br />
Selenge River system: Ero River catchment: stream<br />
Uram; holotype: ZISP 52054; noun in genitive, indeclinable)<br />
10.4.13 Barbatula sturanyi (Steindachner, 1892)<br />
Nemachilus Sturanyi Steindachner, 1892a: 131 (type locality:<br />
FYROM: Lake Ohrid at Pestani, between Ohrid City<br />
and Sveti Naum Monastery; also in Steindachner, 1892b:<br />
378, pl. 2 fig. 3; holotype: NMW 48440; noun in genitive,<br />
indeclinable)<br />
10.4.14 Barbatula tomiana (Ruzsky, 1920)<br />
? Nemacheilus sibiricus Gratzianow, 1907: 167, 168 (type<br />
locality: Russia: Altay: Bija [Biya] River near Bijsk<br />
[Biysk; 52°31'30"N 85°13'30'E]; syntypes: ZMMU<br />
P-2532 [3], Vasil'eva et al., in Pavlinov & Borissenko,<br />
2001: 25; adjective, -us, -a, -um)<br />
Nemacheilus barbatulus tomianus Ruzsky, 1920: 36, figs.<br />
1–2 (type locality: Russia: Siberia: Tomsk oblast: Tom<br />
River at Tomsk, Kuznetsk and Kondoma / brook Kanzas,<br />
a right-hand tributary <strong>of</strong> Balyk-Su, upper reaches <strong>of</strong><br />
Tom River / Ushayka River / Kislovka River near<br />
Tomsk / upper Torsya / upper reaches <strong>of</strong> Ob itself in area<br />
<strong>of</strong> Altay (rivers Katun, Cherga, Ursul, Charysh) / in Yenissei<br />
(rivers Abakan and Minusinka); syntypes: LU; adjective,<br />
-us, -a, -um)<br />
? Barbatula barbatula toni morpha tigris Gundriser, 1975:<br />
24 (infrasubspecific, name not available; locality: Russia:<br />
Tuva: lake and bassin <strong>of</strong> Ii-Kem River [maybe Khemchik,<br />
in Yenisei drainage; 51°16'40"N 90°47'00"E ?])<br />
Taxonomic notes. See under B. toni.<br />
10.4.15 Barbatula toni (Dybowski, 1869)<br />
Cobitis Toni Dybowski, 1869: 957, pl. 18 fig. 10 (type locality:<br />
"common in both river systems" [Russia: Zabaykal-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
80<br />
10.4.14 Barbatula tomiana, CMK 22367, 118.2 mm SL; Russia:<br />
Altai: Ob drainage: Sema.<br />
10.4.15 Barbatula toni, CMK 19616, 79.9 mm SL; Mongolia:<br />
Amur drainage: Onon.<br />
sky Krai [Transbaikal]: Onon and Ingoda Rivers, Amur<br />
River drainage]; syntypes: ZISP ?, MNHN B.738 [8],<br />
B.739 [4], NMW 50473 [2], Bertin & Estève, 1948: 98,<br />
Eschmeyer & Fricke, 2010; noun in genitive, indeclinable)<br />
Taxonomic notes. Obviously a heterogeneous assemblage<br />
<strong>of</strong> several species. Considered as a subspecies <strong>of</strong> B. barbatula<br />
by Prok<strong>of</strong>iev (2003a, 2007a) but the scale <strong>of</strong> analysis is<br />
not detailed enough to be informative. Prok<strong>of</strong>iev (2007a)<br />
recognised a subspecies B. b. toni extending from Japan to<br />
the Urals, with a broad range <strong>of</strong> morphological variation. I<br />
identify as B. toni the species that seems to be restricted to<br />
the upper Amur drainage (the type locality), recognised by<br />
the backward position <strong>of</strong> the dorsal fin, the sharply decreasing<br />
body depth behind the dorsal-fin base, and the distinctive<br />
colour pattern with very irregular bars across the back,<br />
on a yellowish background (see, e.g. Bogutskaya et al., 2008:<br />
339; Kottelat, 2006: 99). I have examined B. 'toni' only from<br />
a few places <strong>of</strong> its range and I have not yet a complete overview<br />
<strong>of</strong> its variability. But my observations on populations<br />
from Mongolia, Korea and northern China show that the<br />
Selenge (Yenisei drainage) has a distinct species (figured as<br />
"B. b. t. infrasubsp. tomianus" by Berg, 1933: 552, 1948:<br />
871). I tentatively retain the name B. tomiana for it, but<br />
N. sibiricus might be an older available name. The B. 'toni'<br />
from the upper Irtysh (Bulgan River) in Mongolia is distinct<br />
and I tentatively retain the name B. markakulensis for it; this<br />
remains to be confirmed by comparison with fresh material<br />
from Markakul Lake. The B. toni from the eastern slope <strong>of</strong><br />
Korea (e.g. Kim & Park, 2007: 193) is apparently the same<br />
as the unnamed species figured by Bogutskaya et al. (2008:<br />
339) and possibly conspecific with populations from Hokkaido,<br />
then B. oreas is the oldest name availabe for it. See<br />
under B. nuda for populations from the western slope <strong>of</strong><br />
Korea and northern China.<br />
10.4.16 Barbatula zetensis (Soric, 2001)<br />
Orthrias barbatulus zetensis Šorić, 2001: 66, fig. 1 (type<br />
locality: Montenegro: Vis brook near Vis village, 5–6 km<br />
west <strong>of</strong> Danilovgrad, tributary <strong>of</strong> Zeta River; holotype:<br />
BIKU; adjective, -is, -is, -e)<br />
10.5 Claea Kottelat, 2011<br />
Oreias Sauvage, 1874: 334 (type species: Oreias dabryi Sau-
10.5.1 Claea dabryi, CMK 13064, 67.8 mm SL; China: Sichuan:<br />
Yangtze drainage: Yalong.<br />
10.5.1 Claea dabryi, MNHN 6276, about 122 mm SL, holotype;<br />
China: Sichuan: Yangtze drainage: Dengchigou.<br />
vage, 1874: 334, by monotypy; a junior synonym <strong>of</strong><br />
Oreias Kaup, 1829: 177, 193 in Aves and Oreias Temminck,<br />
in Temminck & Laugier de Chartrouse, 1839: 108<br />
in Aves). Gender feminine.<br />
Claea Kottelat, 2011a: 384 (replacement name for Oreias<br />
Sauvage, 1874: 334). Gender feminine.<br />
10.5.1 Claea dabryi (Sauvage, 1874)<br />
Oreias dabryi Sauvage, 1874: 334 (type locality: China: eastern<br />
Tibet: torrent at Yao-Tchy [Sichuan: Yaoji, Qiaoji,<br />
Qiaoqi, 30°41'53"N 102°44'48"E; in fact apparently<br />
Baoxing County: Dengchigou, 30°32'07"N 102°56'28"E;<br />
see below; David, 1874: 42, 73]; holotype: MNHN 6276,<br />
Bertin & Estève, 1948: 99, Bănărescu & Nalbant, 1976:<br />
187, figs. 1–4, pl. 1 fig. 1 [plate shows holotype apparently<br />
reconstructed or in one piece; now broken, head<br />
connected to body only by intestine]; noun in genitive,<br />
indeclinable)<br />
Oreias crassipedunculatus Bănărescu & Nalbant, 1976: 189,<br />
figs. 9–12, pl. 1 fig. 4, pl. 2 fig. 7 (type locality: China:<br />
Yunnan: Chatung; holotype: USNM 94731; adjective,<br />
-us, -a, -um)<br />
Oreias furcatus Bănărescu & Nalbant, 1976: 188, figs. 5–8,<br />
pl. 1 figs. 2–3, pl. 2 figs. 6, 8 (type locality: China: Sichuan:<br />
Tsao Po, near Kuan-shien [according to USNM<br />
catalogue] [Kuan-hsien, Guanxian, Dujiangyan County;<br />
30°59'27"N 103°36'32"E]; holotype: USNM 130128;<br />
adjective, -us, -a, -um)<br />
Taxonomic notes. This species is placed in Schistura by Zhu<br />
(1989) and Zhou & Cui (1993). The type locality <strong>of</strong> both<br />
Claea dabryi and the sisorid catfish Chimarrichthys davidi<br />
Sauvage, 1874 is given as Yao-Tchy [Yaoji, Qiaoqi]. The<br />
specimens were collected by A. David in 1869. David was in<br />
Mouping [Muping] (an independant Tibetan princedom) from<br />
1 March to 22 November 1869. Muping more or less corresponds<br />
to present Baoxing County in Sichuan. David published<br />
a narrative <strong>of</strong> his stay in Muping and a day-by-day<br />
description <strong>of</strong> his movements, activities and collections (David,<br />
1874). He stayed at the catholic mission in Dengchigou<br />
(30°32'07"N 102°56'28"E); he was repeatedly ill and with a<br />
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10.6.1 Draconectes narinosus, MHNG 2730.080, 24.7 mm SL,<br />
holotype; Vietnam: Ha Long Bay: cave on Dáo Van Giò island.<br />
(Photograph by Tan Heok Hui).<br />
10.7.1 Dzihunia amudarjensis, ZISP 23898, 65 mm SL; Uzbekistan:<br />
Amu-Darya estuary; syntype <strong>of</strong> Nemachilus choresmi. (Photograph<br />
by A. Prok<strong>of</strong>iev).<br />
10.8.2 Eonemachilus nigromaculatus, YU 720, 69.5 mm SL; China:<br />
Yunnan: Lake Dianchi.<br />
single exception he only made one-day trips around Dengchigou,<br />
all on foot. Although he seems to have walked very<br />
far and hunting on the way, he does not seem to have been to<br />
Qiaoqi (at least 25 km away). Local hunters brought him<br />
various material from Qiaoqi but he did not report receiving<br />
fishes from there. But he mentioned (1874: 42) that his hunters<br />
brought him two fish species from "our torrent" [implied,<br />
the torrent in Dengchigou], chepadze ("a siluroid that stucks<br />
on stones using its flat belly, <strong>of</strong> which it makes a sucker";<br />
apparently Chimarrichthys davidi) and si-yu (a cyprinid, likely<br />
a schizothoracine). On p. 66, he mentioned that the fish<br />
had decayed and, on p. 73, that on 9 October 1869 he collected<br />
again, himself, in Dengchigou, the chepadze and silien-yu<br />
and two small species <strong>of</strong> loaches. One <strong>of</strong> the loaches<br />
is most likely C. dabryi. Page 75, David mentioned that a<br />
man sent to fish in the river <strong>of</strong> "Ta-hong-miao", 2 days away,<br />
did not bring anything new. I consider that the type locality<br />
<strong>of</strong> Cl. dabryi and Ch. davidi is Dengchigou, not Qiaoqi.<br />
10.5.2 Claea niulanjiangensis (Chen, Lu & Mao, 2006)<br />
Schistura niulanjiangensis Chen, Lu & Mao, 2006: 54, fig.<br />
(type locality: China: Yunnan: Zhanyi County: Niulan<br />
River (25°59'N 103°36'E) [Yangtze drainage]; holotype:<br />
FACQR 9206085; adjective, -is, -is, -e)<br />
10.6 Draconectes Kottelat, 2012<br />
Draconectes Kottelat, 2012b: 342 (type species: Draconectes<br />
narinosus Kottelat, 2012b: 342, by original designation).<br />
Gender masculine.<br />
10.6.1 Draconectes narinosus Kottelat, 2012<br />
Draconectes narinosus Kottelat, 2012b: 342, fig. 1 (type<br />
locality: Vietnam: Quang Ninh Province: Ha Long Bay:
island Dáo Van Giò: phreatic lake in cave Dông Duc Tiên,<br />
20°50.34'N 107°16.77'E; holotype: MHNG 2730.080;<br />
adjective, -us, -a, -um)<br />
10.7 Dzihunia Prok<strong>of</strong>iev, 2001<br />
Dzihunia Prok<strong>of</strong>iev, 2001b: 209 (type species: Nemacheilus<br />
amudarjensis Rass, 1929: 253, by original designation).<br />
Gender feminine.<br />
10.7.1 Dzihunia amudarjensis (Rass, 1929)<br />
Nemachilus amudarjensis Rass, 1929: 253, figs. 1–2 (type<br />
locality: Amu-Darya (Oxus) near Termes [Uzbekistan:<br />
Termez; 37°13'N 67°17'E], near Tschardschui [Turkemenistan:<br />
Chardjui, Chardzhou, now Türkmenabat;<br />
39°05'N 63°34'E] and upper reaches; syntypes: ZMMU<br />
2554 [14], ZISP 22042 [2], 20715 [2], 20732 [2]; adjective,<br />
-is, -is, -e)<br />
Nemachilus amudarjensis choresmi Berg, 1932a: 149, fig. 2<br />
(type locality: Uzbekistan: Amu-Darya estuary, about 10<br />
km from confluence <strong>of</strong> Taldyk [43°42'N 59°12'E] and<br />
Medeli arms; syntypes [5]: ZISP 23866 [1], 23867 [1],<br />
23868 [1], 23898 [2], Prok<strong>of</strong>iev, 2001b: 210, Eschmeyer<br />
& Fricke, 2010; also in Berg, 1932b: 559, fig. 490;<br />
noun in genitive, indeclinable)<br />
10.7.2 Dzihunia ilan (Turdakov, 1936)<br />
Nemachilus amudarjensis ilan Turdakov, 1936: 207, 210,<br />
figs. 3a-b, 4 (type locality: Uzbekistan: Samarkand Province:<br />
Kara-Darya near Dagbit [Dagbid, Daxbet, Dahbed,<br />
11 km from Samarkand, 39°45'50"N 66°54'45"E], a tributary<br />
<strong>of</strong> Zeravshan River below Samarkand [39°39'15"N<br />
66°57'35"E]; lectotype: ZISP 30501, designated by Prok<strong>of</strong>iev,<br />
2009b: 891; noun in apposition, indeclinable)<br />
10.7.3 Dzihunia turdakovi Prok<strong>of</strong>iev, 2003<br />
Dzihunia turdakovi Prok<strong>of</strong>iev, 2003c: 57, figs. 1–2 (type<br />
locality: Kyrgyzstan: Ters River, Talas basin; holotype:<br />
ZISP 36723; noun in genitive, indeclinable)<br />
10.8 Eonemachilus Berg, 1938<br />
Eonemachilus Berg, 1938: 314 (type species: Nemacheilus<br />
nigromaculatus Regan, 1904: 192, by original designation).<br />
Gender masculine.<br />
Taxonomic notes. Some authors have recognised a Y. nigromaculatus<br />
group within Yunnanilus (e.g. Chen, Yang &<br />
Yang, 2012). Within this group, Y. nigromaculatus and<br />
Y. yangzonghaiensis are very conspicuous, with a deep body,<br />
mouth terminal and somewhat upturned, a colour pattern <strong>of</strong><br />
bold dark bars or elongated blotches on a whitish or pale<br />
yellowish background, and their lacustrine habitat. I consider<br />
them as a distinct genus. Judging from the figure in the<br />
original description, Y. longidorsalis is possibly related to<br />
and tentatively placed in Eonemachilus; it is not lacustrine.<br />
The remaining species <strong>of</strong> the Y. nigromaculatus group do<br />
not seem to constitute a monophyletic group. Some <strong>of</strong> them<br />
may represent one or several new genera and others may be<br />
more related with species now in Heminoemcaheilus. For<br />
the time being, they are all retained in Yunnanilus.<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
82<br />
10.8.1 ? Eonemachilus longidorsalis (Li, Tao & Lu, in<br />
Li, Tao, Mao & Lu, 2000)<br />
Yunnanilus longidorsalis Li, Tao & Lu, in Li, Tao, Mao &<br />
Lu, 2000: 350, figs. 4–6 (type locality: China: Yunnan:<br />
Luoping County: pool Agang Longtan, 25°04'N 104°07'E<br />
[Nanpangjiang drainage]; holotype: HRAS or ASIZB<br />
9411001; compound noun, indeclinable)<br />
10.8.2 Eonemachilus nigromaculatus (Regan, 1904)<br />
Nemachilus nigromaculatus Regan, 1904: 192 (type locality:<br />
China: Yunnan: "Sea <strong>of</strong> Tien" at Yunnan Fu [Lake<br />
Dianchi at Kunming]; lectotype: BMNH 1904.1.26.38,<br />
designated by Kottelat & Chu, 1988a: 68; adjective, -us,<br />
-a, -um)<br />
10.8.3 Eonemachilus yangzonghaiensis (Cao & Zhu, in<br />
Zheng, 1989)<br />
Yunnanilus nigromaculatus yangzonghaiensis Cao & Zhu,<br />
in Zheng, 1989: 45, fig. 23 (type locality: China: Yunnan:<br />
Yangzong-hai Lake; holotype: IHB 63V1074; adjective,<br />
-is, -is, -e)<br />
10.9 Hedinichthys Rendahl, 1933<br />
Hedinichthys Rendahl, 1933: 26 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus yarkandensis<br />
Day, 1877b: 796, by original designation). Gender<br />
masculine.<br />
10.9.1 Hedinichthys grummorum Prok<strong>of</strong>iev, 2010<br />
Hedinichthys grummorum Prok<strong>of</strong>iev, 2010: 897, fig. 26 (type<br />
locality: China: Xinjiang: Lemjin Oasis in northern part<br />
<strong>of</strong> Turpan Depression [Lamjin, 42°52'N 89°58'E]; holotype:<br />
ZISP 9237; noun in genitive, indeclinable)<br />
10.9.2 Hedinichthys macropterus (Herzenstein, 1888)<br />
Nemachilus yarkandensis macropterus Herzenstein, 1888:<br />
79 (type locality: China: Gansu: Ssa-tschsheú [Sachow<br />
Oasis; Dunhuang, 40 10'N 94 50'E] and in the vicinity <strong>of</strong><br />
Mumin city; syntypes: ZISP 7260 [6+], 7845 [5], 7847<br />
[1]; compound adjective, -us, -a, -um)<br />
Nemachilus yarkandensis nordkansuensis Li & Chang, 1974:<br />
416 (type locality: China: Kansu [Gansu]: Ansi-Hsien<br />
[Anxi, now Guazhou County, 39°50'N 97°34'E]: Shulagol<br />
River [Shule He], Shuang-Ta-Bu [Shuangtapu, about<br />
40°33'N 96°10'E]; holotype: ASIZB G65-52; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. Recognised as distinct from H. yarkandensis<br />
following Prok<strong>of</strong>iev (2010: 896), although the text<br />
(or the translation) is ambiguous.<br />
10.9.3 Hedinichthys minutus (Li, in Li, Tai, Chang, Ma,<br />
Ho & Kao, 1966)<br />
Nemachilus minutus Li, in Li, Tai, Chang, Ma, Ho & Kao,<br />
1966: 46, fig. 1 (type locality: China: Sinkiang [Xinjiang]:<br />
Kalagu reservoir, Pichong [Piqan County, 42°52'N 90°<br />
10'E]; holotype: ASIZB 64-1963; adjective, -us, -a, -um)<br />
10.9.4 Hedinichthys yarkandensis (Day, 1877)<br />
Nemacheilus yarkandensis Day, 1877b: 796 (type locality:
China: Xinjiang: Yarkand River drainage: Pas Robat,<br />
Yankihissar [Yengisar, 38°55'33"N 76°10'17"E] and<br />
Kashgar [39°28'N 75°59'E], "all from waters in connection<br />
with the Yarkand and Yangihissar or Great Easterly<br />
River"; syntypes: ZSI 1228–1229 [2], 1471–1472 [2],<br />
BMNH 1889.2.1.1718–1735 [18], NMW 48453, Whitehead<br />
& Talwar, 1976: 157; also in Day, 1878b: 14, pl. 5<br />
fig. 3; adjective, -is, -is, -e)<br />
Nemachilus tarimensis Kessler, 1879: 300 (type locality:<br />
China: Xinjiang: lower Tarim River and Lob-Nor Lake<br />
[Lop Nur]; syntypes: ZISP 4204 [6+], 4205 [6+], according<br />
to Herzenstein, 1888: 74; Kessler indicated "more<br />
than 20"; reprinted as Kessler, 1880: 259; adjective, -is,<br />
-is, -e)<br />
Nemachilus yarkandensis longibarbus Herzenstein, 1888:<br />
78, pl. 4 fig. 1 (type locality: China: Xinjiang: Oasis Nija<br />
[Niya, Minfeng, 37°03'50"N 82°41'20"E]; Dol Oasis<br />
[about 50 km east <strong>of</strong> Hotan; 37°10°N 80°17'E], Sampula<br />
Oasis [Sampul, Shampula, Shanpuluxian, about 50 km<br />
southeast <strong>of</strong> Hotan; 37°01'11"N 80°06'36"E]; Upper<br />
Tarim River; syntypes: ZISP 7298 [4, in part], 7361 [3],<br />
7375 [1] [ZISP 7298, 7361 listed under both N. y. longibarbus<br />
and N. y. brevibarbus]; compound adjective,<br />
-us, -a, -um)<br />
Nemachilus yarkandensis brevibarbus Herzenstein, 1888:<br />
78, pl. 2 fig. 1 (type locality: China: Xinjiang: Aksu-darja<br />
[Aksu He; stream entering Yarkand River from North<br />
at 40°27'46"N 80°52'10"E]; Nija Oasis [Niya, Minfeng,<br />
37°03'50"N 82°41'20"E]; Tschertschen-darja [Charchan-<br />
Darya, Qarqan He stream; Qarqan town or Qiemo County:<br />
38°08'09"N 85°31'48"E; Qarqan He sometimes enters<br />
Lop Nor from SW]; Dol Oasis [about 50 km east <strong>of</strong><br />
Hotan; 37°10°N 80°17'E], Sampula Oasis [Sampul,<br />
Shampula, Shanpuluxian, about 50 km southeast <strong>of</strong> Hotan;<br />
37°01'11"N 80°06'36"E]; syntypes: ZISP 7287 [1],<br />
7298 [4, in part], 7346 [2], 7348 [3], 7361 [3] [ZISP<br />
7298, 7361 listed under both N. y. brevibarbus and N. y.<br />
longibarbus]; compound adjective, -us, -a, -um)<br />
10.10 Heminoemacheilus Zhu & Cao, 1987<br />
Heminoemacheilus Zhu & Cao, 1987: 324 (type species:<br />
Heminoemacheilus zhengbaoshani Zhu & Cao, 1987:<br />
324, by original designation). Gender masculine.<br />
Taxonomic notes. See comments under Eonemachilus and<br />
Yunnanilus.<br />
10.10.1 Heminoemacheilus hyalinus Lan, Yang & Chen,<br />
1996<br />
Heminoemacheilus hyalinus Lan, Yang & Chen, 1996: 109,<br />
fig. 1 (type locality: China: Guangxi: Du'an County: underground<br />
river at Baoan town [24°07'N 107°51'E; Romero<br />
et al., 2009: 251]; holotype: KIZ 9409005; adjective,<br />
-us, -a, -um)<br />
10.10.2 Heminoemacheilus zhengbaoshani Zhu & Cao,<br />
1987<br />
Heminoemacheilus zhengbaoshani Zhu & Cao, 1987: 324,<br />
figs. 1–4 (type locality: China: Guangxi: Du'an County:<br />
outlet <strong>of</strong> subterranean waters near Du'an [23°56'N<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
83<br />
108°05'E; Romero et al., 2009: 252]; holotype: ASIZB<br />
741802 [61075, Eschmeyer & Fricke, 2010]; noun in<br />
genitive, indeclinable)<br />
10.11 Homatula Nichols, 1925<br />
Homatula Nichols, 1925b: 2 (subgenus <strong>of</strong> Barbatula Linck,<br />
1790: 38; type species: Nemacheilus potanini Günther,<br />
1896: 218, by original designation). Gender feminine.<br />
Taxonomic notes. Usually identified as Paracobitis by Chinese<br />
authors. Many species referred to the genus, especially<br />
cave species, do not seem to belong to this genus.<br />
10.11.1 Homatula acuticephala (Zhou & He, 1993)<br />
Paracobitis acuticephala Zhou & He, 1993: 5, fig. 1 (type<br />
locality: China: Yunnan: Eryuan County: Haixihai Lake<br />
near Niujie; holotype: YU 784141; compound adjective,<br />
-us, -a, -um)<br />
10.11.2 Homatula anguillioides (Zhu & Wang, 1985)<br />
Paracobitis anguillioides Zhu & Wang, 1985: 210, figs. 1–<br />
4 (type locality: China: Yunnan: Eryuan County: Longtang<br />
Spring, near Yousuo (26°N 99°50'E) [Mekong drainage];<br />
holotype: NGI 820134 [now IHB 820134], Hu &<br />
Zhang, 2010: 55, fig. 4a; adjective, indeclinable)<br />
10.11.3 Homatula berezowskii (Günther, 1896)<br />
Nemacheilus berezowskii Günther, 1896: 217, pl. 2 fig. c<br />
(type locality: China: Gansu: Hui-hsien [Hui Xian, about<br />
34°N 106°E, Yangtze drainage]; holotype: ZISP 10990,<br />
Bănărescu & Nalbant, 1974: 96; noun in genitive, indeclinable)<br />
10.11.4 Homatula erhaiensis (Zhu & Cao, 1988)<br />
Paracobitis erhaiensis Zhu & Cao, 1988: 95, figs. 1–2 (type<br />
locality: China: Yunnan: Wase, eastern bank <strong>of</strong> Erhai<br />
Lake; holotype: IHB 64VI0012; adjective, -is, -is, -e)<br />
10.11.5 Homatula laxiclathra Gu & Zhang, 2012<br />
Homatula laxiclathra Gu & Zhang, 2012: 592, fig. 1a (type<br />
locality: China: Shaanxi: Wei River, a tributary <strong>of</strong> Huang<br />
He, at Zhouzhi County; holotype: IHB 73V10738; adjective,<br />
-us, -a, -um [said to be used as an adjective, base<br />
on Latin words laxus and clathrus [barred]; clatrus is a<br />
noun and the adjective should have been clatratus)<br />
Taxonomic notes. See under H. variegata, a possible senior<br />
synonym.<br />
10.11.6 Homatula longidorsalis (Yang, Chen & Kottelat,<br />
1995)<br />
Paracobitis variegatus longidorsalis Yang, Chen & Kottelat,<br />
1995: 63, fig. 3 (type locality: China: Yunnan: Yiliang<br />
County: Jiuxiang, Pearl River drainage, 25°06'N<br />
103°24'E; holotype: KIZ 874048; compound noun, indeclinable)<br />
10.11.7 Homatula nanpanjiangensis (Min, Chen & Yang,<br />
2010)<br />
Paracobitis nanpanjiangensis Min, Chen & Yang, 2010: 200,<br />
fig. 1 (type locality: China: Yunnan: Qujing City: Luo-
10.10.2 Heminoemacheilus zhengbaoshani, ASIZB 102318, 53.4<br />
mm SL; China: Guangxi: Du’an (retouched).<br />
10.9.4 Hedinichthys yarkandensis, ZISP 7298/4, 207 mm SL;<br />
China: Xinjiang: Niya Oasis; syntype <strong>of</strong> H. y. longibarbus. (From<br />
Herzenstein, 1888: pl. 2 fig. 1).<br />
ping County: Niujie River (24°57'46.9"N 104°13'<br />
09.2"E), a tributary <strong>of</strong> Nanpanjiang River, at Niujie village;<br />
holotype: KIZ 1994000023; adjective, -is,<br />
-is, -e)<br />
10.11.8 Homatula oligolepis (Cao & Zhu, in Zheng, 1989)<br />
Paracobitis variegatus oligolepis Cao & Zhu, in Zheng,<br />
1989: 48, fig. 27 (type locality: China: Yunnan: Yangzong-hai<br />
Lake; holotype: IHB 63V553; compound noun,<br />
indeclinable)<br />
10.11.9 Homatula potanini (Günther, 1896)<br />
Nemacheilus potanini Günther, 1896: 218 (type locality:<br />
China: Ya River; syntypes: ZISP 10005 [3]; noun in genitive,<br />
indeclinable)<br />
Nomenclatural notes. Bănărescu & Nalbant (1974) mentioned<br />
specimen ZISP 10005, 95.0 mm, as holotype. Günther<br />
(1896) had three specimens but did not explicitly designate<br />
one as type, therefore all are syntypes. The fact that he only<br />
gave measurements <strong>of</strong> a single specimen does not make it<br />
the holotype.<br />
10.11.10 Homatula pycnolepis Hu & Zhang, 2010<br />
Homatula pycnolepis Hu & Zhang, 2010: 52, fig. 1 (type<br />
locality: China: Yunnan: Jianchuan County: Yangbi River,<br />
a tributary <strong>of</strong> upper Mekong, at Shaxi Town; about<br />
26°19'N 99°51'E; holotype: IHB 814045; compound<br />
noun, indeclinable)<br />
10.11.11 Homatula variegata (Dabry de Thiersant, in<br />
Sauvage & Dabry de Thiersant, 1874)<br />
Nemacheilus variegatus Dabry de Thiersant, in Sauvage &<br />
Dabry de Thiersant, 1874: 14 (type locality: China:<br />
Schensi [Shaanxi] [apparently Yenkiatsoun, which is<br />
Baoji Prefecture, Mei County [34°17'N 107°45'E]; in<br />
Tsang-Yu torrent, Wei He system, Huang He drainage;<br />
see below]; lectotype: MNHN 7854, designated by Bănărescu<br />
& Nalbant, 1974: 96 [as MNHN 7934]; adjective,<br />
-us, -a, -um)<br />
Nemachilus oxygnathus Regan, 1908: 357 (type locality:<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
84<br />
10.11.11 Homatula variegata, CMK 13086, 124.0 mm SL; China:<br />
Sichuan: Yangtze drainage: Yalong.<br />
10.11.11 Homatula variegata, MNHN 7854, 96.8 mm SL, lectotype;<br />
China: Shaanxi: Huang He drainage.<br />
China: Yunnan: Yunnan Fu [Kunming]; lectotype: BMNH<br />
1908.2.27.23, designated by Bănărescu & Nalbant, 1974:<br />
96; compound adjective, -us, -a, -um)<br />
Taxonomic notes. The type locality <strong>of</strong> Nemacheilus variegatus<br />
indicated by Sauvage & Dabry de Thiersant (1874) is<br />
"China"; the jar containing the lectotype bears a label indicating<br />
"Schensi" and it had been accepted that the type locality<br />
was in the Yangtze drainage (e.g. Hu & Zhang, 2010). On<br />
this basis, Hu & Zhang (2010: 58) commented that the Han<br />
River (in Yangtze drainage) and the Wei River (in Huang He<br />
drainage) are inhabited by different species and that the one<br />
in the Wei River is unnamed. Gu & Zhang (2012) described<br />
the species from the Wei River as H. laxiclathra.<br />
Although not mentioned in the original description, the<br />
MNHN database indicates the type locality as Yen-kia-tsoun.<br />
The specimens were collected by David and his travel report<br />
(David, 1875: 261, map) shows that Yen-kia-tsoun, a<br />
place now in Baoji Prefecture, is in the Wei drainage. See<br />
Triplophysa bleekeri for discussion <strong>of</strong> this locality.<br />
This suggests that H. variegata could be the valid name<br />
<strong>of</strong> the species from the Huang He and H. laxiclathra would<br />
then be a junior synonym. A comparison <strong>of</strong> a photograph <strong>of</strong><br />
the lectotype <strong>of</strong> H. variegata, the morphometric data I took<br />
in 1985 (in Yang, Chen & Kottelat, 1994: 636) and the diagnostic<br />
characters listed by Gu & Zhang (2012) indicates that<br />
the head length (15.7% SL) fits in the range <strong>of</strong> H. laxiclathra<br />
(15.9–21.1) rather than that <strong>of</strong> the H. 'variegata' (18.3–<br />
22.7), as does the depth <strong>of</strong> the caudal peduncle (6.9% SL in<br />
lectotype, 8.4–11.4 in H. laxiclathra and 9.6–13.6 in H. 'variegata')<br />
and the dorsal adipose crest extends forward until<br />
about above the anal-fin base (vs. midlength <strong>of</strong> anal-fin base<br />
in H. laxiclathra and in front <strong>of</strong> anal-fin origin in H. 'variegata').<br />
The colour pattern is almost indistinct and when I<br />
examined the lectotype (around 1985) I noted that the bars<br />
were narrower than the interspaces in the posterior part <strong>of</strong><br />
the body, which agrees with H. 'variegata' and not with<br />
H. laxiclathra. I do not have information on the remaining<br />
diagnostic characters (squamation in pre-dorsal area and<br />
shape <strong>of</strong> intestine). These apparent contradictions and the<br />
possible ambiguity about the type locality lead me to retain<br />
the name H. variegata for the Yangtze species until the lectotype<br />
is re-examined. If it would turn out that the lectotype<br />
belongs to the Huang He species, then the name H. oxygnathus<br />
is apparently available for the Yangtze species.<br />
Nomenclatural notes. For N. oxygnathus, Bănărescu &
10.12.1 Ilamnemacheilus longipinnis, NMC 79-966, 36.0 mm SL,<br />
holotype; Iran: Meymeh River, a former tributary <strong>of</strong> Tigris-Euphrates<br />
drainage. (Photograph by Zdenek Lajbner).<br />
10.13.1 Indoreonectes evezardi, about 38 mm SL; India: Maharashtra:<br />
Bhima River near Pune. (Photograph by Beta Mahatvaraj).<br />
Nalbant (1974) mentioned "syntypes BMNH 1908.2.27–24,<br />
[...], 2 spec., 114.0 and 89.5 mm, the first named here declared<br />
lectotype". The catalogue number they give is obviously<br />
incorrect. The 114.0 mm SL specimen they designated<br />
as lectotype actually bears number BMNH 1908.2.27.23.<br />
10.11.12 Homatula wujiangensis (Ding & Deng, 1990)<br />
Paracobitis wujiangensis Ding & Deng, 1990: 287, figs. 1–<br />
3 (type locality: China: Sichuan: Wujiang River; holotype:<br />
SPNRI 64274; adjective, -is, -is, -e)<br />
10.12 Ilamnemacheilus Coad & Nalbant, 2005<br />
Ilamnemacheilus Coad & Nalbant, 2005: 303 (type species:<br />
Ilamnemacheilus longipinnis Coad & Nalbant, 2005: 304,<br />
by original designation). Gender masculine.<br />
10.12.1 Ilamnemacheilus longipinnis Coad & Nalbant,<br />
2005<br />
Ilamnemacheilus longipinnis Coad & Nalbant, 2005: 304,<br />
fig. 1 (type locality: Iran: Meymeh River, a former tributary<br />
<strong>of</strong> Tigris-Euphrates River, 17 km west <strong>of</strong> Dehloran<br />
city and about 21 km east <strong>of</strong> Iraqi border, 32°45'30"N<br />
47°05'30"E; holotype: NMC 79-966; compound noun,<br />
indeclinable)<br />
10.13 Indoreonectes Rita & Banarescu, in Rita, Banarescu<br />
& Nalbant, 1978<br />
Indoreonectes Rita & Bănărescu, in Rita, Bănărescu & Nalbant,<br />
1978: 185 (subgenus <strong>of</strong> Oreonectes Günther, 1868:<br />
369; type species: Oreonectes keralensis Rita & Nalbant,<br />
in Rita, Bănărescu & Nalbant, 1978: 186, by original<br />
designation). Gender masculine.<br />
10.13.1 Indoreonectes evezardi (Day, 1872)<br />
Nemacheilus Evezardi Day, 1872: 182 (type locality: India:<br />
Maharasthra: Puna [Poona, Pune, 18°28'N 73°48'E];<br />
holotype: ZSI 2678 [lost], Whitehead & Talwar, 1976:<br />
157; noun in genitive, indeclinable)<br />
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85<br />
10.14.4 Indotriplophysa leptosoma, ZISP 7261, 144 mm SL; China:<br />
Qinghai: near Xugui. (From Herzenstein, 1888: pl. 1 fig. 2).<br />
10.14.4 Indotriplophysa leptosoma, ZISP 7261, 144 mm SL. (From<br />
Herzenstein, 1888: pl. 1 fig. 2).<br />
10.13.2 Indoreonectes keralensis (Rita & Nalbant, in Rita,<br />
Banarescu & Nalbant, 1978)<br />
Oreonectes keralensis Rita & Nalbant, in Rita, Bănărescu &<br />
Nalbant, 1978: 186, fig. 1 (type locality: India: Kerala:<br />
Pampadanpara, in a small rivulet tributary to Periyar River;<br />
holotype: ISBB 2925; adjective, -is, -is, -e)<br />
10.14 Indotriplophysa Prok<strong>of</strong>iev, 2010<br />
Indotriplophysa Prok<strong>of</strong>iev, 2010: 896 (subgenus <strong>of</strong> Triplophysa<br />
Rendahl, 1933: 21; type species: Nemacheilus<br />
tenuis Day, 1877b: 796, by original designation). Gender<br />
feminine.<br />
Nomen nudum<br />
Triplophysa kokshaalensis Prok<strong>of</strong>iev, 2010: 896<br />
10.14.1 Indotriplophysa choprai (Hora, 1934)<br />
Nemachilus choprai Hora, 1934: 310, fig. 9 (type locality:<br />
Pakistan: Chitral: Lutkuh River near hot springs; holotype:<br />
ZSI: 11301/1; noun in genitive, indeclinable)<br />
10.14.2 Indotriplophysa crassicauda (Herzenstein, 1888)<br />
Nemachilus Stoliczkae crassicauda Herzenstein, 1888: 23,<br />
pl. 7 fig. 3 (type locality: China: Qinghai: Zaidam [Quaidam<br />
basin]: Bajan-gol River [37°18'N 96°50'E]; holotype:<br />
ZISP 7221, Prok<strong>of</strong>iev, 2007b: 16, fig. 7a; compound<br />
noun, indeclinable)<br />
10.14.3 Indotriplophysa eugeniae (Prok<strong>of</strong>iev, 2002)<br />
Triplophysa eugeniae Prok<strong>of</strong>iev, 2002b: 473, fig. 5 [p. 431<br />
<strong>of</strong> translation] (type locality: China: Kam Valley, Ha-Chu<br />
River [error for Hi-Chu; upstream <strong>of</strong> Kap-chzha-kamba<br />
(Kozlov, 1902b: xxviii); Hi-Chu enters Yangtze from<br />
North at about 33°32'58"N 96°41'08"E]; holotype: ZISP<br />
12575; noun in genitive, indeclinable)
Taxonomic notes. Kam or Kami <strong>of</strong> Russian authors is Kham,<br />
a former eastern province <strong>of</strong> Tibet, corresponding to today's<br />
eastern Xizang, western Sichuan and northwestern Yunnan<br />
and including headwaters <strong>of</strong> Yangtze, Mekong, Salween, Irrawaddy,<br />
and Brahmaputra. Kozlov was in upper Mekong<br />
and upper Yangtze in 1900. The type locality is Hi-Chu [name<br />
in ZISP accession book, not Ha-Chu]. The Hi-Chu is a northern<br />
tributary <strong>of</strong> Tongtiang He (the name <strong>of</strong> a stretch <strong>of</strong> the<br />
upper Yangtze). Kozlov travelled from 13 to 20 July 1900<br />
along the Hi-Chu from Kap-chzha-kamba to the source (Kozlov,<br />
1902b: xxviii). See also maps in Kozlov (1902a, 1948).<br />
10.14.4 Indotriplophysa leptosoma (Herzenstein, 1888)<br />
Nemachilus Stoliczkae leptosoma Herzenstein, 1888: 23,<br />
pl. 1 fig. 2 (type locality: China: Tibet: Schuga River<br />
[based on map in Przheval'skii, 1883: Qinghai, river flowing<br />
near Xugui and entering Golmud He at 35°57'N<br />
94°48'E]; holotype: ZISP 7261, Prok<strong>of</strong>iev, 2007b: 15,<br />
fig. 6a; compound noun, indeclinable)<br />
Nemachilus Stoliczkae productus Herzenstein, 1888: 23, pl. 1<br />
fig. 5 (type locality: China: Qinghai: "Fontes fl. Chuanche"<br />
[sources <strong>of</strong> Chuan-che River, Huang He; then considered<br />
to be lakes Zhaling Hu [or Ngoring Hu, 34°52'N<br />
97°10'E] and Eling Hu [or Gyaring Hu, 34°52'N 97°38'E];<br />
holotype: ZISP 7329, Prok<strong>of</strong>iev, 2007b: 15, fig. 6b; simultaneous<br />
subjective synonym <strong>of</strong> Nemachilus stoliczkae<br />
leptosoma Herzenstein, 1888: 23, first reviser [apparently<br />
Zhu, 1989: 110] gave precedence to N. s. leptosoma;<br />
adjective, -us, -a, -um)<br />
10.14.5 Indotriplophysa tenuicauda (Steindachner, 1866)<br />
Cobitis tenuicauda Steindachner, 1866: 792, pl. 17 fig. 3<br />
(type locality: India: Jammu & Kashmir: Rupshu: rivulet<br />
near Hanle monastery; syntypes: NMW 48570 [6], Eschmeyer<br />
& Fricke, 2010; compound noun, indeclinable)<br />
? Nemachilus ladacensis Günther, 1868: 356 (type locality:<br />
India: Kashmir: Ladakh; holotype: BMNH 1860.3.19.<br />
789, Kullander et al., 1999: 142, 145; adjective, -is, -is, -e)<br />
Taxonomic notes. Kullander et al. (1999: 145) examined<br />
the holotype <strong>of</strong> N. ladacensis and commented that it might<br />
be a synonym <strong>of</strong> I. tenuicauda; both nominal species are<br />
described from Ladakh. Li et al. (2007: 55) considered<br />
N. ladacensis as a valid species present in the Tarim basin;<br />
they do not mention I. tenuicauda and do not seem to have<br />
had access to material.<br />
10.14.6 Indotriplophysa tenuis (Day, 1877)<br />
Nemacheilus tenuis Day, 1877b: 796 (type locality: Tadjikistan:<br />
Aktash [12600 ft] where water <strong>of</strong> Ak-su passes to<br />
Oxus [Amu-Darya drainage, Murgab system, 3840 masl;<br />
37°35'13"N 74°53'44"E; Trotter, 1875: (28), (65)] / China:<br />
Xinjiang: Yankihissar (4320 ft) [Yengisar, 38°55'33"N<br />
76°10'17"E] where rivers go to Yarkand River; syntypes:<br />
ZSI 684–686, 710–711, 709 [lost], BMNH 1889.2.1.<br />
1716–1717, NMW 48447, Whitehead & Talwar, 1976:<br />
157; also in Day, 1878b: 15, pl. 5 fig. 4; adjective, -is,<br />
-is, -e)<br />
Taxonomic notes. The widely disjunct localities suggest that<br />
two species are involved. A lectotype designation and redescription<br />
are needed.<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
86<br />
10.14.7 Indotriplophysa yasinensis (Alcock, 1898)<br />
Nemachilus yasinensis Alcock, 1898: 38, pl. 2 figs. 2, 2a<br />
(type locality: Pakistan: Gilgit-Baltistan: Yasin River at<br />
8500 feet [Yasin: 36°22'N 73°20'E]; holotype (?): ZSI<br />
F 14181, according to Menon & Yazdani, 1968: 125; adjective,<br />
-is, -is, -e)<br />
Nemachilus kashmirensis Hora, 1922: 76 (type locality: India:<br />
Jammu & Kashmir: mouth <strong>of</strong> trout farm at Harwan<br />
[34°09'30"N 74°54'16"E] / Verinag spring [33°32'17"N<br />
75°14'49"E] / Kukarnag spring [33°33'48"N<br />
75°20'11"E]; syntypes [17+]: ZSI 10121/1 [9], 10122/1<br />
[9], Menon & Yazdani, 1968: 123; adjective, -is, -is, -e)<br />
? Nemachilus ajmonis Di Caporiacco, 1933: 342, fig. (type<br />
locality: India: Kashmir: pond between Sonamarg<br />
[34°18'04"N 75°17'30"E] and Baltal [34°15'21"N<br />
75°25'05"E]; syntypes [2]: MZUF 55881 [1], Vanni,<br />
1991: 221; adjective, -is, -is, -e)<br />
? Nemachilus hutchinsoni Hora, 1936: 314, figs. 6–7, pl. 12<br />
figs. 1–2 (type locality: India: Kashmir: Ladakh: 3 miles<br />
west <strong>of</strong> Mugleb / pond between Durbuk and Tangtse /<br />
Tsar Tso / Lukung; syntypes [6]: ZSI F 12201/1 [1],<br />
12202/1 [3], 12203/1 [1], Menon & Yazdani, 1968: 122;<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. Synonymy follows Prok<strong>of</strong>iev (2007b: 3,<br />
2010: 896), itself partly based on Menon (1987: 217).<br />
10.15 Iskandaria Prok<strong>of</strong>iev, 2009<br />
Iskandaria Prok<strong>of</strong>iev, 2009b: 889 (type species: Nemachilus<br />
kuschakewitschi Herzenstein, 1890: 61, by original<br />
designation). Gender feminine.<br />
10.15.1 Iskandaria kuschakewitschi (Herzenstein, 1890)<br />
Nemachilus Kuschakewitschi Herzenstein, 1890: 61 [p. 139<br />
<strong>of</strong> reprint] (type locality: Uzbekistan: Syr-Darya drainage:<br />
Margelan [40°28'16"N 71°43'29"E] and Andizhan<br />
[40°47'N 72°202 E]; syntypes: ZISP 4561 [6+], 8744<br />
[6+], 8745 [6+], ? BMNH 1891.10.7.72 [2], Eschmeyer<br />
& Fricke, 2010; noun in genitive, indeclinable)<br />
Nemachilus kuschakewitschi badamensis Turdakov, 1947a:<br />
56 (type locality: Kazakhstan: Yuzhno-Kazakhstansjkaya<br />
oblast: Syr-Darya drainage: Arys' basin: Badam River at<br />
Shymkent [42°19'00"N 69°35'45"E]; syntypes [56]:<br />
ZMMU P-3709 [14], P-3710 [1], Vasilieva et al., in Pavlinov<br />
& Borissenko, 2001: 26; adjective, -is, -is, -e)<br />
10.15.2 Iskandaria pardalis (Turdakov, 1941)<br />
Nemachilus pardalis Turdakov, 1941: 219, fig. 1 (type locality:<br />
Tajikistan: Amu-Darya drainage: Kafirnigan River<br />
system [K<strong>of</strong>arnihon], Dyushambinka River from Stalinabad<br />
[Dushanbe; 38°32'12"N 68°46'48"E] to 4–5 km<br />
upstream; holotype: ZMMU 3791; noun in apposition,<br />
indeclinable)<br />
Taxonomic notes. Treated as subspecies <strong>of</strong> I. kuschakewitschi<br />
by Prok<strong>of</strong>iev (2009b: 889).<br />
10.16 Labiatophysa Prok<strong>of</strong>iev, 2010<br />
Labiatophysa Prok<strong>of</strong>iev, 2010: 895 (subgenus <strong>of</strong> Triplophysa<br />
Rendahl, 1933: 21; type species: Diplophysa labiata
Kessler, 1874: 59, by original designation). Gender feminine.<br />
Taxonomic notes. See Prok<strong>of</strong>iev (2004a) for revision.<br />
Species incertae sedis<br />
10.16.1 Labiatophysa nasalis (Kessler, 1876)<br />
Diplophysa nasalis Kessler, 1876: 27, pl. 3 fig. 2 (type locality:<br />
China: Nei Mongol: Lake Dalai-Nor [Dalai Nur;<br />
43°18'N 116°40'E]; holotype: ZISP 2475, Prok<strong>of</strong>iev,<br />
2004a: 203; noun in apposition, indeclinable)<br />
Taxonomic notes. Known only from holotype, which may<br />
belong to Labiatophysa or Barbatula (Prok<strong>of</strong>iev, 2004a:<br />
204). The type locality is 'Lake Dalai-Nor'. There are several<br />
lakes by this name in China and Mongolia [alternative<br />
spellings: Dalaï-Nur, Dalai-Nuur, etc.]. The holotype was<br />
collected by Przewal'skii in 1874 and his travel map<br />
(Przewal'skii, 1876) shows only one Dalai-nor, at 43°18'N<br />
116°40'E (not Hu-Lun Lake, which is another Dalai Nur;<br />
contra Eschmeyer & Fricke, 2010).<br />
10.16.2 Labiatophysa herzensteini (Berg, 1909)<br />
Diplophysa labiata herzensteini Berg, 1909: 10 (type locality:<br />
Kazakhstan: Almaty Province: Tscharyn River<br />
[Charyn], tributary <strong>of</strong> Ili River, Balkash basin, area <strong>of</strong><br />
Semiretschenks; lectotype: ZISP 14530, designated by<br />
Prok<strong>of</strong>iev, 2004a: 191, fig. 26; noun in genitive, indeclinable)<br />
10.16.3 Labiatophysa kaznakowi (Prok<strong>of</strong>iev, 2004)<br />
Triplophysa kaznakowi Prok<strong>of</strong>iev, 2004a: 195, fig. 54 (type<br />
locality: China: Xinjiang-Uighur: Tuguryuk River Valley<br />
(= Nom Gol River flowing from extreme eastern escarpes<br />
<strong>of</strong> Tien-Shan Mountains and disappearing in the<br />
sands <strong>of</strong> Nomin-Gobi desert, ca. 42°N, ca. 96°E); holotype:<br />
ZISP 12485; noun in genitive, indeclinable)<br />
10.16.4 Labiatophysa labiata (Kessler, 1874)<br />
Diplophysa labiata Kessler, 1874: 59, pl. 8 fig. 41 (type locality:<br />
Kyrgyzstan: Urdjar River, tributary <strong>of</strong> Lake Ala-kul<br />
[42°19'03"N 78°32'08"E; in Issyk Kul endorheic basin];<br />
syntypes: ZISP 2321 [4]; adjective, -us, -a, -um)<br />
Nemachilus labiatus conjungens Herzenstein, 1888: 84, pl. 4<br />
fig. 5 (type locality: Kazakhstan and China: Chorgos River<br />
[Khorgas; a tributary <strong>of</strong> Ili River, confluence at<br />
43°49'25"N 80°31'22"E, making the border between<br />
Kazakhstan and China]; syntypes: ZISP 7843 [1], 7850<br />
[2], Eschmeyer & Fricke, 2010; participle, indeclinable)<br />
Taxonomic notes. Synonymy follows Prok<strong>of</strong>iev (2004a).<br />
10.16.5 Labiatophysa microphthalma (Kessler, 1879)<br />
Diplophysa microphthalma Kessler, 1879: 308 (type locality:<br />
China: Xinjiang: "the stream on which Chami town<br />
[Hami or Kumul; 42°48'00"N 93°27'00"E ] is located [...]<br />
southern slope <strong>of</strong> Altai Range"; lectotype: ZISP 4208,<br />
designated by Prok<strong>of</strong>iev, 2004a: 194, fig. 33; reprinted<br />
as Kessler, 1880: 269; compound adjective, -us, -a, -um)<br />
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10.17 Lefua Herzenstein, 1888<br />
Octonema Herzenstein, in Warpachowski & Herzenstein,<br />
1887: 47 (type species: Octonema pleskei Warpachowski<br />
& Herzenstein, 1887: 48, by subsequent designation<br />
by Berg, 1907a: 437; junior homonym <strong>of</strong> Octonema<br />
Martens, 1869: 608 in Pisces and Octonema Haeckel,<br />
1879: 126 in Coelenterata). Gender neuter.<br />
Lefua Herzenstein, 1888: 3 (replacement name for Octonema<br />
Herzenstein, in Warpachowski & Herzenstein, 1887:<br />
47). Gender feminine.<br />
Elxis Jordan & Fowler, 1903: 768 (type species: Elxis nikkonis<br />
Jordan & Fowler, 1903: 768, by original designation).<br />
Gender feminine [a classical Greek word].<br />
Nomenclatural notes. The type species <strong>of</strong> Octonema is listed<br />
as O. pleskei by monotypy by Eschmeyer & Fricke (2010).<br />
Warpachowski & Herzenstein (1887: 47) listed two included<br />
species, O. pleskei and O. costatum; Berg (1907a: 437)<br />
designated O. pleskei as type species.<br />
10.17.1 Lefua costata (Kessler, 1876)<br />
Diplophysa costata Kessler, 1876: 29, pl. 3 fig. 3 [not 4]<br />
(type locality: "Mongolia: Lake Dalai-Nor at 43°N"<br />
[Berg, 1949: 887; China: Nei Mongol: Dalai Nur;<br />
43°18'N 116°40'E; not Hu-Lun Lake, which is another<br />
Dalai Nur]; holotype: ZISP 2477, Naseka & Bogutskaya,<br />
2004: 285, pl. 2 fig. 6; adjective, -us, -a, -um)<br />
Nemacheilus dixoni Fowler, 1899: 181 (type locality: China:<br />
Pechili: Tan Lan Ho, tributary <strong>of</strong> Shu Lan Ho, about<br />
30 miles northeast <strong>of</strong> Lama-Miau or Dolon-Nor [Doloon-<br />
Nuur, 42°11'22"N 116°28'24"E] [now in Nei Mongol:<br />
Duolun County; Pechili [Zhili], name <strong>of</strong> a former province];<br />
holotype: ANSP 16393; noun in genitive, indeclinable)<br />
Elxis coreanus Jordan & Starks, 1905: 201, fig. 7 (type locality:<br />
Korea: Gensan; holotype: USNM 45243; adjective,<br />
-us, -a, -um)<br />
Lefua andrewsi Fowler, 1922: 1 (type locality: China: Shing<br />
Lung Shan, Eastern Tombs [now Hebei: Zunhua, 40°11'N<br />
117°58'E]; holotype: AMNH 7974; noun in genitive, indeclinable)<br />
10.17.2 Lefua echigonia Jordan & Richardson, 1907<br />
Lefua echigonia Jordan & Richardson, 1907: 263, fig. 1 (type<br />
locality: Japan: Echigo: Nagaoka; holotype: USNM<br />
45243 [in original publication; CAS-SU 20164 according<br />
to Böhlke, 1953: 39]; adjective, -us, -a, -um)<br />
10.17.3 Lefua h<strong>of</strong>fmanni Herre, 1932<br />
Lefua h<strong>of</strong>fmanni Herre, 1932: 429 (type locality: China:<br />
Kwangtung [Guangdong]; holotype: CAS-SU 25727,<br />
Böhlke, 1953: 40, Prok<strong>of</strong>iev, 2005: 431, 437; noun in<br />
genitive, indeclinable)<br />
Taxonomic notes. Prok<strong>of</strong>iev (2005: 437) commented that<br />
this species is neither an Oreonectes nor a Lefua but could<br />
not decide its generic position.<br />
10.17.4 Lefua nikkonis (Jordan & Fowler, 1903)<br />
Elxis nikkonis Jordan & Fowler, 1903: 768, fig. 1 (type locality:<br />
Japan: Hokkaido: Chitose, in Iburi; holotype: CAS-<br />
SU 7848; adjective, -is, -is, -e)
10.16.4 Labiatophysa labiata, ZISP 7850, 126 mm SL; syntype<br />
<strong>of</strong> L. l. conjungens; border between Kazakhstan and China: Xinjiang:<br />
Ili drainage. (From Herzenstein, 1888: pl. 4 fig. 5).<br />
10.16.4 Labiatophysa labiata, ASIZB 53485, 144.0 mm SL.<br />
10.15.1 Iskandaria kuschakewitschi, ZMMU P-9298, about 60 mm<br />
SL; Tajikistan: Amu-Darya drainage: Kara-Su River. (Photograph<br />
by Artem Prok<strong>of</strong>iev).<br />
10.17.5 Lefua pleskei (Herzenstein, in Warpachowski &<br />
Herzenstein, 1887)<br />
Octonema Pleskei Herzenstein, in Warpachowski & Herzenstein,<br />
1887: 48, pl. fig. 5 (type locality: Russia: Primorsky<br />
Kray: Lefu River [Ilistaya; a tributary <strong>of</strong> Lake Khanka]<br />
near Nikolajewka; syntypes: ZISP 7209 [4], BMNH<br />
1891.10.7.73 [1], Naseka & Bogutskaya, 2004: 285, pl. 2<br />
fig. 7; noun in genitive, indeclinable)<br />
10.17.6 Lefua sayu (Herre & Lin, 1936)<br />
Oreonectes sayu Herre & Lin, 1936: 19 (type locality: China:<br />
Chekiang: upper waters <strong>of</strong> the Tsien Tang; holotype:<br />
CAS-SU 32584, Böhlke, 1953: 40; noun in apposition,<br />
indeclinable)<br />
Taxonomic notes. Generic position follows Prok<strong>of</strong>iev (2005:<br />
437).<br />
10.18 Mesonoemacheilus Banarescu & Nalbant, in Singh,<br />
Sen, Banarescu & Nalbant, 1982<br />
Mesonoemacheilus Bănărescu & Nalbant, in Singh, Sen,<br />
Bănărescu & Nalbant, 1982: 202 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
88<br />
10.17.1 Lefua costata, ASIZB 132268, 48.7 mm SL; China: Jilin:<br />
Antu.<br />
10.17.1 Lefua costata (?), CMK 21951, 56.4 mm SL; Korea:<br />
Gangwon.<br />
10.18.5 Mesonoemacheilus petrubanarescui, CMK 5426, 46.7 mm<br />
SL; India: Karnataka: Chandragiri River.<br />
triangularis Day, 1865a: 295, by original designation).<br />
Gender masculine.<br />
10.18.1 Mesonoemacheilus guentheri (Day, 1867)<br />
Nemacheilus guentheri Day, 1867a: 285 (type locality: India:<br />
rapids on slopes <strong>of</strong> Neilgherries [p. 282]; syntypes:<br />
? ZSI 2578 [1], A.959 [1, lost], BMNH 1867.7.24.28 [1],<br />
AMS B.7488 [1], NMW 48412, RMNH 2666, Whitehead<br />
& Talwar, 1976: 157; noun in genitive, indeclinable)<br />
10.18.2 Mesonoemacheilus herrei Nalbant & Banarescu,<br />
in Singh, Sen, Banarescu & Nalbant, 1982<br />
Mesonoemacheilus herrei Nalbant & Bănărescu, in Singh,<br />
Sen, Bănărescu & Nalbant, 1982: 203, figs. 1–6 (type<br />
locality: India: Kerala: Anamallai Hills, Valparai, Puthutotam<br />
Estate; holotype: CAS-SU 41307; noun in genitive,<br />
indeclinable)<br />
10.18.3 ? Mesonoemacheilus menoni (Zacharias & Minimol,<br />
1999)<br />
Noemacheilus menoni Zacharias & Minimol, 1999: 288,<br />
fig. 1 (type locality: India: Kerala: Periyar River, Malappara;<br />
holotype: ZSI/WGRS V/F/1018; noun in genitive,<br />
indeclinable)<br />
10.18.4 Mesonoemacheilus pambarensis (Rema Devi &<br />
Indra, 1994)<br />
Noemacheilus pambarensis Rema Devi & Indra, 1994: 207,<br />
pl. 1 fig. 1 (type locality: India: Kerala: Pambar River at<br />
border <strong>of</strong> Chinnar Sanctuary; holotype: ZSI/SRS F 4095;<br />
adjective, -is, -is, -e)
10.19.1 Metaschistura cristata, FSJF 3277, 66.0 mm SL; Iran: Zanglangou<br />
River near Kazgankala. (Photograph by Jörg Freyh<strong>of</strong>).<br />
10.20.1 Micronemacheilus cruciatus, CMK 15120, 25.7 mm SL;<br />
Vietnam: Hue.<br />
10.21.4 Nemacheilus binotatus, CMK 16044, 34.6 mm SL; Thailand:<br />
Chao Phraya drainage: Nan.<br />
10.18.5 Mesonoemacheilus petrubanarescui (Menon,<br />
1984)<br />
Noemacheilus petrubanarescui Menon, 1984: 46, figs. 1–2<br />
(type locality: India: Karnataka: Netravati River at Dharmasthala;<br />
holotype: ZSI F 559; noun in genitive, indeclinable)<br />
10.18.6 Mesonoemacheilus pulchellus (Day, 1873)<br />
Nemacheilus pulchellus Day, 1873: 528 (type locality: India:<br />
Neilghiries: Bowany River [Bhavani]; syntypes [23]:<br />
ZSI 2581 [1], 2563 [1], BMNH 1889.2.1.1591–1600<br />
[10], AMS B.7739 [1], NMW 48427 [1], RMNH 2664<br />
[1], ZISP 8263 [2], Whitehead & Talwar, 1976: 157; adjective,<br />
-us, -a, -um)<br />
10.18.7 Mesonoemacheilus remadevii Shaji, 2002<br />
Mesonoemacheilus remadevii Shaji, 2002: 217, figs. 2–4<br />
(type locality: India: Kerala: Kunthi River at Poochipara,<br />
Silent Valley National Park; holotype: KFRI F900; noun<br />
in genitive, indeclinable)<br />
10.18.8 Mesonoemacheilus triangularis (Day, 1865)<br />
Nemacheilus triangularis Day, 1865a: 295 (type locality:<br />
India: Kerala: hills at Mundikyum; holotype: ZSI 2681<br />
[lost], Whitehead & Talwar, 1976: 157; also in Day,<br />
1865b: 203, pl. 14 fig. 1; adjective, -is, -is, -e)<br />
Nemacheilus triangularis tambaraparniensis Menon, 1987:<br />
171 (type locality: India: Tamil Nadu: Chittaruvi Forest,<br />
Courtalam (Tambaraparni drainage); holotype: ZSI/SRS<br />
F 576; adjective, -is, -is, -e)<br />
? Nemacheilus periyarensis Kurup & Radhakrishnan, 2005:<br />
75, fig. 1 (type locality: India: Kerala: Thannikkudy, Periyar<br />
Lake; holotype: ZSI/WGRS CLT V/F13030; adjective,<br />
-is, -is, -e)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
89<br />
10.21.1 Nemacheilus anguilla, CMK 14403, 47.2 mm SL; India:<br />
Bengal.<br />
10.21.1 Nemacheilus anguilla, CMK 14403, 47.2 mm SL.<br />
10.19 Metaschistura Prok<strong>of</strong>iev, 2009<br />
Metaschistura Prok<strong>of</strong>iev, 2009b: 892 (type species: Nemacheilus<br />
cristatus Berg, 1898: 18, by original designation).<br />
Gender feminine.<br />
10.19.1 Metaschistura cristata (Berg, 1898)<br />
Nemachilus cristatus Berg, 1898: 18, fig. (type locality:<br />
Turkmenistan: Tedschent River near Aschabad [Askhabad,<br />
Ashgabat; 37°56'N 58°22'E; endorheic drainage,<br />
now connected by canal to Amu-Darya]; syntypes: ZISP<br />
11055, ZMMU P.2555 [59], ISBB 996 [2], Vasilieva,<br />
pers. comm., Eschmeyer & Fricke, 2010; adjective, -us,<br />
-a, -um)<br />
10.20 Micronemacheilus Rendahl, 1944<br />
Micronemacheilus Rendahl, 1944: 45 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus<br />
cruciatus Rendahl, 1944: 37, by original designation).<br />
Gender masculine.<br />
10.20.1 Micronemacheilus cruciatus (Rendahl, 1944)<br />
Nemacheilus cruciatus Rendahl, 1944: 37, fig. 19 (type locality:<br />
Vietnam: Thua Luu, 50 km southeast <strong>of</strong> Hué; holotype:<br />
NHMG; adjective, -us, -a, -um)<br />
10.21 Nemacheilus Bleeker, 1863<br />
Naunacheilus Kuhl & van Hasselt, in van Hasselt, 1823: 132<br />
(nomen nudum)<br />
Noemacheilus Kuhl & van Hasselt, in van Hasselt, 1823:<br />
133, 1824: 376 (nomen nudum, Kottelat, 1987b: 371)<br />
Naemacheilus Valenciennes, in Cuvier & Valenciennes,<br />
1846: 26 (nomen nudum, name cited in synonymy)<br />
Nemacheilus Bleeker, 1863a: 37 (type species: Cobitis fas-
ciata Valenciennes, in Cuvier & Valenciennes, 1846: 25,<br />
by original designation; also in Bleeker, 1863c: 4,<br />
pl. 103). Gender masculine.<br />
Nemacheilos Kner, 1867: 366 (incorrect subsequent spelling)<br />
Nematocheilos Kner, 1867: 366 (incorrect subsequent spelling)<br />
Nemachilus Günther, 1868: 347 (unjustified emendation <strong>of</strong><br />
Nemacheilus Bleeker, 1863a: 37). Gender masculine.<br />
Modigliania Perugia, 1893a: 246 (type species: Modigliania<br />
papillosa Perugia, 1893a: 246, by monotypy). Gender<br />
feminine.<br />
Pogononemacheilus Fowler, 1937: 158 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus<br />
masyae Smith, 1933: 58, by original designation). Gender<br />
masculine.<br />
Nomenclatural notes. Noemacheilus Kuhl & van Hasselt<br />
is a nomen nudum. As a curiosity, it is worth mentioning that<br />
this spelling, which has been used to form numerous genusgroup<br />
names, is most likely erroneous. The name is also<br />
spelled Naunacheilus in the same work. Van Hasselt's text<br />
was not an article but a letter he sent to Temminck. Several<br />
<strong>of</strong> the names in the printed letter are misspelt or have aberrant<br />
or multiple spellings. Most likely the intended spelling<br />
was Naemacheilus and a type setter probably confused the<br />
ligatures æ (ae) and œ (oe) and it became printed oe. Nemacheilus<br />
or næmacheilus has an obvious etymology (νµα,<br />
nema, a thread; χελος, keilos, a lip; Greek), which noemacheilus<br />
does not have.<br />
Species inquirenda<br />
Lepidocephalus weberi Ahl, 1922: 32 (type locality: Indonesia:<br />
Sumatra: Benkulen; syntypes: ZMB 7670 [2,<br />
lost ?], Paepke, 1995: 91; noun in genitive, indeclinable)<br />
Taxonomic notes. Identification at species level will be possible<br />
only if the syntypes can be found, or after some nemacheilines<br />
are collected at the type locality and a neotype designated.<br />
10.21.1 'Nemacheilus' anguilla Annandale, 1919<br />
Nemachilus anguilla Annandale, 1919: 127, pl. 1 fig. 3, pl. 3<br />
fig. 1 (type locality: India: Bombay Presidency [Maharashtra]:<br />
Satara District: Yenna River at Medha; holotype:<br />
ZSI F 9692/1, Menon & Yazdani, 1968: 121; noun<br />
in apposition, indeclinable)<br />
Nemachilus poonaensis Menon, 1949: 226, fig. 1a (type<br />
locality: India: Bombay Presidency [Maharashtra]: Moola-Mootha<br />
River at Poona [Pune, 18°28'N 73°48'E]; holotype:<br />
ZSI F 242/2, Menon & Yazdani, 1968: 124; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. Generic position requires investigation.<br />
Synonymy follows Menon (1987: 154).<br />
10.21.2 Nemacheilus arenicolus Kottelat, 1998<br />
Nemacheilus arenicolus Kottelat, 1998a: 65, fig. 102 (type<br />
locality: Laos: Khammouan Province: Nam Theun basin:<br />
Nam Xot in Ban Nam Xot; 17°53'20"N 105°05'10"E;<br />
holotype: ZRC 41790; originally treated as adjective,<br />
-us, -a, -um)<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
90<br />
'inhabitant <strong>of</strong>' are nouns and do not have to agree in gender.<br />
But when proposed, arenicolus was explicitly proposed as<br />
an adjective and therefore it has to agree in gender with the<br />
genus name.<br />
10.21.3 Nemacheilus banar Freyh<strong>of</strong> & Serov, 2001<br />
Nemacheilus banar Freyh<strong>of</strong> & Serov, 2001: 135, figs. 1–2<br />
(type locality: Vietnam: Kontum Province: middle Pako<br />
River about 50 km north <strong>of</strong> Kontum; 14°39.6'N<br />
107°46.98'E; holotype: ZFMK 28588; noun in apposition,<br />
indeclinable)<br />
10.21.4 Nemacheilus binotatus Smith, 1933<br />
Nemacheilus binotatus Smith, 1933: 61, pl. 1 fig. 4 (type<br />
locality: Thailand: Chiang Mai Province: Mekhan, tributary<br />
<strong>of</strong> Meping, southwest <strong>of</strong> Doi Sutep; holotype:<br />
KUMF 167, Monkolprasit, 1969: 6; adjective, -us, -a,<br />
-um)<br />
10.21.5 Nemacheilus chrysolaimos (Valenciennes, in Cuvier<br />
& Valenciennes, 1846)<br />
Cobitis chrysolaimos Valenciennes, in Cuvier & Valenciennes,<br />
1846: 27, pl. 521 (type locality: Indonesia: Java;<br />
lectotype: MNHN 3961, designated by Kottelat, 1984b:<br />
241; compound noun, indeclinable)<br />
10.21.6 Nemacheilus cleopatra Freyh<strong>of</strong> & Serov, 2001<br />
Nemacheilus cleopatra Freyh<strong>of</strong> & Serov, 2001: 138, figs. 5–<br />
6 (type locality: Vietnam: Gia Lai Province: Sol River<br />
near Plao Ganong about 60 km south <strong>of</strong> Pleiku;<br />
13°31.82'N 108°18.63'E; holotype: ZFMK 27155; noun<br />
in apposition, indeclinable)<br />
10.21.7 'Nemacheilus' corica (Hamilton, 1822)<br />
Cobitis corica Hamilton, 1822: 359, 395 (type locality: India:<br />
Kosi River [Purnia District: Mainayi; Hora, 1935a:<br />
49]; types: NT; noun in apposition, indeclinable)<br />
Schistura punctata M'Clelland, 1839: 308, 442, pl. 53 fig. 4<br />
(type locality: India: Assam / northeastern parts <strong>of</strong> Bengal;<br />
syntypes: material examined by M'Clelland and basis<br />
<strong>of</strong> Cobitis corica Hamilton, 1822: 359; adjective,<br />
-us, -a, -um)<br />
Cobites cinerea Swainson, 1839: 310 (available by indication<br />
to Hamilton, 1822: 359 No. 12 [which is Cobitis<br />
corica]; type locality: India: Kosi River [Purnia District:<br />
Mainayi; Hora, 1935a: 49]; types: NT; adjective, -us, -a,<br />
-um)<br />
Taxonomic notes. Generic position requires investigation.<br />
10.21.8 Nemacheilus elegantissimus Chin & Samat, 1992<br />
Nemacheilus elegantissimus Chin & Samat, 1992: 26, fig. 2<br />
(type locality: Malaysia: Borneo: Sabah: Lahad Datu<br />
District: Sungai Payau, a tributary <strong>of</strong> Sungai Segama,<br />
Danum Valley Conservation Area; holotype: FMNH<br />
100749; adjective, -us, -a, -um)<br />
Nemacheilus ornatissimus Doi, 1997: 16 (nomen nudum;<br />
lapsus for Nemacheilus elegantissimus Chin & Samat,<br />
1992: 26, fig. 1)
10.21.9 Nemacheilus fasciatus (Valenciennes, in Cuvier<br />
& Valenciennes, 1846)<br />
Noemacheilus fasciatus Kuhl & van Hasselt, in van Hasselt,<br />
1823: 133, 1824: 376 (nomen nudum, Kottelat, 1987b:<br />
371)<br />
Cobitis fasciata Valenciennes, in Cuvier & Valenciennes,<br />
1846: 25 (type locality: Indonesia: Java: Buitenzorg<br />
[Bogor]; holotype: MNHN B.2798, Kottelat, 1984b: 247,<br />
fig. 18a [as lectotype], 1987a: 371, Roberts, 1993: 26;<br />
adjective, -us, -a, -um)<br />
Cobitis suborbitalis Valenciennes, in Cuvier & Valenciennes,<br />
1846: 26 (type locality: Indonesia: Java; holotype:<br />
MNHN 3930, Bertin & Estève, 1948: 96, Kottelat, 1984b:<br />
250; simultaneous subjective synonym <strong>of</strong> Cobitis fasciata<br />
Valenciennes, in Cuvier & Valenciennes, 1846: 25, first<br />
reviser [Bleeker, 1860c: 79] gave precedence to C. fasciata;<br />
compound noun, indeclinable)<br />
10.21.10 ? Nemacheilus jaklesii (Bleeker, 1852)<br />
Cobitis Jaklesii Bleeker, 1852: 604 (type locality: Indonesia:<br />
Sumatra: Pajacombo [Payakumbuh]; lectotype:<br />
RMNH 7055, designated by Alfred, 1961: 33; noun in<br />
genitive, indeclinable)<br />
10.21.11 'Nemacheilus' kaimurensis Husain & Tilak, 1998<br />
Nemacheilus kaimurensis Husain & Tilak, 1998: 131, figs.<br />
1–2 (type locality: India: Uttar Pradesh: Sombhadra District:<br />
Kanhar stream, near Kota village, Chopan; holotype:<br />
NRS/ZSI F 1543; adjective, -is, -is, -e)<br />
Taxonomic notes. Generic position requires investigation.<br />
10.21.12 Nemacheilus kapuasensis Kottelat, 1984<br />
Noemacheilus kapuasensis Kottelat, 1984b: 244, fig. 16 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: rocky<br />
channel in main stream <strong>of</strong> Sungai Pinoh at Nanga Saian,<br />
45 km south <strong>of</strong> Nangapinoh, 0°43'S 11°83'E; holotype:<br />
MZB 4004; adjective, -is, -is, -e)<br />
10.21.13 Nemacheilus longipectoralis Popta, 1905<br />
Nemachilus longipectoralis Popta, 1905: 182 (type locality:<br />
Indonesia: Borneo: Kalimantan Timur: upper Mahakam;<br />
lectotype: RMNH 7641, designated by Kottelat,<br />
1984b: 239; also in Popta, 1906: 198, pl. 10 fig. 42; compound<br />
noun, indeclinable)<br />
10.21.14 Nemacheilus longipinnis Ahl, 1922<br />
Nemachilus longipinnis Ahl, 1922: 31 (type locality: Indonesia:<br />
Central Sumatra [upper and middle sections <strong>of</strong><br />
Rokan Kanan, Rokan Kiri and Siak drainages ("Sultanate<br />
<strong>of</strong> Siak" [Kabupaten Siak, Riau Province, Sumatra]<br />
and "Rokan states"; Moszkowski, 1909a-b: maps]; holotype:<br />
ZMB 20547; potentially junior secondary homonym<br />
<strong>of</strong> Acanthocobitis longipinnis Peters, 1861: 712 if<br />
treated as valid in Nemacheilus, which has apparently<br />
never been done; compound noun, indeclinable)<br />
Nemacheilus lactogeneus Roberts, 1989: 107, fig. 82 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: Kapuas<br />
mainstream 58 km northeast <strong>of</strong> Sintang and 1 km dowstream<br />
from Sebruang; holotype: MZB 3542; adjective,<br />
-us, -a, -um)<br />
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10.21.15 Nemacheilus longistriatus Kottelat, 1990<br />
Nemacheilus longistriatus Kottelat, 1990a: 51, fig. 26 (type<br />
locality: Thailand: Loei Province: Mekong main stream<br />
between Chiang Khan (17°50'N 101°45'E) to 70 km<br />
downstream; holotype: CAS 62547; adjective, -us, -a,<br />
-um)<br />
10.21.16 Nemacheilus marang Hadiaty & Kottelat, 2010<br />
Nemacheilus marang Hadiaty & Kottelat, 2010: 41, figs. 2–<br />
3 (type locality: Indonesia: Kalimantan Timur: Kabupaten<br />
Kutai Timur: Kecamatan Kelai: Tepian Langsat village,<br />
Sungai Marang (1°42.465'N 117°45.549'E), Bengalon<br />
drainage; holotype: MZB 13301; noun in apposition,<br />
indeclinable)<br />
10.21.17 Nemacheilus masyae Smith, 1933<br />
Nemacheilus masyae Smith, 1933: 58, fig. 3, pl. 1 fig. 3<br />
(type locality: Thailand: Nakhon Sritamarat: Ban Ta Yai,<br />
Tadi stream; holotype: KUMF 714; masyae is correct<br />
original spelling [Code art. 31.1.1 and Example], masyai<br />
is either an incorrect subsequent spelling or an unjustified<br />
emendation; noun in genitive, indeclinable)<br />
10.21.18 'Nemacheilus' monilis Hora, 1921<br />
Nemachilus monilis Hora, 1921a: 19, fig. 1 (type locality:<br />
India: Bhavani River, 10 miles from Mettupalaiyam; holotype:<br />
ZSI F 9981/1, Menon & Yazdani, 1968: 123; noun<br />
in genitive, indeclinable)<br />
Taxonomic notes. Generic position requires investigation.<br />
10.21.19 Nemacheilus olivaceus Boulenger, 1894<br />
Nemachilus olivaceus Boulenger, 1894a: 250 (type locality:<br />
Malaysia: Borneo: Sabah: Bongon; lectotype: BMNH<br />
1893.5.30.63, designated by Kottelat, 1984b: 234; adjective,<br />
-us, -a, -um)<br />
10.21.20 Nemacheilus ornatus Kottelat, 1990<br />
Nemacheilus ornatus Kottelat, 1990a: 61, fig. 34 (type locality:<br />
Thailand: Surat Thani Province: Khlong Sok at Ban<br />
Khlong Sok, 8°49'N 98°35'E [contra Eschmeyer &<br />
Fricke, 2010, "L. Sonkphan" in original description is<br />
not Lake Sonkphan but the name <strong>of</strong> one <strong>of</strong> the collectors];<br />
holotype: ZRC 38466 [ex ZSM 27469]; adjective,<br />
-us, -a, -um)<br />
10.21.21 Nemacheilus pallidus Kottelat, 1990<br />
Nemacheilus pallidus Kottelat, 1990a: 63, fig. 36 (type locality:<br />
Thailand: Lampang Province: Mae Nam Yom basin:<br />
Huai Mae Phlung from Ban Pong [18°42'N 99°58'E] to<br />
17 km upstream; holotype: ZRC 38468 [ex ZSM 27470];<br />
adjective, -us, -a, -um)<br />
10.21.22 Nemacheilus papillos Tan & Kottelat, 2009<br />
Nemacheilus papillos Tan & Kottelat, 2009: 53, fig. 40 (type<br />
locality: Indonesia: Sumatra: Sumatera Selatan: Sungai<br />
Sentang near Desa Sukajaya, about 5 km from road (turn<strong>of</strong>f<br />
at about 12 km on road from Bayung Lencir to Jambi);<br />
holotype: MZB 10994; noun in apposition, indeclinable)
10.21.9 Nemacheilus fasciatus, CMK 9178, 54.1 mm SL; Indonesia:<br />
Java: Brantas drainage.<br />
10.21.9 Nemacheilus fasciatus, CMK 9237, 61.0 mm SL.<br />
10.21.23 ? Nemacheilus papillosus (Perugia, 1893)<br />
Modigliania papillosa Perugia, 1893a: 246 (type locality:<br />
Indonesia: Sumatra: Balighe, Lake Toba; syntypes:<br />
MCSNG 9230 [6], ZMA 112.874 [2], Tortonese, 1961:<br />
188, Nijssen et al., 1993: 214; adjective, -us, -a, -um)<br />
10.21.24 Nemacheilus paucimaculatus Bohlen & Šlechtová,<br />
2011<br />
Nemacheilus paucimaculatus Bohlen & Šlechtová, 2011b:<br />
201, figs. 1–2 (type locality: Malaysia: Johor: tributary <strong>of</strong><br />
Segamat River upstream <strong>of</strong> Segamat, 2°28'47"N 103°<br />
05'13"E; holotype: ZRC 52361; adjective, -us, -a, -um)<br />
10.21.25 Nemacheilus pfeifferae (Bleeker, 1853)<br />
Cobitis Pfeifferi Bleeker, 1853b: 298 (type locality: Indonesia:<br />
Sumatra: Lake Meninju [Maninjau]; lectotype:<br />
RMNH 7053, designated by Alfred, 1961: 34; incorrect<br />
original spelling, as species is explicitly named for Mrs.<br />
Ida Pfeiffer, spelling must be emended into pfeifferae,<br />
Code arts. 31.1.2, 31.1.3; noun in genitive, indeclinable)<br />
Nemachilus dunckeri Ahl, 1922: 30 (type locality: Indonesia:<br />
Sumatra: Padang; holotype: ZMB 20546; noun in<br />
genitive, indeclinable)<br />
10.21.26 Nemacheilus platiceps Kottelat, 1990<br />
Nemacheilus platiceps Kottelat, 1990a: 66, figs. 38–39 (type<br />
locality: Vietnam: Trang Bom; holotype: NRM 15095;<br />
compound adjective, indeclinable)<br />
10.21.27 Nemacheilus saravacensis Boulenger, 1894<br />
Nemachilus saravacensis Boulenger, 1894a: 251 (type locality:<br />
Malaysia: Borneo: Sarawak: Senah; lectotype;<br />
BMNH 1893.3.6.277, designated by Kottelat, 1984b:<br />
236; adjective, -is, -is, -e)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
92<br />
10.21.28 Nemacheilus selangoricus, CMK 9636, 48.0 mm SL;<br />
Indonesia: Bangka.<br />
10.21.28 Nemacheilus selangoricus Duncker, 1904<br />
Nemachilus selangoricus Duncker, 1904: 175 (type locality:<br />
Malaysia: surroundings <strong>of</strong> Kuala Lumpur; lectotype:<br />
ZMH 386 [formerly 8464], designated by Ladiges et al.,<br />
1958: 159; adjective, -us, -a, -um)<br />
Nemachilus kuiperi de Beaufort, 1939: 190, fig. 1 (type locality:<br />
Indonesia: Billiton [Belitung]; lectotype: ZMA<br />
112.889, designated by Kottelat, 1984b: 255; noun in<br />
genitive, indeclinable)<br />
Nemacheilus trans-lineatus Fowler, 1939: 63, fig. 13 (type<br />
locality: Thailand: waterfall at Trang; holotype: ANSP<br />
68493; must be emended as translineatus, Code art.<br />
32.5.2.3; adjective, -us, -a, -um)<br />
10.21.29 Nemacheilus spiniferus Kottelat, 1984<br />
Noemacheilus spiniferus Kottelat, 1984b: 250, fig. 20 (type<br />
locality: Malaysia: Borneo: Sarawak: Fourth Division:<br />
Sungai Liam, tributary <strong>of</strong> Baram River, 3°19'N 114°45'E;<br />
holotype: ROM 39890; adjective, -us, -a, -um)<br />
10.21.30 'Nemacheilus' stigm<strong>of</strong>asciatus Arunachalam &<br />
Muralidharan, 2009<br />
Nemacheilus stigm<strong>of</strong>asciatus Arunachalam & Muralidharan,<br />
2009: 148, fig. 1 (type locality: India: Karnataka: Thuttinjet,<br />
Seethanathi River, 13°23'54.3"N 75°01'18.2"E;<br />
holotype: ZSI/SRS F7599; adjective, -us, -a, -um)<br />
Taxonomic notes. Generic position requires investigation.<br />
10.21.31 Nemacheilus tebo Hadiaty & Kottelat, 2009<br />
Nemacheilus tebo Hadiaty & Kottelat, 2009a: 120, fig. 3<br />
(type locality: Indonesia: Borneo: Kalimantan Timur:<br />
Berau Regency, Kelai district, Merapun village, Lake<br />
Tebo area, a pond at mouth <strong>of</strong> west cave, 17°00'13"N<br />
54°10'54"E; holotype: MZB 13367; noun in apposition,<br />
indeclinable)<br />
10.21.32 Nemacheilus troglocataractus Kottelat & Géry,<br />
1989<br />
Nemacheilus troglocataractus Kottelat & Géry, 1989: 273,<br />
fig. 1 (type locality: Thailand: Kanchanaburi Province:<br />
Tham Sai Yok Noi, 3 km north-northwest <strong>of</strong> Nam Tok;<br />
14°15'N 99°04'E; holotype: MHNG 2407.54; treated as<br />
a compound adjective, -us, -a, -um)<br />
10.21.33 Nemacheilus tuberigum Hadiaty & Siebert, 2001<br />
Nemacheilus tuberigum Hadiaty & Siebert, 2001: 183, fig. 1<br />
(type locality: Indonesia: Sumatra: Aceh Selatan: Kecamatan<br />
Kluet Selatan, Desa Pucuk Lembang, Gunung<br />
Leuser National Park, forest stream tributary to Sungai
10.22.1 Nemachilichthys ruppelli, CMK 14402, 53.4 mm SL;<br />
India: aquarium-fish trade.<br />
Lembang; holotype: MZB 9356; treated as noun in apposition,<br />
indeclinable)<br />
10.22 Nemachilichthys Day, 1878<br />
Nemachilichthys Day, 1878a: 611 (type species: Cobitis ruppelli<br />
Sykes, 1839a: 162, by monotypy). Gender masculine.<br />
10.22.1 Nemachilichthys ruppelli (Sykes, 1839)<br />
Cobitis Rupelli Sykes, 1839a: 162 (type localites: India:<br />
Beema River at Taimbournee / Mota Mola River near<br />
Poona [Pune, 18°28'N 73°48'E]; syntypes: LU, ? AMS<br />
B.7528 [1], Eschmeyer & Fricke, 2010; also in Sykes,<br />
1839b: 59, 1841: 366, pl. 64, fig. 17; rupelli is an incorrect<br />
original spelling since the species is explicitly named<br />
for Rüppell and must be emended into ruppelli, Code<br />
art. 32.5.1.1; noun in genitive, indeclinable)<br />
? Nemachilichthys shimogensis Narayan Rao, 1920: 62, pl. 2<br />
figs. 5–5b (type locality: India: Mysore [Karnataka]:<br />
Thunga River in Shimoga Town; holotype: BMNH<br />
1919.11.19.13–18 [1 <strong>of</strong> 6], Menon & Yazdani, 1968: 121;<br />
adjective, -is, -is, -e)<br />
Taxonomic notes. Nemachilichthys shimogensis has long<br />
been listed as a synonym <strong>of</strong> N. ruppelli. It is treated as distinct<br />
by Bănărescu & Nalbant (1995: 448) but no proper<br />
comparison has been published. The figures in the respective<br />
original descriptions show some differences, but are not<br />
really conclusive since several figures in Sykes (1841) do<br />
not seem totally accurate. This can be solved only by examination<br />
<strong>of</strong> topotypes <strong>of</strong> both nominal species.<br />
Nomenclatural notes. Narayan Rao explicitly stated that<br />
the holotype and about six paratypes were in BMNH. He<br />
also stated 'examples in ZSI', but these are not explicitly<br />
designated as types and thus are not types. The three 'syntypes'<br />
(ZSI F 9824/1–9826/1 [3]) listed by Menon & Yazdani<br />
(1968: 121) have no type status. There is a single series in<br />
BMNH [1919.11.19.13–18] labelled 'types'. As it seems that<br />
the holotype and the paratypes have been mixed, it is not<br />
possible to identify the holotype.<br />
10.23 Neonoemacheilus Zhu & Guo, 1985<br />
Neonoemacheilus Zhu & Guo, 1985: 321 (type species Nemacheilus<br />
labeosus Kottelat, 1982: 169, by original designation).<br />
Gender masculine.<br />
Infundibulatus Menon, 1987: 177 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus<br />
peguensis Hora, 1929: 321, by original designation).<br />
Gender masculine.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
93<br />
10.23.2 Neonoemacheilus labeosus, CMK 14634, 73.2 mm SL;<br />
Thailand: Salween drainage: Mae Nam Moei.<br />
10.23.2 Neonoemacheilus labeosus, CMK 14634, 73.2 mm SL.<br />
10.23.1 Neonoemacheilus assamensis (Menon, 1987)<br />
Noemacheilus assamensis Menon, 1987: 179 (type locality:<br />
India: Assam: Pagladia River; holotype: ZSI/SRS F 578;<br />
adjective, -is, -is, -e)<br />
10.23.2 Neonoemacheilus labeosus (Kottelat, 1982)<br />
Noemacheilus labeosus Kottelat, 1982: 169, fig. 1 (type<br />
locality: Thailand: Mae Hong Son Province: Salween<br />
River at Mae Sahm Leap, 17°59'N 97°44'E; holotype:<br />
USNM 230060; adjective, -us, -a, -um)<br />
10.23.3 Neonoemacheilus mengdingensis Zhu & Guo, in<br />
Zhu, 1989<br />
Neonoemacheilus mengdingensis Zhu & Guo, in Zhu, 1989:<br />
67, fig. 46 (type locality: China: Yunnan: Gengmaxian<br />
County: Nanding River near Mengding [Salween drainage];<br />
holotype: NGI 806206; adjective, -is, -is, -e)<br />
10.23.4 ? Neonoemacheilus morehensis Arunkumar, 2000<br />
Neonoemacheilus morehensis Arunkumar, 2000a: 44, fig. 1<br />
(type locality: India: Manipur: Lokchao River at Moreh,<br />
110 km south <strong>of</strong> Imphal [Chindwin drainage]; holotype:<br />
MUMF F 620/1A; adjective, -is, -is, -e)<br />
10.23.5 Neonoemacheilus peguensis (Hora, 1929)<br />
Nemachilus peguensis Hora, 1929: 321, pl. 14 figs. 1–2 (type<br />
locality: Burma: Pegu Yoma range; holotype: ZSI F<br />
11057/1, Menon & Yazdani, 1968: 123; adjective, -is,<br />
-is, -e)<br />
10.24 Oreonectes Günther, 1868<br />
Oreonectes Günther, 1868: 369 (type species: Oreonectes<br />
platycephalus Günther, 1868: 369, by monotypy). Gender<br />
masculine.<br />
Octonema Martens, 1869a: 608 (subgenus <strong>of</strong> Homaloptera
van Hasselt, 1823: 133; type species: Homaloptera rotundicauda<br />
Martens, 1869a: 608, by monotypy). Gender<br />
neuter.<br />
Species incertae sedis<br />
10.24.1 'Oreonectes' elongatus Tang, Zhao & Zhang, 2012<br />
Oreonectes elongatus Tang, Zhao & Zhang, 2012: 484, fig.<br />
2 (type locality: China: Guangxi: Huanjiang Co.: Donglei<br />
Cave in Mulun Township; holotype: ASIZB 189288;<br />
adjective, -us, -a, -um)<br />
Taxonomic notes. The original description and figure suggest<br />
this is not a species <strong>of</strong> Oreonectes since it has a forked<br />
(vs. rounded) caudal fin and the dorsal-fin origin clearly in<br />
front <strong>of</strong> the pelvic-fin origin (vs. conspicuously behind the<br />
pelvic-fin base).<br />
Species incertae sedis<br />
10.24.2 'Oreonectes' furcocaudalis Zhu & Cao, 1987<br />
Oreonectes furcocaudalis Zhu & Cao, 1987: 326, figs. 5–6<br />
(type locality: China: Guangxi: Ronshui County: an outlet<br />
<strong>of</strong> subterranean waters near Rongshui [25°04'N<br />
109°13'E; Romero et al., 2009: 253]; holotype: IHB<br />
83V331; compound noun, indeclinable)<br />
Taxonomic notes. The original description and figure suggest<br />
this is not a species <strong>of</strong> Oreonectes since it has a forked<br />
(vs. rounded) caudal fin and the dorsal-fin origin clearly in<br />
front <strong>of</strong> the pelvic-fin origin (vs. conspicuously behind the<br />
pelvic-fin base).<br />
Species incertae sedis<br />
10.24.3 'Oreonectes' macrolepis Huang, Du, Chen &<br />
Yang, 2009<br />
Oreonectes macrolepis Huang, Du, Chen & Yang, 2009: 446,<br />
fig. 1 (type locality: China: Guangxi: Maonan County:<br />
Huanjiang City: Da Cai Xiang [24°45'41"N<br />
108°22'08"E], Shen Long Gong He stream [a cave ?],<br />
Pearl River drainage; holotype: KIZ 2008008131; compound<br />
noun, indeclinable)<br />
Taxonomic notes. The original description and figure suggest<br />
this is not a species <strong>of</strong> Oreonectes since it has a forked<br />
(vs. rounded) caudal fin and the dorsal-fin origin clearly in<br />
front <strong>of</strong> the pelvic-fin origin (vs. conspicuously behind the<br />
pelvic-fin base).<br />
Species incertae sedis<br />
10.24.4 'Oreonectes' microphthalmus Du, Chen & Yang,<br />
2008<br />
Oreonectes microphthalmus Du, Chen & Yang, 2008: 28,<br />
fig. 4 (type locality: China: Guangxi: Du'an County<br />
[24°15'N 107°05'E; Romero et al., 2009: 254]; holotype:<br />
KIZ 2004009395; also spelt microphthalma p. 29, as first<br />
reviser I select microphthalmus as correct original spelling;<br />
compound adjective, -us, -a, -um)<br />
Taxonomic notes. The original description and figure suggest<br />
this is not a species <strong>of</strong> Oreonectes since it has a forked<br />
(vs. rounded) caudal fin and the dorsal-fin origin clearly in<br />
front <strong>of</strong> the pelvic-fin origin (vs. conspicuously behind the<br />
pelvic-fin base).<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
94<br />
Species incertae sedis<br />
10.24.5 'Oreonectes' translucens Zhang, Zhao & Zhang,<br />
2006<br />
Oreonectes translucens Zhang, Zhao & Zhang, 2006: 612,<br />
fig. 1 (type locality: China: Guangxi: Du'an County:<br />
Xia'ao Cave near Xia'ao town [24°15'N 107°05'E; Romero<br />
et al., 2009: 255]; holotype: ASIZB 94785, Tang, Zhao<br />
& Zhang, 2012: 489, fig. 5; participle, indeclinable)<br />
Taxonomic notes. The original description and figure suggest<br />
this is not a species <strong>of</strong> Oreonectes since it has a forked<br />
(vs. rounded) caudal fin and the dorsal-fin origin clearly in<br />
front <strong>of</strong> the pelvic-fin origin (vs. conspicuously behind the<br />
pelvic-fin base). Treated as synonym <strong>of</strong> Triplophysa longibarbata<br />
by Du et al. (2008: 33), treated as valid following<br />
Tang, Zhao & Zhang (2012: 486).<br />
10.24.6 Oreonectes anophthalmus Zheng, in Zheng, 1981<br />
Oreonectes anophthalmus Zheng, in Zheng, 1981: 162,<br />
fig. 134 (type locality: China: Guangxi: Wuming County:<br />
Qi Feng Shan, Taiji Cave [23'06"N 108°40'E; Romero<br />
et al., 2009: 252]; holotype: ASIZB 77001 [Zhang,<br />
1996: 500]; compound adjective, -us, -a, -um)<br />
10.24.7 Oreonectes guananensis Yang, Wei, Lan & Yang,<br />
2011<br />
Oreonectes guananensis Yang, Wei, Lan & Yang, 2011: 72,<br />
fig. 1 (type locality: China: Guangxi: Huanjiang County:<br />
Changmei town [24°57'19"N 108°28'14"E], Guan'an village<br />
[24°56'15"N 108°23'29"E] [Chinese text: tributary<br />
<strong>of</strong> Xiaohuanjiang, Xijiang drainage]; holotype: KIZ<br />
201003067; adjective, -is, -is, -e)<br />
10.24.8 Oreonectes luochengensis Yang, Wu, Wei & Yang,<br />
2011<br />
Oreonectes luochengensis Yang, Wu, Wei & Yang, 2011:<br />
209, figs. 1–2 (type locality: China: Guangxi: Luocheng<br />
County: cave approximately 2 km southeast <strong>of</strong> Tianhe<br />
town, within Tianhe drainage, itself in Xijiang drainage;<br />
holotype: KIZ 2010003073; adjective, -is, -is, -e)<br />
10.24.9 Oreonectes platycephalus Günther, 1868<br />
Oreonectes platycephalus Günther, 1868: 369 (type locality:<br />
China: Hongkong Mountains; syntypes: BMNH<br />
1848.7.12.6–7 [2], 1855.3.27.16–18 [3], 1858.9.19.155–<br />
173 [19], Eschmeyer & Fricke, 2010; compound adjective,<br />
-us, -a, -um)<br />
Homaloptera rotundicauda Martens, 1869a: 608 (type locality:<br />
China: Hong Kong; holotype: ZMB 6842; compound<br />
noun, indeclinable)<br />
Oreonectes yenlingi Lin, 1932b: 380 (type locality: China:<br />
Kwangtung [Guangdong]: Canton: White Cloud Mountain;<br />
holotype: FESC 1098; noun in genitive, indeclinable)<br />
10.24.10 Oreonectes polystigmus Du, Chen & Yang, 2008<br />
Oreonectes polystigmus Du, Chen & Yang, 2008: 30, fig. 7<br />
(type locality: China: Guangxi: Guilin City, Dabu village;<br />
holotype: KIZ 2001060507; compound adjective,<br />
-us, -a, -um)
10.24.11 Oreonectes retrodorsalis Lan, Yang & Chen,<br />
1995<br />
Oreonectes retrodorsalis Lan, Yang & Chen, 1995: 366, fig. 1<br />
(type locality: China: Guangxi: Nandan County: Liuzhai<br />
town, Longli village, outlet <strong>of</strong> underground river, 25°10'N<br />
107°10'E; holotype: KIZ 9110001; compound noun, indeclinable)<br />
10.25 Oxynoemacheilus Banarescu & Nalbant, 1966<br />
Oxynoemacheilus Bănărescu & Nalbant, 1966a: 153 (subgenus<br />
<strong>of</strong> Nemacheilus Bleeker, 1863a: 37; type species:<br />
Cobitis persa Heckel, 1848b: 266, by original designation).<br />
Gender masculine.<br />
Nun Bănărescu & Nalbant, in Bănărescu, Nalbant & Goren,<br />
1982: 23 (type species: Cobitis galilaea Günther, 1864:<br />
493, by original designation). Gender masculine.<br />
Nanoschistura Erk'akan, Nalbant & Özeren, 2007: 82, fig. 11<br />
(nomen nudum)<br />
Taxonomic notes. A very heterogenous (and likely polyphyletic)<br />
genus in which two main 'groups' can be recognised:<br />
species with a relatively contrasted pattern, relatively more<br />
slender body and caudal peduncle, forked caudal fin, and in<br />
some species a suborbital flap in males; and species with a<br />
stouter body and caudal peduncle, a dull brown marmorated<br />
pattern, a truncate or emarginate caudal fin, and no suborbital<br />
flap in males. This last group has <strong>of</strong>ten been included<br />
in Barbatula. Synonymy and included species partly based<br />
on Freyh<strong>of</strong> et al. (2012).<br />
10.25.1 Oxynoemacheilus anatolicus Erk'akan, Özeren<br />
& Nalbant, 2008<br />
Oxynemacheilus anatolica Erk'akan, Özeren & Nalbant,<br />
2008b: 117, fig. 4 (type locality: Turkey: tributary <strong>of</strong><br />
Karamanli Reservoir, Burdur [37°24'34.33"N<br />
29°49'54.94"E]; holotype: HUIC AKD 13; adjective,<br />
-us, -a, -um)<br />
10.25.2 Oxynoemacheilus angorae (Steindachner, 1897)<br />
Nemachilus Angorae Steindachner, 1897: 693, pl. 4 fig. 4a–<br />
c (type locality: Turkey: Tabakane-Su and Tschibuk-<br />
Tschai, both in the vicinity <strong>of</strong> Angora [Ankara]; syntypes:<br />
NMW [16]; noun in genitive, indeclinable)<br />
10.25.3 Oxynoemacheilus araxensis (Banarescu & Nalbant,<br />
in Banarescu, Nalbant & Balik, 1978)<br />
Orthrias angorae araxensis Bănărescu & Nalbant, in<br />
Bănărescu, Nalbant & Balik, 1978: 259, fig. 2, pl. 20<br />
figs. 1–4 (type locality: Turkey: Kandili Karassu, Upper<br />
Araxes drainage [actually Kandiili is a city on the upper<br />
Karasu, in the Tigris-Euphrates drainage; Erk'akan &<br />
Kuru, 1986a: 108]; holotype: ZMH 4827; adjective, -is,<br />
-is, -e)<br />
10.25.4 Oxynoemacheilus argyrogramma (Heckel, 1848)<br />
Cobitis argyrogramma Heckel, 1848a: 239, pl. 18 fig. 3 (type<br />
locality: Syria: Aleppo; syntypes: NMW 48541 [8], 59913<br />
[4], Eschmeyer & Fricke, 2010; compound noun, indeclinable)<br />
Noemacheilus insignis euphraticus Bănărescu & Nalbant,<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
95<br />
1964: 175, pl. 7 figs. 11–12 (type locality: Turkey: eastern<br />
Anatolia: Malay [probably Malatya]; holotype: ZMH<br />
H 1889, Wilkens, 1977: 158; adjective, -us, -a, -um)<br />
Noemacheilus tschaiyssuensis Bănărescu & Nalbant, 1964:<br />
177, pl. 8 fig. 13 (type locality: Turkey: Tchay-Su, between<br />
Keyson (37°30'N 37°50'E) and Gaziantep (37°00'N<br />
37°30'E), Ceyhan drainage; holotype: BMNH<br />
1935.9.12.57; adjective, -is, -is, -e)<br />
Taxonomic notes. Krupp & Schneider (1991: 26) treated<br />
C. argyrogramma as a synonym <strong>of</strong> O. insignis, without discussion.<br />
Placed in Oxynoemacheilus by Prok<strong>of</strong>iev (2009b:<br />
880), but also as a possible synonym <strong>of</strong> Paracobitis tigris.<br />
Noemacheilus insignis euphraticus was treated as a valid<br />
species by Nalbant & Bianco (1999: 110) and as subspecies<br />
<strong>of</strong> O. brandtii by Prok<strong>of</strong>iev (2009b: 880). Synonymy follows<br />
Freyh<strong>of</strong> et al. (2012).<br />
10.25.5 Oxynoemacheilus banarescui (Delmastro, 1892)<br />
Orthrias brandti banarescui Delmastro, 1982: 53, figs. 1–3<br />
(type locality: Turkey: Devrekani creek near Devrekani,<br />
Kocacay basin; holotype: MCSNC; noun in genitive, indeclinable)<br />
10.25.6 Oxynoemacheilus bergianus (Derzhavin, 1934)<br />
Nemachilus bergianus Derzhavin, 1934: 109, fig. 8 (type<br />
locality: Iran: Kisum in Sefi-rud basin and Shah-rud, a<br />
tributary <strong>of</strong> Sefid-rud; syntype: ZISP 25433 [1], Eschmeyer<br />
& Fricke, 2010; adjective, -us, -a, -um)<br />
10.25.7 Oxynoemacheilus brandtii (Kessler, 1877)<br />
Nemachilus Brandtii Kessler, 1877: 174, pl. 6 fig. 23 (type<br />
locality: Georgia: Kara River [Kura] at Tiflis [Tbilisi];<br />
syntypes [5]: BMNH 1897.7.5.39 [1], Eschmeyer &<br />
Fricke, 2010; noun in genitive, indeclinable)<br />
? Nemachilus brandti gibbusnazus Elanidze, 1983: 209 (type<br />
locality: Georgia: upper reaches <strong>of</strong> Alazani River at Alvani<br />
village [42°03'06"N 45°14'15"E], a left bank tributary<br />
<strong>of</strong> Kura; syntypes [60]: LU; compound noun, indeclinable)<br />
10.25.8 Oxynoemacheilus bureschi (Drensky, 1928)<br />
Nemacheilus Bureschi Drensky, 1928: 160, 179, fig. 1 (type<br />
locality: Bulgaria: Radomir District: Struma River, near<br />
Semen railway station / tributaries <strong>of</strong> Struma River in<br />
Küstendil and Dupniza districts [Kjustendil: 42°17'60"N<br />
22°41'24"E; Dupnitsa: 42°16'12"N 23°7'12"E]; syntypes:<br />
NHMS [2]; noun in genitive, indeclinable)<br />
Orthrias brandti macedonicus Šorić, 2000: 83, fig. 1 (type<br />
locality: FYROM: steam Dosnica near Demir Kapija,<br />
Vardar drainage; holotype: BIKU; adjective, -us, -a,<br />
-um)<br />
10.25.9 Oxynoemacheilus ceyhanensis (Erk'akan, Nalbant<br />
& Özeren, 2007)<br />
Schistura ceyhanensis Erk'akan, Nalbant & Özeren, 2007:<br />
80, fig. 10 (type locality: Turkey: Kahramanmaras Province:<br />
Elbistan District: Yalak village, 38°39'N 36°37'E;<br />
holotype: HUIC CEY-1; adjective, -is, -is, -e)<br />
Taxonomic notes. The coordinates <strong>of</strong> the type locality are<br />
the same as for the holotypes <strong>of</strong> Barbatula paucilepis, Schis-
10.24.9 Oreonectes platycephalus, CMK 7207, 43.8 mm SL; China:<br />
Hongkong.<br />
tura evreni and S. seyhanicola, although the localities are<br />
different.<br />
10.25.10 Oxynoemacheilus cinicus (Erk'akan, Nalbant &<br />
Özeren 2007)<br />
Barbatula cinica Erk'akan, Nalbant & Özeren, 2007: 73,<br />
fig. 4 (type locality: Turkey: road from Kütahya to Denizli,<br />
Cin Stream, 39°40'N 29°30'E; holotype: HUIC BM-<br />
3; adjective, -us, -a, -um)<br />
10.25.11 Oxynoemacheilus cyri (Berg, 1910)<br />
Nemachilus tigris cyri Berg, 1910a: 0165, 0170 (type locality:<br />
Turkey: Kars Province: Okam village, Gel plain, Kura<br />
drainage; syntypes: ZISP 13291 [6+], 16885 [2], Eschmeyer<br />
& Fricke, 2010; available by indication to Nemacheilus<br />
tigris <strong>of</strong> Berg, 1906b: 39 [Berg, 1910a originally<br />
refers to the 1906 paper as published in 1907, but he<br />
later corrected the date as 1906; see, e.g. Berg, 1949:<br />
878]; also in Berg, 1910b: 128, with locality as "sources<br />
<strong>of</strong> Kura River, Caucasus"; noun in genitive, indeclinable)<br />
Taxonomic notes. Generic position follows Freyh<strong>of</strong> et al.<br />
(2012).<br />
10.25.12 Oxynoemacheilus ercisianus (Erk'akan & Kuru,<br />
1986)<br />
Orthrias angorae ercisianus Erk'akan & Kuru, 1986b: 161,<br />
fig. 1 (type locality: Turkey: Ercis stream, a tributary <strong>of</strong><br />
Lake Van; holotype: HUIC; adjective, -us, -a, -um)<br />
Nemacheilus pulsiz Krupp, 1992: 291 (type locality: Turkey:<br />
Van Gölü basin: road from Ercis to Patnos; holotype:<br />
ZMH 7995)<br />
Taxonomic notes. Generic position follows Freyh<strong>of</strong> et al.<br />
(2012).<br />
10.25.13 Oxynoemacheilus erdali (Erk'akan, Nalbant &<br />
Özeren, 2007)<br />
Barbatula erdali Erk'akan, Nalbant & Özeren, 2007: 78,<br />
fig. 8 (type locality: Turkey: Murat River - Agri, 39°40'N<br />
43°44'E; holotype: HUIC F12; noun in genitive, indeclinable)<br />
10.25.14 Oxynoemacheilus eregliensis (Banarescu &<br />
Nalbant, in Banarescu, Nalbant & Balik, 1978)<br />
Orthrias angorae eregliensis Bănărescu & Nalbant, in Bănărescu,<br />
Nalbant & Balik, 1978: 258, fig. 1 (type locality:<br />
Turkey: central Anatolia: Eregli; holotype: ZMH 1921;<br />
adjective, -is, -is, -e)<br />
10.25.15 Oxynoemacheilus evreni (Erk'akan, Nalbant &<br />
Özeren, 2007)<br />
Schistura evreni Erk'akan, Nalbant & Özeren, 2007: 82,<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
96<br />
fig. 12 (type locality: Turkey: Tekir stream, Göksu basin,<br />
38°39'N 36°37'E; holotype: HUIC SEY-3; noun in<br />
genitive, indeclinable)<br />
Taxonomic notes. In the original description, the coordinates<br />
are the same as for the holotypes <strong>of</strong> Barbatula paucilepis,<br />
Schistura ceyhanensis and S. seyhanicola, although<br />
the localities are different.<br />
10.25.16 Oxynoemacheilus frenatus (Heckel, 1843)<br />
Cobitis frenata Heckel, 1843: 1086, pl. 12 fig. 1 (type locality:<br />
Iraq: Tigris River at Mossul; syntypes: NMW 48552<br />
[5], ? NRM 15477 [1]; invalid lectotype designation by<br />
Nalbant & Bianco, 1999: 112; adjective, -us, -a, -um)<br />
Nemachilus frenatus afrenatus Battalgil, 1941: 183 (type<br />
locality: Turkey: Diyarbakir [Tigris drainage; 37°55'N<br />
40°14'E]; syntypes: LU; adjective, -us, -a, -um)<br />
Nomenclatural notes. Nalbant & Bianco (1999: 112) wrote:<br />
"A lectotype has been selected from type material <strong>of</strong> this<br />
species [B. frenata] (Type locality: Mosul), housed in NMW,<br />
examined by one <strong>of</strong> us (TN). The lectotype corresponds to<br />
the description and the illustration given by Heckel (1843)".<br />
No information is provided that allows to identify this 'lectotype'<br />
among the syntypes and, therefore, the designation is<br />
not valid. The Code art. 74.5 requires that a lectotype must<br />
be "a particular syntype" "unambiguously selected".<br />
10.25.17 Oxynoemacheilus galilaeus (Günther, 1864)<br />
Cobitis galilaea Günther, 1864: 493 (type locality: Israel:<br />
Lake <strong>of</strong> Galilee [Lake Kinneret]; holotype: BMNH<br />
1863.11.3.8; adjective -us, -a, -um)<br />
Nomenclatural notes. The word galilaea can be either the<br />
nominative singular <strong>of</strong> the noun Galilaea or the nominative<br />
singular feminine <strong>of</strong> the adjective galilaeus. Günther (1864)<br />
did not indicate whether the word was treated as a noun or<br />
an adjective and, under a blind application <strong>of</strong> Code art. 31.2.2,<br />
galilaea should then be treated as a noun in apposition and<br />
be invariable. But, art. 31.2.2 says "where it may be regarded<br />
as either and the evidence <strong>of</strong> usage is not decisive". While<br />
galilaea could be regarded grammatically as either noun or<br />
adjective, the usage (syntax and meaning) shows it is an adjective.<br />
For an author <strong>of</strong> that period, it would have been<br />
awkard to use the noun Galilaea in the nominative in apposition<br />
(C. galilaea would then translate as 'Galilee-loach'); he<br />
would have used the genitive (C. galilaeae, which would<br />
translate as 'loach <strong>of</strong> Galilee') or the adjective (C. galilaea<br />
would then translate as 'Galilean loach'). The last option<br />
would have been the best, and from external evidence we<br />
know that Günther later (1868: 355) treated the name as an<br />
adjective when he transferred the species to Nemacheilus<br />
("Nemachilus galilaeus"). Therefore the name is treated here<br />
as an adjective.<br />
10.25.18 Oxynoemacheilus germencicus (Erk'akan, Nalbant<br />
& Özeren, 2007)<br />
Barbatula germencica Erk'akan, Nalbant & Özeren, 2007:<br />
70, fig. 2 (type locality: Turkey: Aydin Province: "Germencik<br />
15 th km", 37°38'N 27°28'E; holotype: HUIC BM-<br />
1; adjective, -us, -a, -um)
10.25.8 Oxynoemacheilus bureschi, CMK 17403, 58.5 mm SL;<br />
FYR <strong>of</strong> Macedonia: Vardar drainage.<br />
10.25.7 Oxynoemacheilus brandti, CMK 19463, 45.4 mm SL;<br />
Turkey: Arax drainage.<br />
10.25.19 Oxynoemacheilus hamwii (Krupp & Schneider,<br />
1991)<br />
Nemacheilus hamwii Krupp & Schneider, 1991: 24, figs. 1–<br />
5 (type locality: Syria: Nahr Afrin in Afrin, 36°31'N<br />
36°52'E; holotype: SMF 17398)<br />
10.25.20 Oxynoemacheilus insignis (Heckel, 1843)<br />
Cobitis insignis Heckel, 1843: 1087, pl. 12 fig. 2 (type locality:<br />
Syria: Damascus [33°30'N 36°20'E]; syntypes: NMW<br />
[4], SMF 166 [2], Eschmeyer & Fricke, 2010; adjective,<br />
-is, -is, -e)<br />
Noemacheilus angorae jordanicus Bănărescu & Nalbant,<br />
1966b: 329, fig. 1, pl. 5 fig. 1 (type locality: Jordan: Wadi<br />
Kufrinja, east bank <strong>of</strong> Jordan River, 1.5 km upstream <strong>of</strong><br />
its delta [32°16'N 35°34'E]; holotype: ZMH H 2738,<br />
Wilkens, 1977: 158; adjective, -us, -a, -um)<br />
? Nemacheilus insignis tortonesei Bănărescu & Nalbant,<br />
1966b: 331, figs. 2–3, pl. 4 figs. 2–4 (type locality: Jordan:<br />
springs <strong>of</strong> Wadi-Um-ed-Dananir, tributary <strong>of</strong> Zerqua,<br />
about 25 km north <strong>of</strong> Amman, Jordan basin [about<br />
32°04'N 35°50'E]; holotype: ZMH H 2742, Wilkens,<br />
1977: 158; noun in genitive, indeclinable)<br />
Orthrias dori Goren & Bănărescu, in Bănărescu, Nalbant &<br />
Goren, 1982: 12, figs. 9–13 (type locality: Israel: Nahal<br />
Malkoah, Bet She'an Valley [about 32°25'N 35°30'E];<br />
holotype: TAU 8278; noun in genitive, indeclinable)<br />
Orthrias pantheroides Goren & Nalbant, in Bănărescu, Nalbant<br />
& Goren, 1982: 16, figs. 14–18, pl. 2 figs. 9–10<br />
(type locality: Israel: Tanuria Springs, Golan Heights<br />
[32°58'N 35°48'E]; holotype: TAU 8292)<br />
Orthrias israeliticus Goren & Nalbant, in Bănărescu, Nalbant<br />
& Goren, 1982: 19, figs. 19–23 (type locality: Israel:<br />
'Ein Rewaia, Bet She'an Valley [32°28'N 35°28'E];<br />
holotype: TAU 3073; adjective, -us, -a, -um)<br />
Taxonomic notes. Synonymy follows Krupp & Schneider<br />
(1989: 380) and Freyh<strong>of</strong> et al. (2012).<br />
10.25.21 Oxynoemacheilus kaynaki Erk'akan, Özeren &<br />
Nalbant, 2008<br />
Oxynemacheilus kaynaki Erk'akan, Özeren & Nalbant,<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
97<br />
10.25.11 Oxynoemacheilus cyri, CMK 18530, 63.5 mm SL; Turkey:<br />
Arax drainage.<br />
10.25.12 Oxynoemacheilus ercisianus, CMK 22393, 71.3 mm SL;<br />
Turkey: Lake Van basin.<br />
2008b: 115, fig. 1 (type locality: Turkey: Goksu River,<br />
Nurhak, Elbistan, Euphrates drainage; 37°53'22.82"N<br />
37°22'19.99"E; holotype: HUIC F-20; noun in genitive,<br />
indeclinable)<br />
10.25.22 Oxynoemacheilus kermanshahensis (Banarescu<br />
& Nalbant, 1966)<br />
Noemacheilus kermanshahensis Bănărescu & Nalbant,<br />
1966a: 151, figs. 1–2, pl. 19 fig. 2 (type locality: Iran:<br />
Kermanshah in Karun River drainage, a tributary <strong>of</strong> lower<br />
Euphrates [Tigris drainage: stream Quareh Su at Kermanshah,<br />
a tributary <strong>of</strong> Shimarek River, itself a tributary<br />
<strong>of</strong> Karkheh; Nalbant & Bianco, 1999: 110; Karkheh is<br />
an endorheic drainage; 32°29'N 48°07'E]; holotype:<br />
ZMUC P 2787; adjective, -is, -is, -e)<br />
10.25.23 Oxynoemacheilus kiabii Golzarianpour, Abdoli<br />
& Freyh<strong>of</strong>, 2011<br />
Oxynoemacheilus kiabii Golzarianpour, Abdoli & Freyh<strong>of</strong>,<br />
2011: 202, figs. 1–3 (type locality: Iran: Hamadan Province:<br />
stream Dehnoo, 3 km west <strong>of</strong> Nahavand on road<br />
from Nahavand to Sarab-e-Gamasiab, 34°10'N 48°24'E;<br />
holotype: ZFMK 41847; noun in genitive, indeclinable)<br />
10.25.24 Oxynoemacheilus kosswigi (Erk'akan & Kuru,<br />
1986)<br />
Orthrias angorae kosswigi Erk'akan & Kuru, 1986a: 107,<br />
fig. 1 (type locality: Turkey: Sivas Province: Yildiz çayi,<br />
Yildizeli [Kizil Irmak drainage, 39°52'N 36°37'E]; holotype:<br />
HUIC; noun in genitive, indeclinable)
10.25.25 Oxynoemacheilus lenkoranensis (Abdurakhmanov,<br />
1962)<br />
? Nemacheilus bergi Gratsianov, 1907: 163, 167 (based on<br />
Nemachilus spec. ? <strong>of</strong> Berg, 1899: 30, 72; type locality:<br />
Azerbaijan: Akstapha River [Akstafa; Aghstev, 41°15'N<br />
45°26'E], right tributary <strong>of</strong> Kura River, Caspian basin;<br />
syntypes [2]: ZMT; noun in genitive, indeclinable)<br />
Nemacheilus angorae lenkoranensis Abdurakhmanov, 1962:<br />
285, fig. 50 (type locality: Azerbaidjan: Lenkoran; holotype:<br />
LU [? Baku Institute <strong>of</strong> Zoology]; adjective, -is,<br />
-is, -e)<br />
? Nemachilus angorae alasanicus Elanidze, 1983: 204 (type<br />
locality: Georgia: upper reaches <strong>of</strong> Alazani River at Alvani<br />
village [42°03'06"N 45°14'15"E], a left bank tributary<br />
<strong>of</strong> Kura; syntypes [54]: LU; adjective, -us, -a, -um)<br />
10.25.26 Oxynoemacheilus leontinae (Lortet, 1883)<br />
Nemachilus leontinae Lortet, 1883: 171, pl. 18 fig. 1 (type<br />
locality: Israel: Lake <strong>of</strong> Tiberias [Lake Kinneret]; lectotype:<br />
MGHNL 3665, designated by Krupp & Schneider,<br />
1989: 393; noun in genitive, indeclinable)<br />
Taxonomic notes. Treated as a synonym <strong>of</strong> O. panthera by<br />
Prok<strong>of</strong>iev (2009b: 880) and Bănărescu et al. (1982: 16).<br />
10.25.27 Oxynoemacheilus mediterraneus (Erk'akan,<br />
Nalbant & Özeren, 2007)<br />
Barbatula mediterraneus Erk'akan, Nalbant & Özeren, 2007:<br />
74, fig. 5 (type locality: Turkey: Egirdir Province: Candir,<br />
Aksu Stream, 37°38'N 30°31'E; holotype: HUIC<br />
AKD-2a; adjective, -us, -a, -um)<br />
Taxonomic notes. Coordinates possibly erroneous; they are<br />
identical to those <strong>of</strong> the type locality <strong>of</strong> Seminemacheilus<br />
ispartensis, which has different locality data.<br />
10.25.28 Oxynoemacheilus merga (Krynicki, in Nordmann,<br />
1840)<br />
Cobitis merga Krynicki, in Nordmann, 1840: 470 (type locality:<br />
Russia: Stavropol Krai: North Caucasian Federal<br />
District: Podcoumok River near de Paitigorsk [Podkumok,<br />
tributary <strong>of</strong> Kuman, near Pyatigorsk; 44°2'0"N<br />
43°03"00"E]; types: LU; noun in apposition, indeclinable)<br />
Cobitis nurga Valenciennes, in Cuvier & Valenciennes, 1846:<br />
24 (unjustified emendation <strong>of</strong> Cobitis merga Krynicki,<br />
1840: 470)<br />
Nemacheilus terekensis Gratzianow, 1907: 163, 167 (type<br />
locality: Russia: North Osetia-Alanya: Terek River at<br />
Vladjkavkaz [Vladikavkas]; syntypes: ZMMU P-3318<br />
[24], Vasilieva et al., in Pavlinov & Borissenko, 2001:<br />
25; adjective, -is, -is, -e)<br />
10.25.29 Oxynoemacheilus namiri (Krupp & Schneider,<br />
1991)<br />
Nemacheilus namiri Krupp & Schneider, 1991: 28, figs. 7–<br />
13 (type locality: Syria: Orontes River at Jisr ash-Shughur,<br />
35°48'N 36°19'E; holotype: SMF 17837; noun in<br />
apposition, indeclinable)<br />
Taxonomic notes. Generic position follows Freyh<strong>of</strong> et al.<br />
(2012).<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
98<br />
10.25.30 'Oxynoemacheilus' oxianus (Kessler, 1877)<br />
Nemachilus Brandtii var. oxiana Kessler, 1877: 177 (type<br />
locality: Uzbekistan: lower reaches <strong>of</strong> Amu-Darya; holotype:<br />
ZISP; adjective, -us, -a, -um)<br />
Nemachilus flavus Berg, 1905: 193, pl. 6 figs. 3–5 (type locality:<br />
Uzbekistan: Amu-Darya below Chardzkui [Chardjui,<br />
Chardzhou, now Türkmenabat, Turkemenistan;<br />
39°05'00"N 63°34'00"E]; lectotype: ZISP 13282, designated<br />
by Berg, 1949: 879 [caption <strong>of</strong> fig. 625], or if not<br />
valid from Russian text, from translation in Berg, 1965:<br />
444; adjective, -us, -a, -um)<br />
Nemacheilus oxianus natio zeravschani Turdakov, 1936: 199<br />
(an infrasubspecific name, not available; locality: Uzbekistan:<br />
Zeravshan drainage: Ak-Darya River)<br />
Taxonomic notes. Apparently represents an unnamed genus<br />
(see Prok<strong>of</strong>iev, 2009b: 896).<br />
10.25.31 Oxynoemacheilus panthera (Heckel, 1843)<br />
Cobitis panthera Heckel, 1843: 1087, pl. 12 fig. 2 (type locality:<br />
Syria: Damascus [33°30'N 36°20'E]; syntypes: NMW<br />
48565 [3], 15478 [1]; noun in apposition, indeclinable)<br />
Cobitis leopardus Heckel, 1843: 1089 (type locality: Syria:<br />
Damascus [33°30'N 36°20'E]; holotype: NMW 48559;<br />
simultaneous subjective synonym <strong>of</strong> Cobitis panthera<br />
Heckel, 1843: 1087; first reviser [Günther, 1868: 355]<br />
gave precedence to C. panthera; also in Heckel, 1848a:<br />
241, pl. 18, fig. 4; noun in apposition, indeclinable)<br />
10.25.32 Oxynoemacheilus paucilepis (Erk'akan, Nalbant<br />
& Özeren, 2007)<br />
Barbatula paucilepis Erk'akan, Nalbant & Özeren, 2007: 79,<br />
fig. 9 (type locality: Turkey: Sivas Province: Mancilik<br />
stream, Gürün, 38°39'N 37°38'E; holotype: HUIC F2;<br />
also spelt paucllepis on p. 70, fig. 1, an obvious inadvertent<br />
error, thus incorrect original spelling [Code art.<br />
32.5.1]; compound noun, indeclinable)<br />
Taxonomic notes. In the original description, this species is<br />
said to be "close" to and compared with Paracobitis tigris<br />
but not with other species <strong>of</strong> Barbatula. The coordinates <strong>of</strong><br />
the type locality are the same as for the holotypes <strong>of</strong> Schistura<br />
ceyhanensis, S. evreni and S. seyhanicola, although they<br />
have different locality data. Generic position follows Freyh<strong>of</strong><br />
et al. (2012).<br />
10.25.33 Oxynoemacheilus persa (Heckel, 1848)<br />
Cobitis persa Heckel, 1848b: 266 (type locality: Iran: springs<br />
around Persepolis [Marvdasht Plain, about 70 km northeast<br />
<strong>of</strong> Shiraz; 29°56'04"N 52°53'29"E]; holotype: NMW<br />
48567, Eschmeyer & Fricke, 2010; noun in apposition,<br />
indeclinable)<br />
Orthrias farsicus Nalbant & Bianco, 1999: 111, fig. 6 (type<br />
locality: Iran: Kor River near Persepolis [Marvdasht<br />
Plain, about 70 km northeast <strong>of</strong> Shiraz; 29°56'04"N 52°<br />
53'29"E]; holotype: IZA 7823; adjective, -us, -a, -um)<br />
Taxonomic notes. Synonymy follows Freyh<strong>of</strong> et al. (2012).<br />
10.25.34 Oxynoemacheilus phoxinoides (Erk'akan, Nalbant<br />
& Özeren, 2007)<br />
Barbatula phoxinoides Erk'akan, Nalbant & Özeren, 2007:<br />
75, fig. 6 (type locality: Turkey: Iznik Province: 40°41'N
29°30'E; holotype: HUIC MAR-7g; adjective, indeclinable)<br />
10.25.35 Oxynoemacheilus pindus (Economidis, 2005)<br />
Orthrias pindus Economidis, 1991: 26, p. 42 (fig. 4), 1992:<br />
75, 49 (nomen nudum; locality: Greece: Epiros: Aoos<br />
River drainage)<br />
Barbatula pindus Economidis, 2005: 67, figs. 1–2 (type<br />
locality: Greece: Epiros: Aoos drainage, stream Sarantoporos<br />
near village Drosopygi; holotype: DZAUT 1977-<br />
215a; noun in apposition, indeclinable)<br />
10.25.36 Oxynoemacheilus samanticus (Banarescu &<br />
Nalbant, in Banarescu, Nalbant & Balik, 1978)<br />
Orthrias brandti samantica Bănărescu & Nalbant, in Bănărescu,<br />
Nalbant & Balik, 1978: 263, fig. 4. (type locality:<br />
Turkey: tributary <strong>of</strong> Samanti, between Pinarbasi and<br />
Sarizi, Seyhan drainage; holotype: ZMH 3633; adjective,<br />
-us, -a, -um)<br />
10.25.37 Oxynoemacheilus seyhanensis (Banarescu, 1968)<br />
Noemacheilus tigris seyhanensis Bănărescu, 1968: 355, pl. 3<br />
figs. 2–3 (type locality: Turkey: between Viransehir and<br />
Kazancik, upper Seyhan drainage; holotype: ZMH H<br />
4014, Wilkens, 1977: 159; adjective, -is, -is, -e)<br />
Taxonomic notes. Generic position and validity follow Freyh<strong>of</strong><br />
et al. (2012).<br />
10.25.38 Oxynoemacheilus seyhanicola (Erk'akan, Nalbant<br />
& Özeren, 2007)<br />
Schistura seyhanicola Erk'akan, Nalbant & Özeren, 2007:<br />
81, fig. 11 (type locality: Turkey: dam bridge near Adana,<br />
Seyhan drainage, 38°39'N 36°37'E; holotype: HUIC SEY-<br />
1; also spelt seyhanicolo on p. 82, fig. 11, an obvious<br />
inadvertent error, thus incorrect original spelling [Code<br />
art. 32.5.1]; noun in apposition, indeclinable)<br />
Taxonomic notes. The coordinates <strong>of</strong> the type locality are<br />
the same as for the holotypes <strong>of</strong> Barbatula paucilepis, Schistura<br />
evreni and S. ceyhanensis, although the localities are<br />
different.<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
'inhabitant <strong>of</strong>' are nouns and indeclinable.<br />
10.25.39 Oxynoemacheilus simavicus (Balik & Banarescu,<br />
in Banarescu, Nalbant & Balik, 1978)<br />
Orthrias brandti simavica Balik & Bănărescu, in Bănărescu,<br />
Nalbant & Balik, 1978: 261, fig. 3 (type locality: Turkey:<br />
"Simav creek, Kütahya" [seems impossible; Simav<br />
stream, flows through Balikesir (39°39'N 27°53'E), joins<br />
Koca and flows to Sea <strong>of</strong> Marmara; Kütahya (39°25'N<br />
29°56'E) is in Sakarya drainage, which flows to Black<br />
Sea; see Stoumboudi et al., 2006: 143]; holotype: ISBB<br />
2976; adjective, -us, -a, -um)<br />
10.25.40 Oxynoemacheilus theophilii Stoumboudi, Kottelat<br />
& Barbieri, 2006<br />
Oxynoemacheilus theophilii Stoumboudi, Kottelat & Barbieri,<br />
2006: 140, fig. 8 (type locality: Greece: Lesbos<br />
Island: Tsingou springs, in Evergetoulas drainage; holotype:<br />
MHNG 2679.009; noun in genitive, indeclinable)<br />
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Barbatula bergamensis Erk'akan, Nalbant & Özeren, 2007:<br />
71, fig. 3 (type locality: Turkey: Bergama Province:<br />
Kozak (Madra) creek, 39°40'N 27°28'E; holotype: HUIC<br />
KE-1; adjective, -is, -is, -e)<br />
10.25.41 Oxynoemacheilus tigris (Heckel, 1843)<br />
Cobitis tigris Heckel, 1843: 1088, pl. 12 fig. 4 (type locality:<br />
Syria: Kueik stream near Aleppo; syntypes: NMW<br />
48441 [2], 49444 [2], 49445 [4], 49446 [2], SMF 405<br />
[3], Eschmeyer & Fricke, 2010; noun in apposition, indeclinable)<br />
Taxonomic notes. Generic position follows Freyh<strong>of</strong> et al.<br />
(2012).<br />
10.25.42 Oxynoemacheilus tongiorgii (Nalbant & Bianco,<br />
1999)<br />
Seminemacheilus tongiorgii Nalbant & Bianco, 1999: 113,<br />
fig. 11 (type locality: Iran: large spring near Darab<br />
[28°44'56"N 54°33'05"E], Kul River drainage; holotype:<br />
IZA 801; noun in genitive, indeclinable)<br />
10.26 Paracobitis Bleeker, 1863<br />
Paracobitis Bleeker, 1863a: 37 (type species: Cobitis malapterura<br />
Valenciennes, in Cuvier & Valenciennes, 1846:<br />
88, by original designation; also in Bleeker, 1863c: 3).<br />
Gender feminine.<br />
Pseudodon Kessler, 1874: 40 (type species: Cobitis longicauda<br />
Kessler, 1872: 65, by monotypy; junior homonym<br />
<strong>of</strong> Pseudodon Gould, 1845: 161 in Mollusca). Gender<br />
masculine.<br />
Adiposia Annandale & Hora, 1920: 182 (type species: Nemacheilus<br />
macmahoni Chaudhuri, 1910b: 341, by original<br />
designation). Gender feminine.<br />
Taxonomic notes. See Homatula for the Chinese species<br />
usually placed in Paracobitis.<br />
10.26.1 Paracobitis boutanensis (M'Clelland & Griffith,<br />
in M'Clelland, 1842)<br />
Cobitis boutanensis M'Clelland & Griffith, in M'Clelland,<br />
1842: 586 (type locality: "Boutan, on the Mishmee mountains"<br />
[erroneous, Bolas Pass, Helmand drainage, according<br />
to Hora, 1928: 482; Pakistan: Balochistan: Bholan<br />
or Bolan Pass, 28°01'N 67°04'E]; holotype: BMNH<br />
1860.3.14.775; adjective, -is, -is, -e)<br />
Taxonomic notes. M'Clelland indicated the type locality as<br />
'Boutan, on the Mishmee mountains'. Hora (1928: 482) corrected<br />
it as "Bolas Pass, Helmand". It seems to be Bolan or<br />
Bohlan Pass, see above. Griffith (in M'Clelland, 1842: 562)<br />
listed the species he obtained in the rivers <strong>of</strong> Bolan pass; the<br />
list does not mention loaches but it is probably not exhaustive.<br />
10.26.2 Paracobitis ghazniensis (Banarescu & Nalbant,<br />
1966)<br />
Noemacheilus ghazniensis Bănărescu & Nalbant, 1966a:<br />
162, fig. 7, pl. 20 figs. 4–5 (type locality: East Afghanistan:<br />
Ghazni [33°33'N 68°26'E], on Ghazni River, tributary<br />
<strong>of</strong> Ab-i-istadah Lake, Helmand drainage; holotype:<br />
BMNH 1944.4.1.1; adjective, -is, -is, -e)
10.26.3 Paracobitis iranica, CMK 21898, 60.8 mm SL; Iran:<br />
Qomtra.<br />
10.26.3 Paracobitis iranica Nalbant & Bianco, 1999<br />
Paracobitis iranica Nalbant & Bianco, 1999: 114, fig. 13<br />
(type locality: Iran: Qom River near Qom City [34°38'N<br />
50°53'E]; holotype: IZA 7831; adjective, -us, -a, -um)<br />
10.26.4 Paracobitis longicauda (Kessler)<br />
Cobitis longicauda Kessler, 1872: 65, pl. 11 figs. 30–31 (type<br />
locality: Uzbekistan: Ak-Darya [tributary <strong>of</strong> Zeravshan];<br />
holotype: ZISP 2686; compound noun, indeclinable)<br />
Taxonomic notes. Treated as valid species by Prok<strong>of</strong>iev<br />
(2009b: 888).<br />
10.26.5 Paracobitis malapterura (Valenciennes, in Cuvier<br />
& Valenciennes, 1846)<br />
Cobitis malapterura Valenciennes, in Cuvier & Valenciennes,<br />
1846: 88, pl. 523 (type locality: "sent from Syria"; syntypes:<br />
MNHN 3962 [1], B-3070 [1], Bertin & Estève,<br />
1948: 96; Eschmeyer & Fricke, 2010; compound adjective,<br />
-us, -a, -um [as shown in French translation by Valenciennes])<br />
Taxonomic notes. The type locality is usually listed as "Syria".<br />
But Valenciennes explicitly mentioned that "Mr Aucher<br />
Eloy (1792–1838) has sent, from Syria, a loach" and not that<br />
the loach was from Syria. Aucher Eloy was a botanist who<br />
travelled throughout most <strong>of</strong> Southwest Asia and sold natural<br />
history specimens. The material was apparently received<br />
in Paris in 1840 (after his death) and his last travels have<br />
been especially in Iran and as far east as Balochistan. He does<br />
not seem to have travelled much in Syria (Aucher Eloy, 1843).<br />
Nalbant & Bianco (1999: 114) listed and figured a specimen<br />
in MNHN (no catalogue number listed) as holotype <strong>of</strong><br />
Cobitis malapterura. It is nowhere stated in the original description<br />
that Valenciennes based his description on a single<br />
specimen (his use <strong>of</strong> singular is because he constantly referred<br />
to the species, not to individuals). Thus there is no<br />
reason not to consider the two specimens MNHN 3962 and<br />
B-3070 as syntypes.<br />
Figure 12 in Nalbant & Bianco (1999: 114) shows both a<br />
photograph <strong>of</strong> the "holotype" and the drawing <strong>of</strong> the same<br />
specimen in Cuvier & Valenciennes (1846: pl. 523). They<br />
disagree in showing the eye more backwards and protruding<br />
over the dorsal head pr<strong>of</strong>ile in the drawing (vs. not in the<br />
photograph), the dorsal-fin origin clearly closer to the base<br />
<strong>of</strong> the caudal fin than to the tip <strong>of</strong> the snout (vs. about halfway),<br />
differences in the shape <strong>of</strong> the adipose crest, and the<br />
shape <strong>of</strong> the belly in the drawing is a free reconstruction.<br />
10.26.6 Paracobitis rhadinaea (Regan, 1906)<br />
Nemachilus rhadinaeus Regan, 1906a: 8 (type locality: Afghanistan:<br />
affluents <strong>of</strong> Helmand River; syntypes: BMNH<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
100<br />
10.27.1 Paranemachilus genilepis, ASIZB 60382, 79.0 mm SL,<br />
holotype; China: Guangxi: Changping. (Photograph by Zhao Ya-<br />
Hui; caudal peduncle retouched by author).<br />
1905.11.29.28–29 [2], ZSI F 1240/1 [1], Menon &<br />
Yazdani, 1968: 120; adjective, -us, -a, -um)<br />
Nemachilus macmahoni Chaudhuri, 1910b: 341 (type locality:<br />
Afghanistan: affluents <strong>of</strong> Helmand River; holotype:<br />
ZSI F 1222/1, Menon & Yazdani, 1968: 120; noun in<br />
genitive, indeclinable)<br />
Taxonomic notes. Both N. rhadineus and N. macmahoni<br />
have been collected by the Seistan Arbitration Commission<br />
and have no other locality information besides "affluents <strong>of</strong><br />
the Helmand". All headwaters <strong>of</strong> Helmand River seem to be<br />
in Afghanistan, although the BMNH register indicates the<br />
locality <strong>of</strong> BMNH 1905.11.29.28–29 as Iran.<br />
10.26.7 Paracobitis smithi (Greenwood, 1976)<br />
Noemacheilus smithi Greenwood, 1976: 130, fig. 1 (type<br />
locality: Iran: Zagros Mountains: Kaaje-Ru [33°05'N<br />
48°36'E] near Baq-e-Loveh Oasis; holotype: BMNH<br />
1976.6.28.1; noun in genitive, indeclinable)<br />
Taxonomic notes. I tentatively follow Nalbant & Bianco<br />
(1999: 115) who placed this species in Paracobitis, but without<br />
discussion.<br />
10.26.8 Paracobitis vignai Nalbant & Bianco, 1999<br />
Paracobitis vignai Nalbant & Bianco, 1999: 115, figs. 14<br />
(type locality: Iran: Seistan: Nahr-Taheri, Zabol<br />
[31°01'48"N 61°29'42"E]; holotype: IZA 7838; noun in<br />
genitive, indeclinable)<br />
10.27 Paranemachilus Zhu, 1983<br />
Paranemachilus Zhu, 1983: 311 (type species: Paranemachilus<br />
genilepis Zhu, 1983: 311, by original designation).<br />
Gender masculine.<br />
Nomenclatural notes. Contrary to statement in Romero et<br />
al. (2009: 257), the original spelling Paranemachilus must<br />
be retained. The spelling Paranemachilus is used five times<br />
in the paper and this cannot be called a lapsus or an inadvertent<br />
error in the sense <strong>of</strong> Code art. 32.5.1. Also, neither<br />
Nemacheilus nor Nemachilus is used in the paper and comment<br />
that the spelling Paranemacheilus was intended is not<br />
supported; it is also irrelevant in regard to the Code.<br />
10.27.1 Paranemachilus genilepis Zhu, 1983<br />
Paranemachilus genilepis Zhu, 1983: 311, figs. 1–4 (type<br />
locality: China: Guangxi: Fusui County: subterranean<br />
stream in Changping [22°42'N 107°53'E; Romero et al.,<br />
2009: 257]; holotype: ASIZB 60382 [ex 790002]; compound<br />
noun, indeclinable)
10.28.2 Paraschistura bampurensis, CMK 21897, 42.6 mm SL;<br />
Iran: Hormuzgan.<br />
10.28 Paraschistura Prok<strong>of</strong>iev, 2009<br />
Paraschistura Prok<strong>of</strong>iev, 2009b: 891 (type species: Nemachilus<br />
sargadensis Nikolski, 1900: 415, by original designation).<br />
Gender feminine.<br />
10.28.1 Paraschistura alepidota (Mirza & Banarescu, in<br />
Mirza, Banarescu & Nalbant, 1970)<br />
Noemacheilus rupecola alepidotus Mirza & Bănărescu, in<br />
Mirza, Bănărescu & Nalbant, 1970: 55, fig. 10 (type locality:<br />
Pakistan: Madyan River, Swat drainage [Madyan<br />
town: 35°08'25"N 72°32'14"E]; holotype: ISBB 1356;<br />
adjective, -us, -a, -um)<br />
10.28.2 Paraschistura bampurenris (Nikolski, 1900)<br />
Nemacheilus bampurensis Nikolski, 1900: 414 (type locality:<br />
Southeastern Iran: Sistan and Baluchestan Province:<br />
Kjuagun and Kashkin, near Bazman [village, maybe at<br />
27°39'30"N 59°58'30"E, near volcano Kuh-e Bazman,<br />
28°07'N 60°00'E], Bampur drainage, Jaz Murian endorheic<br />
basin; syntypes: ZISP 11698 [6], 11699 [6]; adjective,<br />
-is, -is, -e)<br />
10.28.3 Paraschistura chrysicristinae (Nalbant, 1998)<br />
Schistura chrysicristinae Nalbant, 1998: 372, fig. 1 (type<br />
locality: Turkey: Batman River (tributary <strong>of</strong> upper Tigris)<br />
at Catalköprü, about 18 km east <strong>of</strong> Silvan; holotype:<br />
ISBB uncat.; noun in genitive, indeclinable)<br />
Taxonomic notes. Generic placement based on geographic<br />
location and advice from J. Freyh<strong>of</strong> (pers. comm.).<br />
10.28.4 Paraschistura kessleri (Günther, 1889)<br />
Nemachilus kessleri Günther, 1889b: 109 (type locality:<br />
Afghanistan: Nushki [in fact Pakistan: Balochistan;<br />
29°33'N 66°01'E, Pishin Lora endorheic basin]; syntypes<br />
[8]: BMNH 1886.9.21.177–180 [4], ZSI 11487–11490<br />
[3]; noun in genitive, indeclinable)<br />
10.28.5 Paraschistura lepidocaulis (Mirza & Nalbant, in<br />
Mirza, Nalbant & Banarescu, 1981)<br />
Schistura kessleri lepidocaulis Mirza & Nalbant, in Mirza,<br />
Nalbant & Bănărescu, 1981: 110, figs. 9–13 (type locality:<br />
Pakistan: Kurram Tribal Agency: Parachinar<br />
[33°53'57"N 70°06'03"E], Kurram River drainage, a right<br />
tributary <strong>of</strong> Indus; holotype: ISBB 3366; compound noun,<br />
indeclinable)<br />
Taxonomic notes. Usually listed as a synonym or subspecies<br />
<strong>of</strong> S. kessleri, which geographical distance and isolation<br />
in widely disjunct basins makes unlikely.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
101<br />
10.29.2 Petruichthys brevis, NRM 28473, 28.6 mm SL (male,<br />
above) and 32.1 mm SL (female, below); Myanmar: Inlé Lake.<br />
10.28.6 Paraschistura lindbergi (Banarescu & Mirza,<br />
1965)<br />
Noemacheilus lindbergi Bănărescu & Mirza, 1965: 265,<br />
figs. 1–4 (type locality: Afghanistan: Siaw, between Farah<br />
[32°20'37"N 62°07'10"E] and Dilaram [Delaram;<br />
32°09'50"N 63°25'51"E], at 780 m, in a rivulet in Farah<br />
Rud drainage, Helmand basin; holotype: HUJ 1572 [lost,<br />
Golani, 2006: 23]; noun in genitive, indeclinable)<br />
Nemacheilus lindbergi haarlovi Bănărescu & Nalbant,<br />
1966a: 176, figs. 14–15, pl. 21 fig. 6 (type locality: Afghanistan:<br />
Kandahar Province: Pirzada [Khugiana;<br />
31°38'18"N 65°07'30"E], West <strong>of</strong> Kandahar, Helmand<br />
basin; holotype ZMUC P 27111 [ex 2811]; noun in genitive,<br />
indeclinable)<br />
Taxonomic notes. Nemacheilus linbergi haarlovi is sometimes<br />
listed as a synonym or subspecies <strong>of</strong> S. kessleri, which<br />
geographical distance and isolation in widely disjunct endorheic<br />
basins makes very unlikely.<br />
10.28.7 Paraschistura microlabra (Mirza & Nalbant, in<br />
Mirza, Nalbant & Banarescu, 1981)<br />
Schistura microlabra Mirza & Nalbant, in Mirza, Nalbant<br />
& Bănărescu, 1981: 126, figs. 56–60 (type locality: Pakistan:<br />
Ali Masjid, Khyber Pass, Kabul River drainage;<br />
holotype: ISBB 3869; compound noun, indeclinable)<br />
10.28.8 Paraschistura naseeri (Ahmad & Mirza, 1963)<br />
Noemacheilus punjabensis naseeri Ahmad & Mirza, 1963:<br />
78, figs. 3–4 (type locality: Pakistan: Swat State: Madyan<br />
stream at Madyan [35°08'25"N 72°32'14"E]; syntypes<br />
[5]: LU; noun in genitive, indeclinable)<br />
10.28.9 Paraschistura nielseni (Nalbant & Bianco, 1999)<br />
Schistura nielseni Nalbant & Bianco, 1999: 119, fig. 22 (type<br />
locality: Iran: Shah Bazar; holotype: ZMC P.27109; noun<br />
in genitive, indeclinable)<br />
Taxonomic notes. Generic placement follows J. Freyh<strong>of</strong><br />
(pers. comm.).<br />
10.28.10 Paraschistura pakistanica (Mirza & Banarescu,<br />
in Mirza, Banarescu & Nalbant, 1969)<br />
Noemacheilus pakistanicus Mirza & Bănărescu, in Mirza,<br />
Bănărescu & Nalbant, 1969: 87, pl. 1 figs. 1–3 (type local-
ity: Pakistan: Hindubagh, Zhob River drainage; holotype:<br />
ISBB 2005; adjective, -us, -a, -um)<br />
10.28.11 Paraschistura prashari (Hora, 1933)<br />
Nemachilus prashari Hora, 1933: 189, pl. 5 figs. 1–2 (type<br />
locality: Pakistan: Northwest Frontier Province [Khyber<br />
Pakhtunkhwa Prov.]: spring 300 yards north <strong>of</strong> Kohat<br />
City; holotype: ZSI F 10719/1; noun in genitive, indeclinable)<br />
10.28.12 Paraschistura punjabensis (Hora, 1923)<br />
Nemachilus punjabensis Hora, 1923: 384, figs. 2–3 (type<br />
locality: Pakistan: Punjab Province: Jhelum District: Pind<br />
Dadan Khan Tehsil [district; Pind Dadan Khan town:<br />
32°35'11"N 73°02'42"E]; syntypes: ZSI F10348/1 [6],<br />
Eschmeyer & Fricke, 2010; adjective, -is, -is, -e)<br />
? Noemacheilus sargadensis paludani Bănărescu & Nalbant,<br />
1966a: 167, figs. 9–10, pl. 21 figs. 1–2 (type locality:<br />
Northeastern Afghanistan: small tributary <strong>of</strong> Pech River<br />
[Kabul River drainage] at Gusalek; holotype: ZMUC<br />
P 27101 [ex 2801]; noun in genitive, indeclinable)<br />
Noemacheilus alepidotus nalbanti Bănărescu & Mirza, 1972:<br />
121, pl. 1 figs. 1–2 (type locality: Pakistan: Rawlakot,<br />
Azad Kashmir, Jhelum River drainage; holotype: ISBB<br />
2490; noun in genitive, indeclinable)<br />
Taxonomic notes. Synonymy follows Kullander et al. (1999:<br />
137).<br />
Nomenclatural notes. Menon & Yazdani (1968: 124) listed<br />
6 syntypes <strong>of</strong> N. punjabensis (ZSI F 10348) collected on<br />
16 July 1922 by Hora in 'stream below Watta Hills, salt range,<br />
Punjab'. Although this locality data is not in agreement with<br />
Hora's original data, they do not provide explanation.<br />
10.28.13 Paraschistura sargadensis (Nikolski, 1900)<br />
Nemachilus sargadensis Nikolski, 1900: 415 (type locality:<br />
Persia: Sija-Rischan in Sargado [according to Berg, 1949:<br />
876: Iran: Sargad Country in Kirman, near Kuh-i-tuftan<br />
volcano [Taftan, 28°36'N 61°8'E; Sarhad Plateau]; syntypes:<br />
ZISP 11700 [31], Eschmeyer & Fricke, 2010; adjective,<br />
-is, -is, -e)<br />
? Nemachilus turcmenicus Berg, 1932a: 149, fig. 1 (type<br />
locality: Turkmenistan: creek near railway station Gjaurs<br />
[Gevers, Gäwers; 37°47'46"N 58°43'43"E] [Kelte-China<br />
stream, Cherokh, Eschmeyer & Fricke, 2010 [Kelte-<br />
Çynar]]; syntypes: ZISP 11064 [3]; adjective, -us, -a, -um)<br />
Taxonomic notes. Geography suggests that N. turcmenicus<br />
is unlikely to be a synonym <strong>of</strong> S. sargadensis.<br />
10.28.14 Paraschistura turcomana (Nikolski, 1947)<br />
Nemachilus kessleri turcomanus Nikolski, 1947: 32, fig. 3<br />
(type locality: Turkmenistan: Murgab River drainage:<br />
Mary Province: Kushka River near Kushka city [35°16'N<br />
62°20'E] / Kushka River near Margunovskyi [Merv,<br />
Margiana, Mary, 37°35'45"N 61°49'30"E] [Murgab River:<br />
endorheic basin earlier connected with Amu Darya];<br />
syntypes: ZMMU P-5734 [3], P-5735 [1], Vasilieva et<br />
al., in Pavlinov & Borissenko, 2001: 27; adjective, -us,<br />
-a, -um)<br />
Taxonomic notes. Usually listed as a subspecies or synonym<br />
<strong>of</strong> S. kessleri, which geographical distance and isolation in<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
102<br />
widely disjunct endorheic basins, on either side <strong>of</strong> the Hindu<br />
Kush range, make unlikely.<br />
10.29 Petruichthys Menon, 1987<br />
Petruichthys Menon, 1987: 181 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemachilus brevis<br />
Boulenger, 1893: 203, by original designation). Gender<br />
masculine.<br />
Taxonomic notes. See comments under Yunnanilus.<br />
Species inquirenda et incertae sedis<br />
10.29.1 Petruichthys salmonides (Chaudhuri, 1911)<br />
Nemachilus salmonides Chaudhuri, 1911: 18, pl. 1 figs. 3–<br />
3a (type locality: China: Yunnan: Tengyueh district:<br />
Mongpan [Mengban, 23°07'N 100°23'E, Mekong drainage;<br />
Coggin Brown, 1910]; holotype: ZSI F 4732/1,<br />
Menon & Yazdani, 1968: 124; adjective, indeclinable)<br />
10.29.2 Petruichthys brevis (Boulenger, 1893)<br />
Nemachilus brevis Boulenger, 1893: 203 (type locality: Burma:<br />
Southern Shan States: Inlé Lake, Fort Stedman; lectotype:<br />
BMNH 1893.6.30.83, designated by Kottelat &<br />
Chu, 1988a: 79; adjective, -is, -is, -e)<br />
10.30 Physoschistura Banarescu & Nalbant, in Singh,<br />
Sen, Banarescu & Nalbant, 1982<br />
Physoschistura Bănărescu & Nalbant, in Singh, Sen, Bănărescu<br />
& Nalbant, 1982: 208 (type species: Nemacheilus<br />
brunneanus Annandale, 1918: 45, by original designation).<br />
Gender feminine.<br />
10.30.1 Physoschistura brunneana (Annandale, 1918)<br />
Nemachilus brunneanus Annandale, 1918: 44, pl. 2 fig. 2<br />
(type locality: Burma: Yawnghwe Valley and Inlé Lake;<br />
holotype: ZSI F 9406/1; adjective, -us, -a, -um)<br />
10.30.2 Physoschistura chindwinensis Lokeshwor & Vishwanath,<br />
2012<br />
Physoschistura chindwinensis Lokeshwor & Vishwanath,<br />
2012a: 231, fig. 1 (India: Manipur: Lokchao River at<br />
Moreh [Irrawaddy drainage]; holotype: MUMF 11077)<br />
10.30.3 Physoschistura elongata Sen & Nalbant, in Singh,<br />
Sen, Banarescu & Nalbant, 1982<br />
Physoschistura elongata Sen & Nalbant, in Singh, Sen,<br />
Bănărescu & Nalbant, 1982: 211, figs. 18–21 (type locality:<br />
India: Meghalaya: Barapani, 20 km from Shillong;<br />
holotype: ZSI/ERS 3063; junior secondary homonym <strong>of</strong><br />
Aborichthys elongatus Hora, 1921c: 735 when both<br />
placed in Nemacheilus by Menon, 1987: 131, 184; junior<br />
secondary homonym <strong>of</strong> Nemachilus kungessanus<br />
elongatus Herzenstein, 1888: 44, when placed in Nemacheilus<br />
by Menon, 1987: 131; reinstated according to<br />
Code art. 59.4; adjective, -us, -a, -um)<br />
Noemacheilus barapaniensis Menon, 1987: 131 (replacement<br />
name for Physoschistura elongata Sen & Nalbant,<br />
in Singh, Sen, Bănărescu & Nalbant, 1982: 211; adjective,<br />
-is, -is, -e)
10.30.4 Physoschistura pseudobrunneana Kottelat, 1990<br />
Physoschistura pseudobrunneana Kottelat, 1990a: 81, fig.<br />
49 (type locality: Thailand: Chiang Rai Province: Nam<br />
Mae Lao at km 62 and 65 along road from Chiang Mai to<br />
Chiang Rai; holotype: ZRC 38473 [ex ZSM 27471]; adjective,<br />
-us, -a, -um)<br />
10.30.5 Physoschistura raoi (Hora, 1929)<br />
Nemachilus raoe Hora, 1929: 332, pl. 15 figs. 7–8 (type<br />
locality: Burma: Northern Shan States: Mongyai [22°26'N<br />
98°03'E]; holotype: ZSI F 11062/1, Menon & Yazdani,<br />
1968: 124; incorrect original spelling, must be emended<br />
into raoi, Code art. 31.1.2; noun in genitive, indeclinable)<br />
10.30.6 Physoschistura rivulicola (Hora, 1929)<br />
Nemachilus rivulicola Hora, 1929: 324, pl. 15 figs. 3–4 (type<br />
locality: Burma: Southern Shan States: Yawnghwe Valley<br />
and He-Ho plain; holotype: ZSI F 11060/1; noun in<br />
apposition, indeclinable)<br />
Nomenclatural notes. Words ending in -cola and meaning<br />
'inhabitant <strong>of</strong>' are nouns and indeclinable.<br />
10.30.7 Physoschistura shanensis (Hora, 1929)<br />
Nemachilus shanensis Hora, 1929: 322, fig. 2, pl. 15 figs. 5–<br />
6 (type locality: Burma: Southern Shan States: Thale-ú<br />
stream near Fort Stedman; holotype: ZSI F 11058/1; adjective,<br />
-is, -is, -e)<br />
10.30.8 Physoschistura tuivaiensis Lokeshwor, Vishwanath<br />
& Shanta, 2012<br />
Physoschistura tuivaiensis Lokeshwor, Vishwanath & Shanta,<br />
2012: 6, fig. 1 (type locality: India: Manipur: Churchandpur<br />
District: Tuivai River at Likhailok (24°04'41"N<br />
93°33'67"E, 635 masl), Brahmaputra drainage; holotype:<br />
MUMF 5089; adjective, -is, -is, -e)<br />
10.30.9 Physoschistura yunnaniloides Chen, Kottelat &<br />
Neely, 2011<br />
Physoschistura yunnaniloides Chen, Kottelat & Neely, 2011:<br />
180, fig. 1 (type locality: Myanmar: Sagaing Division:<br />
Kalemyo fish markets [Kalaymio, 23°11'20"N<br />
94°04'00"E; Chindwin drainage]; holotype: CAS 88871;<br />
adjective, indeclinable)<br />
10.31 Protonemacheilus Yang & Chu, 1990<br />
Protonemacheilus Yang & Chu, 1990a: 109 (type species:<br />
Protonemacheilus longipectoralis Yang & Chu, 1990a:<br />
110, by original designation; also spelt Protenemacheilus<br />
p. 109, an obvious inadvertent error, thus incorrect original<br />
spelling [Code art. 32.5.1]). Gender masculine.<br />
Taxonomic notes. Possibly a junior synonym <strong>of</strong> Physoschistura<br />
or a senior synonym <strong>of</strong> Pteronemacheilus.<br />
10.31.1 Protonemacheilus longipectoralis Yang & Chu,<br />
1990<br />
Protonemacheilus longipectoralis Yang & Chu, 1990a: 110,<br />
fig. 1 (type locality: China: Yunnan: Luxi County: Mukang,<br />
Fangmaqiao River, 24°30'N 98°37'E [Dehong Pre-<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
103<br />
fecture; Irrawaddy drainage]; holotype: KIZ 193001724<br />
[8310110]; compound noun, indeclinable)<br />
10.32 Pteronemacheilus Bohlen & Šlechtová, 2011<br />
Pteronemacheilus Bohlen & Šlechtová, 2011a: 5 (type species:<br />
Pteronemacheilus lucidorsum Bohlen & Šlechtová,<br />
2011: 6, by original designation). Gender masculine.<br />
10.32.1 Pteronemacheilus lucidorsum Bohlen & Šlechtová,<br />
2011<br />
Pteronemacheilus lucidorsum Bohlen & Šlechtová, 2011a: 6,<br />
figs. 4–5 (type locality: Myanmar: Shan State: stream<br />
Nam Paw west <strong>of</strong> Hsipaw city; 22°37'37"N 97°17'19"E<br />
[Irrawaddy drainage]; holotype: ZRC 52039; compound<br />
noun, indeclinable)<br />
10.32.2 Pteronemacheilus meridionalis (Zhu, 1982)<br />
Nemachilus meridionalis Zhu, 1982a: 108, fig. 5 (type locality:<br />
China: Yunnan: Mengla County: Menglun; holotype:<br />
NPIB 780503 [780530, p. 111]; adjective, -is, -is, -e)<br />
10.33 Qinghaichthys Zhu, 1981<br />
Qinghaichthys Zhu, 1981: 1063 (subgenus <strong>of</strong> Triplophysa<br />
Rendahl, 1933: 21; type species Nemacheilus alticeps<br />
Herzenstein, 1888: 28, by original designation). Gender<br />
masculine.<br />
Taxonomic notes. See under Triplophysa.<br />
10.33.1 Qinghaichthys alticeps (Herzenstein, 1888)<br />
Nemachilus alticeps Herzenstein, 1888: 28, pl. 6 fig. 3 (type<br />
locality: China: Qinghai: sources in Eastern Zaidam<br />
[Qaidam basin]; lectotype: ZISP 7852 [1 <strong>of</strong> 4], designated<br />
by Prok<strong>of</strong>iev, 2006a: 599; compound adjective, indeclinable)<br />
10.33.2 Qinghaichthys rotundiventris (Wu & Chen, 1979)<br />
Nemacheilus rotundiventris Wu & Chen, 1979: 292, fig. 3<br />
(type locality: China: Qinghai: Jiegu He, tributary <strong>of</strong><br />
upper Jinsha Jiang [Yangtze] in Yushu; syntypes: NPIB<br />
[5]; compound noun, indeclinable)<br />
10.33.3 Qinghaichthys zaidamensis (Kessler, 1876)<br />
Nemachilus zaidamensis Kessler, 1876: 34 (type locality:<br />
China: Qinghai: Zaidam [Tsaidam or Qaidam basin]; syntypes:<br />
ZISP 2487 [2]; adjective, -is, -is, -e)<br />
10.33.4 Qinghaichthys zamegacephalus (Zhao, 1985)<br />
Nemachilus zamegacephalus Zhao, 1985: 53, fig. 1 (type<br />
locality: China: South <strong>of</strong> Xinjiang: Xiaohaizi, a small lake<br />
at Zubashi, 39°30'N 78°40'E, Bachu; syntypes: ASIZB<br />
54884–885 [2], 60189–200 [12]; compound adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Possibly a synonym <strong>of</strong> Q. rotundiventris<br />
according to Prok<strong>of</strong>iev (2010: 894), which geography suggests<br />
is unlikely.
10.30.4 Physoschistura pseudobrunneana, USNM 295772, 26.5<br />
mm SL; Thailand: Mekong drainage, Mae Nam Fang.<br />
10.30.4 Physoschistura pseudobrunneana, CMK 4111, 23.2 mm SL.<br />
10.34 Schistura McClelland, 1838<br />
Schistura McClelland, 1838: 944, 947 (type species: Schistura<br />
rupecula McClelland, 1838: 948, by subsequent<br />
designation by Jordan, 1919: 195; also in M'Clelland,<br />
1839: 306, 439). Gender feminine.<br />
Acoura Swainson, 1839: 310 (subgenus <strong>of</strong> Cobitis Linnaeus,<br />
1758: 303; type species: Cobitis obscura Swaison, 1839:<br />
310, by subsequent designation by Swain, 1883: 281; also<br />
spelt Acourus on p. 190, first reviser [Jordan, 1919: 204]<br />
gave precedence to Acoura; as there is no etymology<br />
cannot be treated as an inadvertent error). Gender feminine.<br />
Acura Agassiz, 1846: 7 (unjustified emendation <strong>of</strong> Acoura<br />
Swainson, 1839: 310). Gender feminine.<br />
? Longischistura Bănărescu & Nalbant, 1995: 444 (type species:<br />
Nemacheilus striatus Day, 1867b: 347, by original<br />
designation). Gender feminine.<br />
Taxonomic notes. The description <strong>of</strong> Longischistura is accompanied<br />
by neither a diagnosis nor a discussion and it is<br />
impossible to know why the mentioned characters would<br />
diagnose a distinct lineage; at best they seem to be autapomorphies.<br />
In addition, there is no clear indication <strong>of</strong> what<br />
material has been examined.<br />
Nomenclatural notes. Two species were originally included<br />
in Schistura: S. rupecula and S. montana. Schistura is a<br />
feminine noun because it ends with the Greek word ορ<br />
(tail), which is feminine. Further, the species name montana<br />
is an adjective with feminine ending and also indicates that<br />
the genus name was meant to be feminine (see also under<br />
S. rupecula).<br />
Species incertae sedis<br />
10.34.1 Schistura sonlaensis (Nguyen, Nguyen & Hoang,<br />
2010)<br />
Oreias sonlaensis Nguyen, Nguyen & Hoang, 2010: 45, fig. 1<br />
(type locality: Vietnam: Son La Province: Son La District:<br />
Chieng Xom commune [about 21°24'N 103°56'E],<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
104<br />
10.31.1 Protonemacheilus longipectoralis, KIZ 8310110, holotype,<br />
46.5 mm SL; China: Yunnan: Irrawaddy drainage. (Photograph<br />
by Chen Xiao-Yong).<br />
10.31.1 Protonemacheilus longipectoralis, KIZ uncat., about 70<br />
mm SL; China: Yunnan: Irrawaddy drainage. (Photograph by Chen<br />
Xiao-Yong).<br />
lake Born Hau [apparently a resurgence], Chieng Xom<br />
commune [about 21°24'N 103°56'E], in Song Da drainage;<br />
holotype: NCNTTSI (?) SL.08.11.001; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. Originally described in Oreias, which in<br />
other papers <strong>of</strong> authors mainly corresponds to Schistura. The<br />
data in the original description does not allow to decide on<br />
generic position. It is not a species <strong>of</strong> Claea (replacement<br />
name for the invalid Oreias).<br />
Species incertae sedis<br />
10.34.2 Schistura trilineata (Nguyen, Nguyen & Hoang,<br />
2010)<br />
Oreias trilineatus Nguyen, Nguyen & Hoang, 2010: 47, fig. 2<br />
(type locality: Vietnam: Son La Province: Son La District:<br />
Chieng Xom commune [about 21°24'N 103°56'E],<br />
lake Born Hau [apparently a resurgence], in Song Da<br />
drainage; holotype: NCNTTSI (?) SL.08.11.011; adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Originally described in Oreias, which in<br />
other papers <strong>of</strong> authors mainly corresponds to Schistura. The<br />
data in the original description does not allow to decide on<br />
generic position. It is not a species <strong>of</strong> Claea (replacement<br />
name for the invalid Oreias).<br />
10.34.3 Schistura acuticephala (Hora, 1929)<br />
Nemachilus acuticephalus Hora, 1929: 328, pl. 14 figs. 5–6<br />
(type locality: Burma: Northern Shan States: Hsipaw<br />
State: Monglong Subdivision: Hwe-gna-sang River, Pazi<br />
Township [Irrawaddy drainage]; holotype: ZSI F 6675/1;<br />
compound adjective, -us, -a, -um)<br />
10.34.4 Schistura afasciata Mirza & Banarescu, in Mirza,<br />
Nalbant & Banarescu, 1981<br />
Schistura afasciata Mirza & Bănărescu, in Mirza, Nalbant<br />
& Bănărescu, 1981: 119, figs. 35–39 (type locality: Northern<br />
Pakistan: Havelian on Dor River, left tributary <strong>of</strong><br />
upper Indus; holotype: ISBB 3368; adjective, -us, -a,<br />
-um)
10.32.2 Pteronemacheilus meridionalis, CMK 14321, 40.4 mm<br />
SL (male, above) and 39.6 mm SL (female, below); Laos: Mekong<br />
drainage: Nam Youan.<br />
10.33.1 Qinghaichthys alticeps, ZISP 7255/15, about 54 mm SL;<br />
China: Qinghai: Lake Qinhai Hu. (From Herzenstein, 1888: pl. 6<br />
fig. 3).<br />
Taxonomic notes. Menon (1987: 68) treated S. afasciata as<br />
synonym <strong>of</strong> S. arifi, without discussion. I tentatively retain<br />
it as valid.<br />
10.34.5 Schistura aizawlensis Lalramliana, 2012<br />
Schistura aizawlensis Lalramliana, 2012: 98, figs. 1–2 (type<br />
locality: India: Mizoram: Muthi River, a tributary <strong>of</strong> Tuirial<br />
River in vicinity <strong>of</strong> Zemabawk, Aizawl, 23°44'54"N 92°<br />
45'27"E; holotype: PUCMF 11026; adjective, is, -is, -e)<br />
10.34.6 Schistura alta Nalbant & Bianco, 1999<br />
Schistura alta Nalbant & Bianco, 1999: 118, fig. 21 (type<br />
locality: Afghanistan: Kajkai [?Kajaki; 32°19'21"N<br />
65°07'07"E], Helmand drainage, northeast <strong>of</strong> Girisk [Girishk,<br />
Gereshk, Geresk, 31°49'N 64°33'E]; holotype: ZMC<br />
P.27115; adjective, -us, -a, -um)<br />
10.34.7 Schistura alticrista Kottelat, 1990<br />
Schistura alticrista Kottelat, 1990a: 98, fig. 68 (type locality:<br />
Thailand: Salween basin: Mae Hong Son Province:<br />
Mae La Ka, Huei Nong Heng, Tambon Muang Bon,<br />
Amphoe Khum Yuan; holotype: ZRC 38469 [ex ZSM<br />
27472]; compound noun, indeclinable)<br />
10.34.8 Schistura altipedunculata (Banarescu & Nalbant,<br />
1968)<br />
Noemacheilus altipedunculatus Bănărescu & Nalbant, 1968:<br />
337, figs. 7–8, pl. 1 fig. 10 (type locality: India: Karnataka:<br />
North Canara [Uttara Kannada]: Mandurli, Kati River,<br />
15°05'N 74°25'E; holotype: ZMH H 3618, Wilkens,<br />
1977: 158; adjective, -us, -a, -um)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
105<br />
10.34.7 Schistura alticrista, CMK 14730, 46.1 mm SL; Thailand:<br />
Salween drainage: Mae Hong Son.<br />
10.34.15 Schistura atra, CMK 17057, 50.8 mm SL; Laos: Mekong<br />
drainage: Nam Theun.<br />
10.34.19 Schistura cf. balteata, CMK 16326, 43.2 mm SL; Myanmar:<br />
aquarium-fish trade.<br />
10.34.9 Schistura amplizona Kottelat, 2000<br />
Schistura amplizona Kottelat, 2000: 54, fig. 24 (type locality:<br />
Laos: Louangnamtha Province: Nam Tha at Ban Finho,<br />
about 14 km north <strong>of</strong> Luang Nam Tha, Mekong drainage;<br />
21°04'44"N 101°24'09"E; holotype: ZRC 45321;<br />
compound noun, indeclinable)<br />
10.34.10 Schistura anambarensis (Mirza & Banarescu,<br />
in Mirza, Banarescu & Nalbant, 1970)<br />
Noemacheilus anambarensis Mirza & Bănărescu, in Mirza,<br />
Bănărescu & Nalbant, 1970: 51, figs. 4–6 (type locality:<br />
Pakistan: Balochistan: Anambar River at Loralai [30°22'N<br />
68°36'E; Indus drainage]; holotype: ISBB 1364; adjective,<br />
-is, -is, -e)<br />
10.34.11 Schistura antennata Freyh<strong>of</strong> & Serov, 2001<br />
Schistura antennata Freyh<strong>of</strong> & Serov, 2001: 140, figs. 8–9<br />
(type locality: Vietnam: Ha Tinh Province: stream at Son<br />
Kim; 18°24.25'N 105°11.10'E; holotype: ZFMK 27934;<br />
adjective, -us, -a, -um)<br />
10.34.12 Schistura aramis Kottelat, 2000<br />
Schistura aramis Kottelat, 2000: 54, fig. 25 (type locality:<br />
Laos: Phongsali Province: Houay Chik, about 2 km east<br />
<strong>of</strong> Muang Mai, Mekong drainage; 21°10'26"N 102°44'<br />
06"E; holotype: ZRC 45323; noun in apposition, indeclinable)<br />
10.34.13 Schistura arifi Mirza & Banarescu, in Mirza,<br />
Nalbant & Banarescu, 1981<br />
Schistura arifi Mirza & Bănărescu, in Mirza, Nalbant &<br />
Bănărescu, 1981: 113, figs. 16–20 (type locality: Paki-
stan: Balochistan: Zhob City [formerly Fort Sandeman;<br />
31°21'N 69°27'E] on Zhob River; holotype: ISBB 3775;<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. Menon (1987: 68) listed S. curtistigma,<br />
S. macrolepis and S. afasciata as synonyms <strong>of</strong> S. arifi, without<br />
discussion. This is not followed. See under the respective<br />
species.<br />
10.34.14 Schistura athos Kottelat, 2000<br />
Schistura athos Kottelat, 2000: 55, fig. 26 (type locality:<br />
Laos: Louangphabang Province: Houay Houn, about<br />
3 km upstream <strong>of</strong> Ban Houay Lek, in gorges, Mekong<br />
drainage; approx. 20°32'32"N 102°40'51"E; holotype:<br />
ZRC 45325; noun in apposition, indeclinable)<br />
Oreias punctatus Nguyen, 2005: 570, fig. 15 (type locality:<br />
Vietnam: Dien Bien Province: Dien Bien district: Nua<br />
Ngam, Nam Ngam stream, Mekong drainage; holotype:<br />
NCNTTSI; adjective, -us, -a, -um)<br />
10.34.15 Schistura atra Kottelat, 1998<br />
Schistura atra Kottelat, 1998a: 69, fig. 107 (type locality:<br />
Laos: Khammouan Province: Nam Theun basin: Upper<br />
Nam Theun at 17°59'31"N 105°27'49"E, Mekong drainage;<br />
holotype: ZRC 41792; adjective, ater, atra, atrum)<br />
10.34.16 Schistura aurantiaca Plongsesthee, Page &<br />
Beamish, 2011<br />
Schistura aurantiaca Plongsesthee, Page & Beamish, 2011:<br />
170, figs. 1–2 (type locality: Thailand: Kanchanaburi<br />
Province: Thong Pha Phum, Mae Khlong basin, Kwai<br />
Noi River system, Khayeng River at bridge on Route<br />
3272, 14°39'35"N 98°32'01"E; holotype: UF 178532;<br />
adjective, -us, -a, -um)<br />
10.34.17 Schistura bachmaensis Freyh<strong>of</strong> & Serov, 2001<br />
Schistura bachmaensis Freyh<strong>of</strong> & Serov, 2001: 143, figs. 11–<br />
12 (type locality: Vietnam: Thua Thien Hue Province:<br />
stream Suoi Voi at Thua Luu, 55 km south <strong>of</strong> Hue;<br />
16°14.58'N 107°59.39'E; holotype: ZFMK 27942; adjective,<br />
-is, -is, -e)<br />
10.34.18 Schistura bairdi Kottelat, 2000<br />
Schistura bairdi Kottelat, 2000: 55, fig. 27 (type locality:<br />
Laos: Champasak Province: Mekong mainstream at Ban<br />
Hang Khone below Khone Falls; holotype: ZRC 45327;<br />
noun in genitive, indeclinable)<br />
10.34.19 Schistura balteata (Rendahl, 1948)<br />
Nemacheilus balteatus Rendahl, 1948: 42, figs. 20–22 (type<br />
locality: Burma: Tenasserim: Malwedaung; holotype:<br />
NRM 14741; adjective, -us, -a, -um)<br />
10.34.20 Schistura baluchiorum (Zugmayer, 1912)<br />
Nemachilus baluchiorum Zugmayer, 1912: 599 (type locality:<br />
Pakistan: Balochistan: Panjgur [26°58'00"N<br />
64°05'60"E]; syntypes [13]: ZSM [12, lost; pers. obs.],<br />
NMW 19851 [1], Neumann, 2006: 267; noun in genitive,<br />
indeclinable)<br />
Taxonomic notes. Treated as synonym <strong>of</strong> Paraschistura<br />
bampurensis by Nalbant & Bianco (1999: 118), without ex-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
106<br />
planation. Retained in Schistura following Prok<strong>of</strong>iev (2009b:<br />
892).<br />
10.34.21 Schistura bannaensis Chen, Yang & Qi, 2005<br />
Schistura bannaensis Chen, Yang & Qi, 2005: 147, fig. 1<br />
(type locality: China: Yunnan: Xishuangbanna: Mengla<br />
County: Nanla River drainage; holotype: KIZ 200107;<br />
adjective, -is, -is, -e)<br />
10.34.22 Schistura beavani (Günther, 1868)<br />
Nemachilus beavani Günther, 1868: 350 (type locality: India:<br />
Kossye River; holotype: BMNH 1867.5.12.35; noun<br />
in genitive, indeclinable)<br />
10.34.23 Schistura bella Kottelat, 1990<br />
Schistura bella Kottelat, 1990a: 104, fig. 72 (type locality:<br />
Thailand: Chiang Mai Province: Mae Nam Fang, 35 km<br />
south <strong>of</strong> Fang on road to Chiang Mai, Mekong drainage;<br />
holotype: ZRC 38465 [ex ZSM 27473]; adjective, -us,<br />
-a, -um)<br />
10.34.24 Schistura bhimachari (Hora, 1937)<br />
Nemachilus bhimachari Hora, 1937: 13, fig. 5 (type locality:<br />
India: Karnataka: Tunga River at Shimoga<br />
[13°55'00"N 75°34'00"E]; holotype: ZSI F 12178/1,<br />
Menon & Yazdani, 1968: 122; noun in genitive, indeclinable)<br />
Taxonomic notes. Treated as synonym <strong>of</strong> S. striata by Menon<br />
(1987: 113), which I do not follow. The figure <strong>of</strong><br />
S. striata in Day (1878a: pl. 153 fig. 8, reproduced by Menon,<br />
1987: pl. 5 fig. 1) shows a fish with much more slender<br />
body, elongated caudal peduncle, slightly emarginate caudal<br />
fin (vs. deeply forked) than the figure <strong>of</strong> S. bhimachari<br />
in Hora (1937: 13).<br />
10.34.25 Schistura bolavenensis Kottelat, 2000<br />
Schistura bolavenensis Kottelat, 2000: 56, fig. 28 (type locality:<br />
Laos: Champasak Province: Huay Makchan-Gnai (Xe<br />
Nam Noy basin), south <strong>of</strong> Ban Taot, at turn<strong>of</strong>f to Xe Nam<br />
Noy Project, on road from Pakse to Attapu, Mekong<br />
drainage; 15°04'14"N 106°32'33"E; holotype: ZRC<br />
45328; adjective, -is, -is, -e)<br />
10.34.26 Schistura breviceps (Smith, 1945)<br />
Noemacheilus breviceps Smith, 1945: 308, fig. 63 (type locality:<br />
Thailand: Chiang Mai Province: Menam Mao, a tributary<br />
<strong>of</strong> Menam Fang, an affluent <strong>of</strong> Mekong; holotype:<br />
USNM 177751; compound adjective, indeclinable)<br />
10.34.27 Schistura bucculenta (Smith, 1945)<br />
Noemacheilus bucculentus Smith, 1945: 326, fig. 74 (type<br />
locality: Laos: Huey Nam Puat at Ban Nam Puat [Mekong<br />
drainage]; holotype: USNM 107942; adjective, -us, -a, -um)<br />
10.34.28 Schistura callichroma (Zhu & Wang, 1985)<br />
Noemacheilus callichromus Zhu & Wang, 1985: 214, figs.<br />
13–17 (type locality: China: Yunnan: Jingdong County:<br />
Babianjiang River [Red River drainage], 24°20'N<br />
100°10'E; holotype: NGI 810233; compound adjective,<br />
-us, -a, -um])
10.34.29 Schistura callidora Bohlen & Šlechtová, 2011<br />
Schistura callidora Bohlen & Šlechtová, 2011a: 2, figs. 1–2<br />
(type locality: Myanmar: Shan State: mouth <strong>of</strong> Nam Paw<br />
at confluence with Myit Nge River at Hsipaw city,<br />
22°37'24"N 97°18'12"E [Irrawaddy drainage]; holotype:<br />
ZRC 52037; compound noun, proposed as a noun in apposition,<br />
indeclinable)<br />
10.34.30 Schistura carbonaria Freyh<strong>of</strong> & Serov, 2001<br />
Schistura carbonaria Freyh<strong>of</strong> & Serov, 2001: 145, figs. 14–<br />
15 (type locality: Vietnam: Thua Thien Hue Province:<br />
stream Khe Vinh An at Nam Dong village about 40 km<br />
south <strong>of</strong> Hue; 16°11.58'N 107°43.76'E; holotype: ZFMK<br />
26593; adjective, -us, -a, -um)<br />
10.34.31 Schistura carletoni (Fowler, 1924)<br />
Nemacheilus carletoni Fowler, 1924: 68, fig. 2 (type locality:<br />
India: Himachal Pradesh: Kooloo Valley [Kullu Valley:<br />
Kullu town: 31°57'36"N 77°06'00"E]; holotype:<br />
MCZ 18748; noun in genitive, indeclinable)<br />
10.34.32 Schistura cataracta Kottelat, 1998<br />
Schistura cataracta Kottelat, 1998a: 70, fig. 109 (type locality:<br />
Laos: Khammouan Province: Nam Theun, waterfall<br />
at 18°01'40"N 104°58'54"E; holotype: ZRC 41793; noun<br />
in apposition, indeclinable)<br />
10.34.33 Schistura caud<strong>of</strong>urca (Mai, 1978)<br />
Barbatula caud<strong>of</strong>urca Mai, 1978: 233, fig. 106 (type locality:<br />
Vietnam: Nam Cong, Song Ma; syntypes: DVZUT;<br />
compound noun, indeclinable)<br />
Noemacheilus laterivittatus Zhu & Wang, 1985: 213, figs. 9–<br />
12 (type locality: China: Yunnan: Jingdong County: Babianjiang<br />
River [Red River drainage], 24°20'N 100° 10'E;<br />
holotype: NGI 810205; adjective, -us, -a, -um)<br />
10.34.34 Schistura chapaensis (Rendahl, 1944)<br />
Nemacheilus chapaensis Rendahl, 1944: 35, fig. 18 (type<br />
locality: Vietnam: Lao Cai Province: Chapa [Sapa] in<br />
Fansipan Range [Red River drainage]; holotype: NHMG;<br />
adjective, -is, -is, -e)<br />
10.34.35 Schistura chindwinica (Tilak & Hussain, 1990)<br />
Nemacheilus chindwinicus Tilak & Husain, 1990: 51, figs. 1–<br />
5 (type locality: India: Manipur: tributary stream <strong>of</strong><br />
Chindwin [erroneous; India: Manipur: Brahmaputra basin:<br />
Barak drainage, Lankha stream, a tributary <strong>of</strong> Irang<br />
River; Vishwanath & Nebeshwar, 2004: 329]; holotype:<br />
ZSI/NRS F 613; adjective, -us, -a, -um)<br />
10.34.36 Schistura cincticauda (Blyth, 1860)<br />
Cobitis cincticauda Blyth, 1860: 172 (type locality: Thailand:<br />
Tak Province: Amphoe Tha Song Yang, Huei<br />
Jawang, Salween drainage [original type locality: Burma:<br />
Tenasserim provinces (Pegu according to Day, 1870a:<br />
553)]; neotype: ZSM 27474, designated by Kottelat,<br />
1990a: 118; compound noun, indeclinable)<br />
10.34.37 Schistura clatrata Kottelat, 2000<br />
Schistura clatrata Kottelat, 2000: 56, fig. 29 (type locality:<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
107<br />
Laos: Xekong Province: Houay Cha Ngao, an east side<br />
tributary <strong>of</strong> Xe Kong, entering it about 8 km upriver <strong>of</strong><br />
Muang Kaleum, Mekong drainage; 15°46'08"N 106°<br />
45'54"E; holotype: ZRC 45330; adjective, -us, -a, -um)<br />
10.34.38 Schistura conirostris (Zhu, 1982)<br />
Nemachilus conirostris Zhu, 1982a: 104, fig. 1 (type locality:<br />
China: Yunnan: Jinghong County, Lancang-jiang drainage<br />
[Mekong]; holotype: NPIB 780464; compound noun,<br />
indeclinable)<br />
10.34.39 Schistura coruscans Kottelat, 2000<br />
Schistura coruscans Kottelat, 2000: 57, fig. 30 (type locality:<br />
Laos: Saisomboun Special Zone: Houay Sala Yai, a<br />
tributary <strong>of</strong> Nam San; 18°35'17"N 103°05'00"E, Mekong<br />
drainage; holotype: ZRC 45332; participle, indeclinable,<br />
proposed as a noun in apposition)<br />
10.34.40 Schistura crabro Kottelat, 2000<br />
Schistura crabro Kottelat, 2000: 57, fig. 31 (type locality: Laos:<br />
Bolikhamsai Province: Nam Ngiap, Mekong drainage;<br />
holotype: ZRC 45334; noun in apposition, indeclinable)<br />
10.34.41 Schistura crypt<strong>of</strong>asciata Chen, Kong & Yang,<br />
2005<br />
Schistura crypt<strong>of</strong>asciata Chen, Kong & Yang, 2005: 28, fig. 2<br />
(type locality: China: Yunnan: Lincang prefecture:<br />
Yongde County: Great Snow Mountain township: Nanting<br />
River (23°58"N 99°39'E), a tributary <strong>of</strong> Salween;<br />
holotype: KIZ 20026453; adjective, -us, -a, -um)<br />
10.34.42 Schistura curtistigma Mirza & Nalbant, in Mirza,<br />
Nalbant & Banarescu, 1981<br />
Schistura curtistigma Mirza & Nalbant, in Mirza, Nalbant<br />
& Bănărescu, 1981: 114, figs. 21–25 (type locality: Pakistan:<br />
North-West Frontier Province: Bannu<br />
[32°59'20"N 70°36'20"E] on Kurram River, a right-side<br />
tributary <strong>of</strong> Upper Indus; holotype: ISBB 3776; compound<br />
noun, indeclinable)<br />
Taxonomic notes. Menon (1987: 68, 74) treated this species<br />
as synonym <strong>of</strong> S. arifi except for the Ravi River record<br />
<strong>of</strong> Mirza et al. (1981: 114). Mirza et al. gave neither a figure<br />
nor data for the Ravi sample; Menon listed it in the synonymy<br />
<strong>of</strong> S. horai (p. 74), but without explanation. The reported<br />
differences between S. arifi and S. curtistigma indeed are<br />
small, and inconclusive. I tentatively retain the species as<br />
valid.<br />
10.34.43 Schistura dalatensis Freyh<strong>of</strong> & Serov, 2001<br />
Schistura dalatensis Freyh<strong>of</strong> & Serov, 2001: 150, figs. 19–<br />
20 (type locality: Vietnam: Lam Dong Province: Dai Tan<br />
River about 30 km south <strong>of</strong> Da Lat; 11°46.64'N 108°<br />
19.28'E; holotype: ZFMK 23906; adjective, -is, -is, -e)<br />
10.34.44 Schistura daubentoni Kottelat, 1990<br />
Schistura daubentoni Kottelat, 1990a: 118, fig. 85 (type<br />
locality: Cambodia: bridge on Prek Preas at km 369 on<br />
road from Saigon to Stung Treng, near Ph Khsang Hong,<br />
Mekong drainage, 13°09'N 106°09'E; holotype: MNHN<br />
1988-88; noun in genitive, indeclinable)
10.34.46 Schistura defectiva, CMK 22686, 40.9 mm SL; Laos:<br />
Mekong drainage: Nam Ngum.<br />
10.34.47 Schistura denisoni, CMK 5428, 37.6 mm SL; India:<br />
Karnataka: Gangavaly.<br />
10.34.45 Schistura deansmarti Vidthayanon & Kottelat,<br />
2003<br />
Schistura deansmarti Vidthayanon & Kottelat, 2003: 168,<br />
fig. 6 (type locality: Thailand: Phitsanulok Province:<br />
Thung Salaeng Luang National Park: Tham Phra Sai<br />
Ngam cave, about 200 m from entrance; 16°37'23"N<br />
100°39'47"E; holotype: NIFI 3152; noun in genitive, indeclinable)<br />
10.34.46 Schistura defectiva Kottelat, 2000<br />
Schistura defectiva Kottelat, 2000: 58, fig. 32 (type locality:<br />
Laos: Xiangkhouang Province: Nam Ngum at Ban<br />
Phianglouang, Mekong drainage; 19°31'21"N<br />
103°03'58"E; holotype: ZRC 45335; adjective, -us, -a,<br />
-um)<br />
10.34.47 Schistura denisoni (Day, 1867)<br />
Nemacheilus denisoni Day, 1867a: 287 (type locality: India:<br />
Bowany River [Bhavani], along the base <strong>of</strong> Nilgiris<br />
[p. 282]; syntypes: ZSI A.2682 [1, lost], A.962 [1, lost],<br />
BMNH 1867.7.24.29 [1], NMW 48406 [1], ZMB 10800<br />
[1], RMNH 2661 [1], ZISP 8283 [5], Whitehead & Talwar,<br />
1976: 156; noun in genitive, indeclinable)<br />
Nemacheilus chryseus Day, 1873: 529 (type locality: India:<br />
Bowany River [Bhavani]; syntypes [3]: LU, Whitehead &<br />
Talwar, 1976: 156; adjective, -us, -a, -um)<br />
Nemachilus dayi Hora, 1935a: 57 (new name for Nemachilus<br />
savona [non Hamilton, 1822] <strong>of</strong> Day, 1878a: 619,<br />
pl. 155 fig. 8; type locality: India: West Bengal: hills near<br />
Raniganj [23°37'12"N 87°07'48"E] & Northwestern<br />
Provinces [corresponds to present Uttar Pradesh]; syntypes:<br />
material examined by Day and Hora, details not<br />
given, but includes at least BMNH 1889.2.1.1697–1699<br />
[3]; noun in genitive, indeclinable)<br />
Noemacheilus rendahli Bănărescu & Nalbant, 1968: 336,<br />
figs. 5–6, pl. 1 fig. 8 (type locality: India: Maharasthra:<br />
Nashik District: "Nandur, Madhmeswar" [Nandur-<br />
Madhyameshwar dam on Godavari River, 20°00'51"N<br />
74°07'01"E]; holotype: ZMH H 3621, Wilkens, 1977:<br />
159; noun in genitive, indeclinable)<br />
Noemacheilus denisoni pambaensis Menon, 1987: 103,<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
108<br />
10.34.53 Schistura dorsizona, CMK 22915, 30.5 mm SL; Laos:<br />
Mekong drainage: Nam Theun.<br />
10.34.68 Schistura hoai, CMK 20890, 43.8 mm SL; Laos: Mekong<br />
drainage: Nam Ou.<br />
pl. 10 figs. 5–6 (type locality: India: Kerala: Sabarigiri<br />
[a dam at Moozhiyar on Pamba River, 9°18'30"N<br />
77°04'21"E], southeast <strong>of</strong> Pamba; holotype: ZSI/SRS<br />
F 563; adjective, -is, -is, -e)<br />
Taxonomic notes. Placement <strong>of</strong> N. rendahli in the synonym<br />
<strong>of</strong> S. denisoni follows Menon (1987: 93). This requires confirmation<br />
by the comparison <strong>of</strong> fresh topotypes. My examination<br />
<strong>of</strong> samples from Karnataka and Kerala indicates that<br />
several species are confused under the name S. denisoni. At<br />
least S. mukambbikaensis is a distinct species.<br />
10.34.48 Schistura desmotes (Fowler, 1934)<br />
Nemacheilus desmotes Fowler, 1934: 107, fig. 59 (type locality:<br />
Thailand: Chiang Mai [Chao Phraya drainage]; holotype:<br />
ANSP 60082; noun in apposition, indeclinable)<br />
? Nemacheilus myrmekia Fowler, 1935: 106, fig. 32 (type<br />
locality: Thailand: Hua Hin Province: Keng Sok [about<br />
20 km northwest <strong>of</strong> Hua Hin; near Ban Thung Luang,<br />
12°41'N 99°51'E]; holotype: ANSP 63546; noun in apposition,<br />
indeclinable)<br />
10.34.49 Schistura devdevi (Hora, 1935)<br />
Nemachilus devdevi Hora, 1935a: 54, pl. 3 figs. 5–6 (type<br />
locality: India: small streams below Darjeeling and in<br />
Sikkim; holotype: ZSI F 11752/1; noun in genitive, indeclinable)<br />
10.34.50 Schistura diminuta Ou, Montaña, Winemiller<br />
& Conway, 2011<br />
Schistura diminuta Ou, Montaña, Winemiller & Conway,<br />
2011: 194, fig. 1 (type locality: Cambodia: Stung Treng<br />
Province: Mekong drainage, lower Sekong River in Siem<br />
Pang district, 14º07'11.88"N 106º23'11.36"E; holotype:<br />
ZRC 53105; adjective, -us, -a, -um)<br />
10.34.51 Schistura disparizona Zhou & Kottelat, 2005<br />
Schistura disparizona Zhou & Kottelat, 2005: 17, fig. 1 (type<br />
locality: China: Yunnan: Cangyuan: Nangun River (a<br />
Salween tributary), 0.5 km to Manglai on road from Banhong<br />
to Mangku, 23°16.49'N 99°04.34'E; holotype:<br />
SWFC 00203035; compound noun, indeclinable)
10.34.76 Schistura isostigma, CMK 15651, 25.7 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
10.34.52 Schistura doonensis (Tilak & Husain, 1977)<br />
Noemacheilus doonensis Tilak & Husain, 1977: 133, fig. 1<br />
(type locality: India: Uttar Pradesh: Dehra Dun District:<br />
small stream near village Kandholi, on way to Doonga;<br />
holotype: ZSI/NRS V-1090; adjective, -is, -is, -e)<br />
10.34.53 Schistura dorsizona Kottelat, 1998<br />
Schistura dorsizona Kottelat, 1998a: 73, fig. 112 (type locality:<br />
Laos: Khammouan Province: Xe Bangfai basin: Nam<br />
Phao about 10 km upriver <strong>of</strong> Ban Nape, Mekong drainage;<br />
approximately 18°22'N 105°08'E; holotype: ZRC<br />
41795; compound noun, indeclinable)<br />
10.34.54 Schistura dubia Kottelat, 1990<br />
Schistura dubia Kottelat, 1990a: 123, fig. 89 (type locality:<br />
Thailand: Phrae Province: Nam Mae Kham Mi, near Ban<br />
Mae Krating, road from Nan to Phrae, km 66, Chao<br />
Phraya drainage, 18°22'N 100°25'E; holotype: ZRC<br />
38474 [ex ZSM 27475]; adjective, -us, -a, -um)<br />
10.34.55 Schistura ephelis Kottelat, 2000<br />
Schistura ephelis Kottelat, 2000: 59, fig. 33 (type locality:<br />
Laos: Saisomboun Special Zone: Houay Sala Yai, a tributary<br />
<strong>of</strong> Nam San, Mekong drainage; 18°35'17"N<br />
103°05'00"E; holotype: ZRC 45337; noun in apposition,<br />
indeclinable)<br />
10.34.56 Schistura fasciata Lokeshwor & Viswanath,<br />
2011<br />
Schistura fasciata Lokeshwor & Viswanath, 2011: 1514,<br />
fig. 1 (type locality: India: Manipur: Senapati District:<br />
Barak River at western side <strong>of</strong> Maram Hill, 25°23'<br />
24.66"N 94°04'09.25"E, Brahmaputra drainage; holotype:<br />
MUMF 11010; adjective, -us, -a, -um)<br />
10.34.57 Schistura fascimaculata Mirza & Nalbant, in<br />
Mirza, Nalbant & Banarescu, 1981<br />
Schistura fascimaculata Mirza & Nalbant, in Mirza, Nalbant &<br />
Bănărescu, 1981: 121, figs. 41–45 (type locality: Pakistan:<br />
Khyber-Pakhtunkhwa Province: Kohat District: Hangu<br />
[33°31'48"N 71°03'36"E] on Kohat Toi River, Indus drainage;<br />
holotype: ISBB 3371; adjective, -us, -a, -um)<br />
10.34.58 Schistura fasciolata (Nichols & Pope, 1927)<br />
Homaloptera fasciolata Nichols & Pope, 1927: 339, fig. 11<br />
(type locality: China: Hainan: Nodoa; holotype: AMNH<br />
8365; adjective, -us, -a, -um)<br />
Nemachilus humilis Lin, 1932c: 515 (type locality: China:<br />
Kweichow [Guizhou]: Yunkiang; holotype: BLG K.107;<br />
adjective, -is, -is, -e)<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
109<br />
10.34.72 Schistura imitator, CMK 21686, 45.8 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
10.34.82 Schistura khamtanhi, CMK 21615, 60.1 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
10.34.59 Schistura finis Kottelat, 2000<br />
Schistura finis Kottelat, 2000: 59, fig. 34 (type locality: Laos:<br />
Xiangkhouang Province: Nam Kan, a small creek making<br />
the border between Laos and Vietnam at Ban Xayden,<br />
nam Mo drainage [Song Ca]; 19°28'19"N<br />
104°05'04"E; holotype: ZRC 45339; noun in apposition,<br />
indeclinable)<br />
10.34.60 Schistura fusinotata Kottelat, 2000<br />
Schistura fusinotata Kottelat, 2000: 60, fig. 35 (type locality:<br />
Laos: Xekong Province: Nam Vi at ford downriver<br />
(1.5 km NW) <strong>of</strong> Ban Kasang-Kan, Mekong drainage;<br />
15°17'55"N 106°54'10"E; holotype: ZRC 45341, see also<br />
Kottelat, 2005a: 65; adjective, -us, -a, -um)<br />
10.34.61 Schistura gangetica (Menon, 1987)<br />
Noemacheilus gangeticus Menon, 1987: 83, pl. 12 fig. 6 (type<br />
locality: India: Uttarakhand: Garhwal: Alaknanda River<br />
at Srinagar [30°19'48"N 78°03'36"E]; holotype: ZSI/SRS<br />
F 582; adjective, -us, -a, -um)<br />
10.34.62 Schistura geisleri Kottelat, 1990<br />
Schistura geisleri Kottelat, 1990a: 127, fig. 91 (type locality:<br />
Thailand: Chiang Mai Province: Nam Mae Taeng at<br />
Ban Mae Ta Man, Chao Phraya drainage, 19°12'N<br />
98°53'E; holotype: ZRC 38475 [ex ZSM 27476]; noun<br />
in genitive, indeclinable)<br />
10.34.63 Schistura globiceps Kottelat, 2000<br />
Schistura globiceps Kottelat, 2000: 60, fig. 36 (type locality:<br />
Laos: Louangnamtha Province: unnamed forest creek<br />
tributary to Nam Talan, at about km 60 on road from<br />
Oudomxai to Luang Nam Tha, about 3 km south <strong>of</strong> Ban<br />
Nateuy, Mekong drainage; 20°59'56"N 101°39'47"E;<br />
holotype: ZRC 45343; compound adjective, indeclinable)<br />
10.34.64 ? Schistura hagiangensis (Nguyen, 2005)<br />
Paracobitis hagiangensis Nguyen, 2005: 565, fig. 12 (type<br />
locality: Vietnam: Ha Giang Province: Lo River, near Ha<br />
Giang [Red River drainage]; holotype: NCNTTSI; ad-
jective, -is, -is, -e)<br />
Taxonomic notes. Possibly a junior synonym <strong>of</strong> Schistura<br />
incerta.<br />
10.34.65 Schistura harnaiensis (Mirza & Nalbant, in<br />
Mirza, Banarescu & Nalbant, 1969)<br />
Noemacheilus harnaiensis Mirza & Nalbant, in Mirza, Bănărescu<br />
& Nalbant, 1969: 89, pl. 2 figs. 4–6 (type locality:<br />
Pakistan: Balochistan: Harnai [30°36'0"N 67°33'36"E],<br />
Kaman-Beji River drainage, Indus drainage; holotype:<br />
ISBB 2006; adjective, -is, -is, -e)<br />
10.34.66 Schistura himachalensis (Menon, 1987)<br />
Noemacheilus himachalensis Menon, 1987: 76, pl. 16 fig. 2<br />
(type locality: India: Himachal Pradesh: Kangra District:<br />
Nadaun [Beas drainage; 31°46'48"N 76°21'00"E]; holotype:<br />
ZSI/SRS F 566; adjective, -is, -is, -e)<br />
10.34.67 Schistura hingi (Herre, 1934)<br />
Homaloptera hingi Herre, 1934: 287 (type locality: China:<br />
Hong Kong: mountain torrent at Pok Fu Lam; holotype:<br />
CAS-SU 29086; noun in genitive, indeclinable)<br />
10.34.68 ? Schistura hoai (Nguyen, 2005)<br />
Oreias hoai Nguyen, 2005: 568, fig. 14 (type locality: Vietnam:<br />
Dien Bien Province: Dien Bien district: Nua Ngam,<br />
Nam Ngam stream, Mekong drainage; holotype:<br />
NCNTTSI; noun in genitive, indeclinable)<br />
Taxonomic notes. The original description includes few informative<br />
data and the accompanying figure is <strong>of</strong> very poor<br />
quality. However, recent sampling in Laos very close to the<br />
border with Vietnam and in the catchment <strong>of</strong> the same stream<br />
as the type locality yielded a single nemacheilid species that<br />
could match the figure in the original description, especially<br />
the shape <strong>of</strong> the caudal fin, the bars in front <strong>of</strong> the dorsal fin<br />
much narrower than those behind, and the shape <strong>of</strong> the black<br />
bar at the base <strong>of</strong> the caudal fin. This species had been earlier<br />
identified as S. poculi (e.g. Kottelat, 2001b: 113), from<br />
which it differs by details <strong>of</strong> the colour pattern and the shape<br />
<strong>of</strong> the black bar at the base <strong>of</strong> the caudal fin (compare figures<br />
here).<br />
10.34.69 Schistura horai (Menon, 1951)<br />
Nemachilus horai Menon, 1951: 227, pl. 7 figs. 1–3 (type<br />
locality: India: Himachal Pradesh: Kangra District: Bener<br />
Khand [Baner River, Indus drainage], south <strong>of</strong> Kangra<br />
[32°06'00"N 76°16'12"E]; holotype: ZSI F 637/2,<br />
Menon & Yazdani, 1968: 122; noun in genitive, indeclinable)<br />
10.34.70 Schistura huapingensis (Wu & Wu, 1992)<br />
Nemacheilus obscurus huapingensis Wu & Wu, 1992: 145<br />
(type locality: China: Yunnan: Huaping County, Jinshajiang<br />
drainage [Yangtze]; syntypes: NPIB [3]; adjective,<br />
-is, -is, -e)<br />
10.34.71 Schistura huongensis Freyh<strong>of</strong> & Serov, 2001<br />
Schistura huongensis Freyh<strong>of</strong> & Serov, 2001: 155, figs. 24–<br />
25 (type locality: Vietnam: Thua Thien Hue Province:<br />
stream Khe Vinh An at Nam Dong village, about 40 km<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
110<br />
south <strong>of</strong> Hue; 16°11.58'N 107°43.76; holotype: ZFMK<br />
26810; adjective, -is, -is, -e)<br />
10.34.72 Schistura imitator Kottelat, 2000<br />
Schistura imitator Kottelat, 2000: 61, fig. 37 (type locality:<br />
Laos: Xekong Province: Xe Namnoy, rapids about 1 km<br />
upriver <strong>of</strong> Tad Feak waterfall, Mekong drainage;<br />
15°14'09"N 106°44'55"E; holotype: ZRC 45344; noun<br />
in apposition, indeclinable)<br />
10.34.73 Schistura implicata Kottelat, 2000<br />
Schistura implicata Kottelat, 2000: 62, fig. 38 (type locality:<br />
Laos: Houaphan Province: Nam Poun at confluence<br />
with Nam Xang, downstream <strong>of</strong> Ban San, Nam Ma drainage;<br />
20°19'25"N 104°31'36"E; holotype: ZRC 45346;<br />
adjective, -us, -a, -um)<br />
10.34.74 Schistura incerta (Nichols, 1931)<br />
Barbatula incerta Nichols, 1931b: 458, fig. 2 (type locality:<br />
China: Kwangtung [Guangdong]: Lung T'au Shaan, Tso<br />
Tsz Hoh; holotype: AMNH 9690; adjective, -us, -a, -um)<br />
? Barbatula uniformis Mai, 1978: 235, fig. 107 (type locality:<br />
northern Vietnam; syntypes: DVZUT; adjective, -is,<br />
-is, -e)<br />
10.34.75 Schistura irregularis Kottelat, 2000<br />
Schistura irregularis Kottelat, 2000: 62, fig. 39 (type locality:<br />
Laos: Xiangkhouang Province: Nam Khan at Muang<br />
Hian, Mekong drainage; 20°05'18"N 103°22'09"E; holotype:<br />
ZRC 45348; adjective, -is, -is, -e)<br />
10.34.76 Schistura isostigma Kottelat, 1998<br />
Schistura isostigma Kottelat, 1998a: 76, fig. 114 (type locality:<br />
Laos: Nam Leuk about 1 km downstream <strong>of</strong> confluence<br />
with Nam Ngang [error for Nam Gnong], Mekong<br />
drainage; 18°22'03"N 103°06'03"E; holotype: ZRC<br />
41797; compound noun, proposed as a noun in apposition,<br />
indeclinable)<br />
10.34.77 Schistura jarutanini Kottelat, 1990<br />
Schistura jarutanini Kottelat, 1990b: 49, pl. 1 fig. 1 (type<br />
locality: Thailand: Kanchanaburi Province: Amphoe Sri<br />
Sawat, Tham Ba Dan [cave]; holotype: ZRC 38479 [ex<br />
ZSM 27171]; noun in genitive, indeclinable)<br />
10.34.78 Schistura kangjupkhulensis (Hora, 1921)<br />
Nemachilus kangjupkhulensis Hora, 1921b: 202, pl. 10<br />
figs. 4, 4a (type locality: India: Manipur: Yaribuk [Irrawaddy<br />
drainage]; holotype: ZSI F 10060/1; adjective,<br />
-is, -is, -e)<br />
10.34.79 Schistura kangrae (Menon, 1951)<br />
Nemachilus kangrae Menon, 1951: 229, pl. 7 figs. 4–5 (type<br />
locality: India: Himachal Pradesh: Kangra District:<br />
Baijnath [32°03'N 76°39'E], Indus drainage; holotype:<br />
ZSI F 639/2; noun in genitive, indeclinable)<br />
Taxonomic notes. Treated as synonym <strong>of</strong> S. montana by<br />
Menon (1987: 79), without discussion. With the available<br />
data I cannot follow this synonymy.
10.34.80 Schistura kaysonei Vidthayanon & Jaruthanin,<br />
2002<br />
Schistura kaysonei Vidthayanon & Jaruthanin, 2002: 18,<br />
fig. 1 (type locality: Laos: Khammouan Province: cave<br />
in Phu Tham Nam, Ban Don Yom village in Khammouan<br />
karst, between 18°45' and 18°30'N 104°45' and 105°E<br />
[apparently erroneous, Phu Tham Nam is a hill at<br />
17°29'00"N 104°54'00"E]; holotype: NIFI 3131; name<br />
in genitive, indeclinable)<br />
10.34.81 Schistura kengtungensis (Fowler, 1936)<br />
Nemacheilus kengtungensis Fowler, 1936: 509, fig. 1 (type<br />
locality: Burma: Shan States: Loi Mwe [21°10'N 99°<br />
45'E], Keng Tung, Mekong drainage [see Fowler, 1937:<br />
125]; holotype: ANSP 64157; adjective, -is, -is, -e)<br />
10.34.82 Schistura khamtanhi Kottelat, 2000<br />
Schistura khamtanhi Kottelat, 2000: 63, fig. 40 (type locality:<br />
Laos: Xekong Province: Houay Pao, a west side tributary<br />
<strong>of</strong> Xe Kong, entering it about 16 km upriver <strong>of</strong><br />
Muang Kaleum, Mekong drainage; 15°50'17"N<br />
106°45'40"E; holotype: ZRC 45349, noun in genitive,<br />
indeclinable)<br />
10.34.83 Schistura khugae Vishwanath & Shanta, 2004<br />
Schistura macrocephalus Vishwanath & Shanta, 2004a: 138,<br />
figs. 1–2 (type locality: India: Manipur: Churachandpur<br />
District: Khuga River, a tributary <strong>of</strong> Loktak Lake, Chindwin<br />
drainage; holotype: MUMF 5013; junior primary<br />
homonym <strong>of</strong> Schistura macrocephalus Kottelat, 2000:<br />
65; compound adjective, -us, -a, -um)<br />
Schistura khugae Vishwanath & Shanta, 2004b: 330 (replacement<br />
name for Schistura macrocephalus Vishwanath &<br />
Shanta, 2004a: 138, figs. 1–2; noun in genitive, indeclinable)<br />
10.34.84 Schistura kloetzliae Kottelat, 2000<br />
Schistura kloetzliae Kottelat, 2000: 63, fig. 41 (type locality:<br />
China: Yunnan: Xishuangbanna: Mengla County:<br />
Mengla market, Mekong drainage; holotype: NRM<br />
33199; noun in genitive, indeclinable)<br />
10.34.85 Schistura kodaguensis (Menon, 1987)<br />
Noemacheilus kodaguensis Menon, 1987: 108, pl. 16 fig. 11<br />
(type locality: India: Karnataka: Kotu Hola, 8 km northwest<br />
<strong>of</strong> Merkara; holotype: ZSI/SRS F 565; adjective, -<br />
is, -is, -e)<br />
10.34.86 Schistura kohatensis Mirza & Banarescu, in<br />
Mirza, Nalbant & Banarescu, 1981<br />
Schistura kohatensis Mirza & Bănărescu, in Mirza, Nalbant<br />
& Bănărescu, 1981: 123, figs. 46–50 (type locality: Pakistan:<br />
Kohat District: Hangu [33°31'48"N 71°03'36"E]<br />
on Kohat Toi River, a small right tributary <strong>of</strong> Indus; holotype:<br />
ISBB 3373; adjective, -is, -is, -e)<br />
10.34.87 Schistura kohchangensis (Smith, 1933)<br />
Nemacheilus kohchangensis Smith, 1933: 56, fig. 2, pl. 1<br />
fig. 2 (type locality: Thailand: Trat Province: waterfall<br />
stream on Koh Chang island; holotype: KUMF 169,<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
111<br />
Monkolprasit, 1969: 6; adjective, -is, -is, -e)<br />
Noemacheilus deignani Smith, 1945: 320, fig. 70 (type locality:<br />
Thailand: Chantaburi Province: Kao Sabap, near<br />
Chantabun [Chantaburi]; USNM 107946; name in genitive,<br />
indeclinable)<br />
10.34.88 Schistura koladynensis Lokeshwor & Vishwanath.<br />
2012<br />
Schistura koladynensis Lokeshwor & Vishwanath. 2012b:<br />
140, fig. 1 (type locality: India: Mizoram: Lawntlai District:<br />
Koladyne River at Kolchaw, 22°23'45"N 92°<br />
55'56"E; holotype: MUMF 11048; adjective, is, -is, -e)<br />
10.34.89 Schistura kongphengi Kottelat, 1998<br />
Schistura kongphengi Kottelat, 1998a: 77, fig. 116 (type<br />
locality: Laos: Khammouan Province: upper Nam Theun<br />
at 17°59'31"N 105°27'49"E, Mekong drainage; holotype:<br />
ZRC 41799; noun in genitive, indeclinable)<br />
10.34.90 Schistura kontumensis Freyh<strong>of</strong> & Serov, 2001<br />
Schistura kontumensis Freyh<strong>of</strong> & Serov, 2001: 160, figs. 29–<br />
30 (type locality: Vietnam: Kontum Province: stream Iasia<br />
about 25 km west <strong>of</strong> Kontum; 14°23.63'N 107°<br />
48.64'E; holotype: ZFMK 26687; adjective, -is, -is, -e)<br />
10.34.91 Schistura latidens Kottelat, 2000<br />
Schistura latidens Kottelat, 2000: 64, fig. 42 (type locality:<br />
Laos: Savannakhet Province: Xe Pon between rapids<br />
upstream and downstream <strong>of</strong> Ban Fuang, Mekong drainage;<br />
16°37'06"N 106°33'30"E; holotype: ZRC 45351;<br />
compound noun, indeclinable)<br />
10.34.92 Schistura latifasciata (Zhu & Wang, 1985)<br />
Noemacheilus latifasciatus Zhu & Wang, 1985: 211, figs.<br />
5–6 (type locality: China: Yunnan: Yunxian County: Nanqiao<br />
River, tributary <strong>of</strong> Lancangjiang River [Mekong],<br />
24°N 100°E; holotype: NGI 810367; adjective, -us, -a,<br />
-um)<br />
10.34.93 Schistura leukensis Kottelat, 2000<br />
Schistura leukensis Kottelat, 2000: 64 [not fig. 44 which<br />
shows S. kongphengi] (type locality: Laos: Vientiane<br />
Province: Nam Leuk at dam site, Mekong drainage;<br />
18°26'15"N 102°56'48"E; holotype: ZRC 45353, figured<br />
in Kottelat, 2005a: 65; adjective, -is, -is, -e)<br />
10.34.94 Schistura longa (Zhu, 1982)<br />
Nemachilus longus Zhu, 1982a: 105, fig. 2 (type locality:<br />
China: Yunnan: Lushui County: tributary <strong>of</strong> Nu-jiang<br />
River [Salween]; holotype: NPIB 785032; adjective,<br />
-us, -a, -um)<br />
10.34.95 Schistura machensis (Mirza & Nalbant, in Mirza,<br />
Banarescu & Nalbant, 1970)<br />
Noemacheilus horai machensis Mirza & Nalbant, in Mirza,<br />
Bănărescu & Nalbant, 1970: 54, figs. 7–9 (type locality:<br />
Pakistan: Mach River [Mach, Machh, Muchh; town:<br />
29°52'N 67°20'E], tributary to Bolan River, Indus drainage;<br />
holotype: ISBB 1538; adjective, -is, -is, -e)
10.34.84 Schistura kloetzliae, CMK 14312, 43.9 mm SL; Laos:<br />
Mekong drainage: Nam Youan.<br />
10.34.87 Schistura kohchangensis, CMK 20204, 49.7 mm SL;<br />
Thailand: Chantaburi.<br />
10.34.102 Schistura mahnerti, CMK 14682, 44.8 mm SL; Thailand:<br />
Salween drainage: Mae Nam Moei.<br />
10.34.92 Schistura latifasciata, KIZ uncat., 62.4 mm SL; China:<br />
Yunnan: Mekong drainage: Yangbi.<br />
10.34.96 Schistura macrocephalus Kottelat, 2000<br />
Schistura macrocephalus Kottelat, 2000: 65, fig. 45 (type<br />
locality: Laos: Louangnamtha Province: Nam Youan at<br />
ford south <strong>of</strong> Ban Muang Mon, Mekong drainage; 21°19'<br />
28"N 101°10'19"E; holotype: ZRC 45355; compound<br />
noun, proposed as a noun in apposition, indeclinable)<br />
10.34.97 Schistura macrolepis Mirza & Banarescu, in<br />
Mirza, Nalbant & Banarescu, 1981<br />
Schistura macrolepis Mirza & Bănărescu, in Mirza, Nalbant<br />
& Bănărescu, 1981: 116, figs. 27–31 (type locality:<br />
Pakistan: Dera Ghazi Khan District: Racki Munh, a small<br />
right tributary <strong>of</strong> Indus upwards from confluence with<br />
Sutlej River; holotype: ISBB 3778; compound noun, indeclinable)<br />
Taxonomic notes. Menon (1987: 69) treated this species as<br />
synonym <strong>of</strong> S. arifi except for the Shadiwal paratype, which<br />
he listed in the synonymy <strong>of</strong> S. horai (p. 74), without explanation.<br />
I tentatively retain the species as valid.<br />
10.34.98 Schistura macrotaenia (Yang, 1990)<br />
Nemacheilus macrotaenia Yang, in Chu & Chen, 1990: 36,<br />
fig. 30 (type locality: China: Yunnan: Pingbian County:<br />
Nan-Qi River, 23°00'N 103°18'E [Red River drainage];<br />
holotype: KIZ 8540217; compound noun, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
112<br />
10.34.99‘Schistura’ maculiceps, RMNH 28878, 79.0 mm SL; Indonesia:<br />
Borneo: Kapuas.<br />
10.34.105 Schistura melarancia, ZRC 45357, 40.2 mm SL, holotype;<br />
Laos: Mekong drainage: Nam Ou.<br />
10.34.111 Schistura mukambbikaensis, CMK 5427, 32.6 mm SL;<br />
India: Karnataka: Chandragiri.<br />
10.34.99 'Schistura' maculiceps (Roberts, 1989)<br />
Nemacheilus maculiceps Roberts, 1989: 108, fig. 83 (type<br />
locality: Indonesia: Borneo: Kalimantan Barat: rocky<br />
channel in mainstream <strong>of</strong> Sungai Pinoh, 37 km south <strong>of</strong><br />
Nangapinoh; holotype: MZB 3543; compound adjective,<br />
indeclinable)<br />
Taxonomic notes. Apparently represents a distinct genus.<br />
10.34.100 Schistura maepaiensis Kottelat, 1990<br />
Schistura maepaiensis Kottelat, 1990a: 143, fig. 105 (type<br />
locality: Thailand: Mae Hong Son Province: Nam Mae<br />
Cha at Ban Pha Bong, 19°12'N 97°59'E, 12 km south <strong>of</strong><br />
Mae Hong Son, Salween drainage; holotype: ZRC 38476<br />
[ex ZSM 27479]; adjective, -is, -is, -e)<br />
10.34.101 Schistura magnifluvis Kottelat, 1990<br />
Schistura magnifluvis Kottelat, 1990a: 146, fig. 107 (type<br />
locality: Thailand: Loei Province: Mekong main stream<br />
between Chiang Khan [17°50'N 101°45'E] to 70 km<br />
downstream by road; holotype: CAS 62549; compound<br />
noun, indeclinable)<br />
10.34.102 Schistura mahnerti Kottelat, 1990<br />
Schistura mahnerti Kottelat, 1990a: 151, fig. 110 (type locality:<br />
Thailand: Mae Hong Son Province: 20 km north <strong>of</strong><br />
Mae Sariang, Salween drainage; holotype: ZRC 38477<br />
[ex ZSM 27478]; noun in genitive, indeclinable)<br />
10.34.103 Schistura malaisei Kottelat, 1990<br />
Schistura malaisei Kottelat, 1990a: 154, fig. 113 (type locality:<br />
Burma: Putao, Irrawaddy drainage; holotype: NRM<br />
25083; noun in genitive, indeclinable)
10.34.104 Schistura manipurensis (Chaudhuri, 1912)<br />
Nemachilus manipurensis Chaudhuri, 1912: 443, pl. 40<br />
fig. 4–4b, pl. 41 fig. 1–1b (type locality: India: Manipur<br />
[Irrawaddy drainage]; syntypes: ZSI [3]; adjective, -is,<br />
-is, -e)<br />
10.34.105 Schistura melarancia Kottelat, 2000<br />
Schistura melarancia Kottelat, 2000: 65, fig. 46 (type locality:<br />
Laos: Oudomxai Province: Nam Kouat near Ban Nam<br />
Kouat, a tributary <strong>of</strong> Nam Nga, Mekong drainage;<br />
20°34'35"N 102°07'56"E; holotype: ZRC 45357; name<br />
in genitive, indeclinable)<br />
Oreias lineatus Nguyen, 2005: 567, fig. 13 (type locality:<br />
Vietnam: Dien Bien Province: Dien Bien district: Nua<br />
Ngam, Nam Ngam stream, Mekong drainage; holotype:<br />
NCNTTSI; adjective, -us, -a, -um)<br />
10.34.106 Schistura menanensis (Smith, 1945)<br />
Noemacheilus menanensis Smith, 1945: 310, fig. 65 (type<br />
locality: Thailand: Nan Province: Menam Kon, tributary<br />
<strong>of</strong> Menam Nan, Chao Phraya drainage; holotype: USNM<br />
117753; adjective, -is, -is, -e)<br />
10.34.107 Schistura minuta Vishwanath & Shanta Kumar,<br />
2006<br />
Schistura minutus Vishwanath & Shanta Kumar, 2006: 211,<br />
fig. 1 (type locality: India: Manipur: Tamenglong District:<br />
Iyei River, Noney, a tributary <strong>of</strong> Barak River, Brahmaputra<br />
drainage; holotype: ZSI FF3749; adjective, -us,<br />
-a, -um)<br />
10.34.108 Schistura mobbsi Kottelat & Leisher, 2012<br />
Schistura mobbsi Kottelat & Leisher, 2012: 240, fig. 3 (type<br />
locality: Vietnam: Thai Nguyen Province: Phuong Hoang<br />
Cave, about 32 km from Bac Song (Lang Song Prov.),<br />
probably draining to Cau River, draining to Hai Phong;<br />
21°46'31''N 106°07'10''E; holotype: MHNG 2732.044;<br />
noun in genitive, indeclinable)<br />
10.34.109 Schistura moeiensis Kottelat, 1990<br />
Schistura moeiensis Kottelat, 1990a: 164, fig. 120 (type<br />
locality: Thailand: Tak Province: Amphoe Mae Sot: Tambon<br />
Kanei Jiu, Huei Kit, Salween drainage; holotype:<br />
ZRC 38471 [ex ZSM 27480]; adjective, -is, -is, -e)<br />
10.34.110 Schistura montana McClelland, 1838<br />
Schistura montana McClelland, 1838: 947, pl. 55 fig. 1 (type<br />
locality: India: Himachal Pradesh: Simla [Shimla,<br />
31°06'12"N 77 10'20"E]; syntypes: ? BMNH 1860.3.19.<br />
118–119 [2]; also in M'Clelland, 1839: 307, 440, pl. 57<br />
fig. 1; adjective, -us, -a, -um)<br />
10.34.111 Schistura mukambbikaensis (Menon, 1987)<br />
Noemacheilus denisoni mukambbikaensis Menon, 1987:<br />
101, pl. 10 fig. 7 (type locality: India: Karnataka: Kashi<br />
stream, a tributary <strong>of</strong> Kollur River, Mukambbika, 145<br />
km northeast <strong>of</strong> Mangalore; holotype: ZSI/SRS F 568;<br />
adjective, -is, -is, -e)<br />
Taxonomic notes. See under Schistura denisoni.<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
113<br />
10.34.112 Schistura multifasciata (Day, 1878)<br />
Nemachilus multifasciatus Day, 1878a: 617, pl. 153 fig. 7<br />
(type locality: India: Darjeeling and Assam; syntypes: ZSI<br />
F 2677 [1], BMNH 1889.2.1.1665–1667 [3], NMW<br />
48428 [1], AMS B.7737 [1], Whitehead & Talwar, 1976:<br />
157, Menon & Yazdani, 1968: 123, Eschmeyer & Fricke,<br />
2010; adjective, -us, -a, -um)<br />
Nomenclatural notes. Day (1878a: 617) had at least two<br />
specimens <strong>of</strong> N. multifasciatus as he listed the distribution<br />
as Darjeeling and Assam, stating that the illustrated specimen<br />
came from Darjeeling. Thus the specimen listed as holotype<br />
by Menon & Yazdani (1968: 123) can only be a syntype.<br />
10.34.113 ? Schistura nagaensis (Menon, 1987)<br />
Noemacheilus nagaensis Menon, 1987: 117 (type locality:<br />
India: Nagaland: Phodung River, tributary <strong>of</strong> Tizu River<br />
[Chindwin basin; Vishwanath & Laisram, 2001: 209];<br />
holotype: ZSI/SRS F 574; adjective, -is, -is, -e)<br />
Taxonomic notes. Possibly a synonym <strong>of</strong> S. kangjupkhulensis.<br />
10.34.114 Schistura nagodiensis Sreekantha, Gururaja,<br />
Remadevi, Indra & Ramachandra, 2006<br />
Schistura nagodiensis Sreekantha, Gururaja, Remadevi, Indra<br />
& Ramachandra, 2006: 2211, fig. web suppl. 1a (type<br />
locality: India: Karnataka: Sharavathi River (13°54'40"N<br />
74°53'49"E), Algod, Shimoga; holotype: ZSI/SRS<br />
F-7595; adjective, -is, -is, -e)<br />
10.34.115 Schistura namboensis Freyh<strong>of</strong> & Serov, 2001<br />
Schistura namboensis Freyh<strong>of</strong> & Serov, 2001: 163, figs. 32–<br />
34 (type locality: Vietnam: Dac Lac Province: small<br />
stream at Ea Nuol, about 20 km north-west <strong>of</strong> Buon Ma<br />
Thuot; 12°41.58'N 107°57.74'E; holotype: ZFMK 24094;<br />
adjective, -is, -is, -e)<br />
10.34.116 Schistura nandingensis (Zhu & Wang, 1985)<br />
Noemacheilus nandingensis Zhu & Wang, 1985: 212, figs. 7–<br />
8 (type locality: China: Yunnan: Yunxian County: small<br />
tributary <strong>of</strong> Nanding River [Salween drainage], near<br />
Xingfu, 24°30'N 100°10'E; holotype: NGI 810497; adjective,<br />
-is, -is, -e)<br />
Taxonomic notes. Treated as synonym <strong>of</strong> Pteronemacheilus<br />
meridionalis by Yang (in Chu & Chen, 1990: 49).<br />
10.34.117 Schistura nasifilis (Pellegrin, 1936)<br />
Nemachilus spilopterus var. nasifilis Pellegrin, 1936: 247<br />
(type locality: Vietnam: Annam: Nha Trang: Song Cai<br />
basin: Song Ko, 680 masl / Song Tan, 1400 masl; syntypes:<br />
MNHN 1936.21–25 [19], 1936.26–28 [20],<br />
BMNH 1936.7.30.3 [1], Kottelat, 1990a: 168; compound<br />
noun, indeclinable)<br />
10.34.118 Schistura nicholsi (Smith, 1933)<br />
Nemacheilus nicholsi Smith, 1933: 53, fig. 1, pl. 1 fig. 1<br />
(type locality: Thailand: Pak Jong, circle <strong>of</strong> Nakon Rajasima<br />
[Nakhon Ratchasima Province; Mekong drainage];<br />
holotype: KUMF 170, Monkolprasit, 1969: 6; noun in<br />
genitive, indeclinable)
10.34.124 Schistura obeini, CMK 22841, 77.6 mm SL; Laos:<br />
Mekong drainage: Nam Theun.<br />
10.34.124 Schistura obeini, CMK 22829, 67.4 mm SL.<br />
Nemacheilus thai Fowler, 1934: 104, fig. 56 (type locality:<br />
Thailand: Bua Yai, Mekong drainage; holotype: ANSP<br />
56655; noun in apposition, indeclinable)<br />
? Schistura laterimaculata Kottelat, 1990a: 141, fig. 103<br />
(type locality: Thailand: Chayaphum Province: Kuan<br />
Gielom, Mekong drainage; holotype: ZRC 38470 [ex<br />
ZSM 27477]; adjective, -us, -a, -um)<br />
10.34.119 Schistura nilgiriensis (Menon, 1987)<br />
Noemacheilus nilgiriensis Menon, 1987: 106, pl. 10 figs. 8–<br />
9, pl. 14 figs. 1–2 (type locality: India: Tamil Nadu: Nilgiri<br />
District: small stream joining the outlet stream from<br />
Pykara dam; holotype: ZSI/SRS F 561; adjective,<br />
-is, -is, -e)<br />
10.34.120 Schistura nomi Kottelat, 2000<br />
Schistura nomi Kottelat, 2000: 66, fig. 47 (type locality:<br />
Laos: Attapu Province: Xe Kaman at Muang Xaisettha,<br />
Mekong drainage; 14°48'27"N 106°55'52"E; holotype:<br />
ZRC 45359; noun in genitive, indeclinable)<br />
10.34.121 Schistura notostigma (Bleeker, 1863)<br />
Nemacheilus notostigma Bleeker, 1863b: 254 (type locality:<br />
Ceylon [Sri Lanka]; syntypes: RMNH [3, 57–71 mm<br />
TL]; also in Bleeker, 1864b: 5, pl. 1 fig. 2; compound<br />
noun, indeclinable)<br />
10.34.122 Schistura novemradiata Kottelat, 2000<br />
Schistura novemradiata Kottelat, 2000: 66, fig. 48 (type<br />
locality: Laos: Louangnamtha Province: Nam Luang,<br />
about 2 km upstream <strong>of</strong> Ban Namluang, Mekong drainage;<br />
21°09'05"N 101°20'34"E; holotype: ZRC 45360;<br />
adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
114<br />
10.34.120 Schistura nomi, CMK 21572, 49.0 mm SL; Laos:<br />
Mekong drainage: Xe Kong.<br />
10.34.134 Schistura poculi, CMK 16153, 40.1 mm SL; Thailand:<br />
Chao Phraya drainage: Mae Taeng.<br />
10.34.123 Schistura nudidorsum Kottelat, 1998<br />
Schistura nudidorsum Kottelat, 1998a: 80, fig. 119 (type<br />
locality: Laos: Khammouan Province: Nam Theun drainage:<br />
Nam Phao at waterfall immediately downriver <strong>of</strong><br />
border post on road from Lak Sao to Vinh (Vietnam),<br />
Mekong drainage; 18°23'00"N 105°09'20"E; holotype:<br />
ZRC 41801; compound noun, indeclinable)<br />
10.34.124 Schistura obeini Kottelat, 1998<br />
Schistura obeini Kottelat, 1998a: 83, fig. 122 (type locality:<br />
Laos: Khammouan Province: Nam Theun drainage: Nam<br />
Phao at waterfall immediately downriver <strong>of</strong> border post<br />
on road from Lak Sao to Vinh (Vietnam), Mekong drainage;<br />
18°23'00"N 105°09'20"E; holotype: ZRC 41803;<br />
noun in genitive, indeclinable)<br />
10.34.125 Schistura oedipus (Kottelat, 1988)<br />
Nemacheilus oedipus Kottelat, 1988c: 225, figs. 1–3 (type<br />
locality: Thailand: Mae Hong Son Province: Nam Lang<br />
Cave, 19°31'N 98°09'E, Salween drainage; holotype:<br />
AMS I 25986-002; noun in apposition, indeclinable)<br />
10.34.126 ? Schistura orthocauda (Mai, 1978)<br />
Barbatula orthocauda Mai, 1978: 231, fig. 105 (type locality:<br />
northern Vietnam; holotype: DVZUT; compound<br />
noun, indeclinable)<br />
Taxonomic notes. The type locality information is useless.<br />
Nguyen (2005: 245) identified material from Ky Phu Dai Tu<br />
(Thai Nguyen Province, Vietnam; Red River drainage) as<br />
S. orthocauda. A proper allocation <strong>of</strong> the name will only be<br />
possible after a critical revision <strong>of</strong> the barred Schistura from<br />
the Red River drainage.<br />
10.34.127 Schistura papulifera Kottelat, Harries &<br />
Proudlove, 2007<br />
Schistura papulifera Kottelat, Harries & Proudlove, 2007:<br />
36 fig. 1 (type locality: India: Meghalaya, Maintia Hills,<br />
Krem Umsngat entrance (25º11'14"N 92°21'03"E) to<br />
Synrang Pamiang cave system (main entrance 25º12'48"N<br />
92°21'48"E); holotype: MHNG 2680.074; adjective,<br />
-fer, -fera, -ferum)
10.34.144 Schistura punctifasciata, CMK 17134, 39.6 mm SL;<br />
Laos: Mekong drainage: Xe Bangfai.<br />
10.34.145 Schistura quaesita, CMK 22699, 79.7 mm SL; Laos:<br />
Mekong drainage: Nam Ngum.<br />
10.34.128 Schistura paucicincta Kottelat, 1990<br />
Schistura paucicincta Kottelat, 1990a: 177, fig. 133 (type<br />
locality: Thailand: Tak Province: Salween basin, Huai<br />
Mae Charno, 4 km south <strong>of</strong> Amphoe Mae Ramat on road<br />
1085, Salween drainage, 16°58'N 98°34'E; holotype:<br />
ZRC 38464 [ex ZSM 27481]; compound noun, indeclinable)<br />
10.34.129 Schistura paucifasciata (Hora, 1929)<br />
Nemachilus paucifasciatus Hora, 1929: 330, fig. 3, pl. 15<br />
figs. 1–2 (type locality: Burma: Northern Shan States:<br />
Hsipaw State: Monglong Subdivision: Hwe-gna-sang<br />
River, Pazi Township [Irrawaddy drainage]; holotype:<br />
ZSI F 6314/1; adjective, -us, -a, -um)<br />
10.34.130 Schistura personata Kottelat, 2000<br />
Schistura personata Kottelat, 2000: 67, fig. 49 (type locality:<br />
Laos: Saisomboun Special Zone: Houay Sala Yai,<br />
tributary <strong>of</strong> Nam San, Mekong drainage; 18°35'17"N<br />
103°05'00"E; holotype: ZRC 45362; adjective, -us, -a,<br />
-um)<br />
10.34.131 Schistura pertica Kottelat, 2000<br />
Schistura pertica Kottelat, 2000: 67, fig. 50 (type locality:<br />
Laos: Phongsali Province: Nam Ou at confluence with<br />
Houay Nam, 3 km east-southeast <strong>of</strong> Muang Khoa,<br />
Mekong drainage; 21°04'10.4"N 102°31'43.8"E; holotype:<br />
ZRC 45364; noun in apposition, indeclinable)<br />
10.34.132 Schistura pervagata Kottelat, 2000<br />
Schistura pervagata Kottelat, 2000: 68, fig. 51 (type locality:<br />
Laos: Houaphan Province: small stream tributary <strong>of</strong><br />
Nam Hao, Nam Ma drainage; 20°31'09"N 104°21'44"E;<br />
holotype: ZRC 45366; adjective, -us, -a, -um)<br />
10.34.133 ? Schistura phongthoensis (Nguyen, 2005)<br />
Paracobitis phongthoensis Nguyen, 2005: 564, fig. 11 (type<br />
locality: Vietnam: Lai Chau Province: Phong Tho district:<br />
Muong So [Song Da drainage]; holotype:<br />
NCNTTSI; adjective, -is, -is, -e)<br />
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115<br />
10.34.155 Schistura rupecula, KU 40561, 64.0 mm SL; Nepal:<br />
Kosi drainage: Indrawati. (Photograph by Kevin Conway).<br />
10.34.155 Schistura rupecula, KU 40571, about 40 mm SL. (Photograph<br />
by Kevin Conway).<br />
10.34.134 Schistura poculi (Smith, 1945)<br />
Noemacheilus poculi Smith, 1945: 323, fig. 72 (type locality:<br />
Thailand: Chiang Mai Province: Doi Angka [Doi Inthanon];<br />
holotype: MCZ 35525; noun in genitive, indeclinable)<br />
Taxonomic notes. Schistura poculi as used by Kottelat<br />
(1990a: 186) in fact includes several species (see, e.g. under<br />
S. hoai).<br />
10.34.135 Schistura polytaenia (Zhu, 1982)<br />
Nemachilus polytaenia Zhu, 1982a: 106, fig. 3 (type locality:<br />
China: Yunnan: tributary <strong>of</strong> Irrawaddy River in<br />
Tengchong County; holotype: NPIB 784092; compound<br />
noun, indeclinable)<br />
10.34.136 Schistura porthos Kottelat, 2000<br />
Schistura porthos Kottelat, 2000: 69, fig. 52 (type locality:<br />
Laos: Louangnamtha Province: Nam Tha at Ban Finho,<br />
about 14 km north <strong>of</strong> Luang Nam Tha, Mekong drainage;<br />
21°04'44"N 101°24'09"E; holotype: ZRC 45368;<br />
noun in apposition, indeclinable)<br />
10.34.137 Schistura prashadi (Hora, 1921)<br />
Nemachilus prashadi Hora, 1921b: 203, pl. 10 fig. 2–2a (type<br />
locality: India: Manipur: Thonagpal Tank, Thoubal<br />
Stream and Sikmai Stream [Chinwin drainage]; holotype:<br />
ZSI F 9987/1; noun in genitive, indeclinable)<br />
Nomenclatural notes. Hora (1921b: 203) had material <strong>of</strong><br />
N. prashadi based on 74 specimens from 3 localities, in<br />
Chindwin drainage. He designated ZSI F 9987/1 as holotype<br />
but did not mention its locality. Menon & Yazdani (1968:<br />
124) listed 5 "syntypes", ZSI 9988/1 from Thonagpal Tank,<br />
5 miles from Yaribuk. Menon (1987: 129) mentioned 26<br />
syntypes also as ZSI 9988/1.
10.34.138 Schistura pridii Vidthayanon, 2003<br />
Schistura pridii Vidthayanon, 2003: 307, fig. 1 (type locality:<br />
Thailand: Chiang Mai Province: stream Nam Muen,<br />
a tributary <strong>of</strong> Mae Taeng, upper Chao Phraya drainage;<br />
holotype: NIFI 2909; noun in genitive, indeclinable)<br />
10.34.139 Schistura procera Kottelat, 2000<br />
Schistura procera Kottelat, 2000: 69, fig. 53 (type locality:<br />
Laos: Oudomxai Province: Nam Phak watershed: waterfall<br />
Taad Lak Sip Et, km 11 on road from Oudomxai to<br />
Nambak, Mekong drainage; 20°37'01.8"N 102°00'<br />
12.0"E; holotype: ZRC 45370; adjective, -us, -a, -um)<br />
10.34.140 Schistura prolixifasciata Zheng, Yang & Chen,<br />
2012<br />
Schistura prolixifasciata Zheng, Yang & Chen, 2012b: 64,<br />
fig. 1 (type locality: China: Yunnan: Cangyuan County:<br />
Mengnonghe River, a tributary <strong>of</strong> Nangunhe River (Salween<br />
drainage) at Banlao Village, 23°30'08.9"N 99°01'<br />
42.9" E, 510 masl; holotype: KIZ 200504111365; adjective,<br />
-us, -a, -um)<br />
10.34.141 Schistura pseud<strong>of</strong>asciolata Zhou & Cui, 1993<br />
Schistura pseud<strong>of</strong>asciolata Zhou & Cui, 1993: 89, fig. 10<br />
(type locality: China: Sichuan: Huidong: Canyu River (a<br />
tributary <strong>of</strong> Jinshajiang River [Yangtze]) near Huidong;<br />
26°38'N 102°34'E; holotype: KIZ 90110157; adjective,<br />
-us, -a, -um)<br />
10.34.142 Schistura psittacula Freyh<strong>of</strong> & Serov, 2001<br />
Schistura psittacula Freyh<strong>of</strong> & Serov, 2001: 170, figs. 37–<br />
38 (type locality: Vietnam: Quang Tri Province: middle<br />
Cam Lo River about 20 km west <strong>of</strong> Dong Ha; 16°47.00'N<br />
106°53.86'E; holotype: ZFMK 27015; adjective, -us, -a,<br />
-um)<br />
10.34.143 Schistura pumatensis Nguyen & Nguyen, 2007<br />
Schistura pumatensis Nguyen [X. K.] & Nguyen [H. D.],<br />
2007: 17, fig. (type locality: Vietnam: Nghe An Province:<br />
Khe Bu stream in Pumat National Park, Lam drainage;<br />
holotype: Pumat National Park PM020170; adjective,<br />
-is, -is, -e)<br />
10.34.144 Schistura punctifasciata Kottelat, 1998<br />
Schistura punctifasciata Kottelat, 1998a: 84, fig. 123 (type<br />
locality: Laos: Khammouan Province: Xe Banfai drainage:<br />
Nam Kathang, 5 km north <strong>of</strong> Ban Gnomalat at Ban<br />
Keovilay, Mekong drainage; 17°39'N 105°10'20"E; holotype:<br />
ZRC 41804; adjective, -us, -a, -um)<br />
10.34.145 Schistura quaesita Kottelat, 2000<br />
Schistura quaesita Kottelat, 2000: 70, fig. 54 (type locality:<br />
Laos: Xiangkhouang Province: Nam Ngum, rapids downstream<br />
<strong>of</strong> Ban Latbouak, Mekong drainage; 19°36'28"N<br />
103°14'23"E; holotype: ZRC 45372; adjective, -us, -a,<br />
-um)<br />
10.34.146 Schistura quasimodo Kottelat, 2000<br />
Schistura quasimodo Kottelat, 2000: 70, fig. 55 (type locality:<br />
Laos: Saisomboun Special Zone: Houay Sala Yai, a<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
116<br />
tributary <strong>of</strong> Nam San, Mekong drainage; 18°35'17"N<br />
103°05'00"E; holotype: ZRC 45373; noun in apposition,<br />
indeclinable)<br />
10.34.147 Schistura rajasthanica (Mathur & Yazdani,<br />
1970)<br />
Noemacheilus rajasthanicus Mathur & Yazdani, 1970: 97,<br />
fig. 1 (type locality: India: Rajasthan: Takhat Sagar, Kailana<br />
[26°18'19"N 72°58'23"E], Jodhpur; holotype: ZSI;<br />
adjective, -us, -a, -um)<br />
10.34.148 Schistura rara (Zhu & Cao, 1987)<br />
Noemacheilus rarus Zhu & Cao, 1987: 328, figs. 7–8 (type<br />
locality: China: Guangdong: Ruyuan County [24°52'42"N<br />
113°11'18"E]; holotype: NGI 840130; adjective, -us, -a,<br />
-um)<br />
10.34.149 Schistura reidi (Smith, 1945)<br />
Noemacheilus reidi Smith, 1945: 313, fig. 67 (type locality:<br />
Thailand: Mae Hong Son Province: Huey Mekong Kha,<br />
a mountain torrent at base <strong>of</strong> Doi Mekong Kha, Salween<br />
drainage; holotype: USNM 107944; noun in genitive,<br />
indeclinable)<br />
10.34.150 Schistura reticulata Vishwanath & Nebeshwar,<br />
2004<br />
Schistura reticulata Vishwanath & Nebeshwar, 2004: 324,<br />
fig. 1 (type locality: India: Manipur: Chindwin drainage:<br />
Ukhrul District: Maklang River; holotype: MUMF 4400;<br />
adjective, -us, -a, -um)<br />
10.34.151 Schistura reticul<strong>of</strong>asciata (Singh & Banarescu,<br />
in Singh, Sen, Banarescu & Nalbant, 1982)<br />
Mesonoemacheilus reticul<strong>of</strong>asciatus Singh & Bănărescu, in<br />
Singh, Sen, Bănărescu & Nalbant, 1982: 206, figs. 12–<br />
16 (type locality: India: Meghalaya: Barani, 20 km from<br />
Shillong; holotype: ZSI/ERS 3062; adjective, -us, -a,<br />
-um)<br />
10.34.152 Schistura rikiki Kottelat, 2000<br />
Schistura rikiki Kottelat, 2000: 71, fig. 56 (type locality:<br />
Laos: Attapu Province: Xe Kong between Attapu and<br />
downstream to Ban Ouk, Mekong drainage; 14°44'51"N<br />
106°43'59"E; holotype: ZRC 45375; noun in apposition,<br />
indeclinable)<br />
10.34.153 Schistura robertsi Kottelat, 1990<br />
Schistura robertsi Kottelat, 1990a: 198, fig. 148 (type locality:<br />
Thailand: Phangnga Province: tributary <strong>of</strong> Khlong<br />
Khao Thalu at Ban Bang Kan, 8°33'N 98°28'E, road from<br />
Phangnga to Kapong, km 22; holotype: ZRC 38472 [ex<br />
ZSM 27482]; noun in genitive, indeclinable)<br />
10.34.154 Schistura rosammai (Sen, 2009)<br />
Aborichthys rosammai Sen, 2009: 15, fig. 1, pls. 1–2 (type<br />
locality: India: Assam: Dhemaji District [27°28'27"N<br />
93°33'50"E]: Pabomukh, Subansiri River (27°34'N<br />
94°14'E); holotype: ZSI/ERS V/F/2129; noun in genitive,<br />
indeclinable)
10.34.155 Schistura rupecula McClelland, 1838<br />
Schistura rupecula McClelland, 1838: 948, pl. 55 fig. 3 (type<br />
locality: India: Himachal Pradesh: Simla [Shimla,<br />
31°06'12"N 77 10'20"E]; syntypes: SMF 8993 [6]; also<br />
in M'Clelland, 1839: 309, 441, erratum; adjective, -us,<br />
-a, -um [but see below])<br />
Schistura rupicola M'Clelland, 1839: erratum (unjustified<br />
emendation <strong>of</strong> Schistura rupecula McClelland, 1838:<br />
948)<br />
Nemachilus rupecula var. inglisi Hora, 1935a: 58, pl. 3 figs.<br />
9–10 (type locality: India: Eastern Himalayas, rivers below<br />
Darjeeling and in Sikkim; holotype: ZSI F11755/1;<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. Menon (1987: 52) placed without explanation<br />
N. r. inglisi in the synonymy <strong>of</strong> S. multifasciata, which<br />
is not followed. The figures <strong>of</strong> the two species in Hora<br />
(1935a) and the figure <strong>of</strong> S. multifasciata in Day (1878a:<br />
pl. 153 fig. 7) show conspicuously differently shaped and<br />
patterned species.<br />
Nomenclatural notes. Schistura rupecula first appeared in<br />
McClelland (1838). It was repeated in M'Clelland (1839:<br />
309, 441). The erratum sheet in the second work includes a<br />
correction <strong>of</strong> rupecula into rupicola. Such a correction would<br />
have been valid only if included in the original description.<br />
The correction was linguistically correct but for nomenclature,<br />
formally, it is an unjustified emendation.<br />
Names ending in -cola (<strong>of</strong> which -cula is a misspelling)<br />
are nouns; they are <strong>of</strong>ten mistaken as adjectives. McClelland<br />
(1838) did not indicate if it was an noun or an adjective<br />
and this is not apparent from the species account alone; therefore<br />
it could be considered as a noun in apposition (Code<br />
art. 31.2.2). But elsewhere in both the 1838 and 1839 works<br />
McClelland described a Gonorhynchus rupeculus, in which<br />
rupeculus is an adjective. I take this as informative that, in<br />
Schistura too, rupecula is an adjective.<br />
10.34.156 Schistura russa Kottelat, 2000<br />
Schistura russa Kottelat, 2000: 71, fig. 57 (type locality:<br />
Laos: Louangnamtha Province: Nam Tha at Ban Finho,<br />
about 14 km north <strong>of</strong> Luang Nam Tha, Mekong drainage;<br />
21°04'44"N 101°24'09"E; holotype: ZRC 45377;<br />
adjective, -us, -a, -um)<br />
10.34.157 Schistura savona (Hamilton, 1822)<br />
Cobitis savona Hamilton, 1822: 357, 394 (type locality: India:<br />
Kosi River [at Nathpur near the Nepal frontier; Hora,<br />
1935a: 49]; types: NT; Hamilton's unpublished figure<br />
reproduced in M'Clelland, 1839: pl. 53 fig. 3; noun in<br />
apposition, indeclinable)<br />
Cobites obscura Swainson, 1839: 310 (available by indication<br />
to Hamilton, 1822: 357, No. 9 [which is C. savona];<br />
type locality: India: Kosi River [at Nathpur; Hora, 1935a:<br />
49]; types: NT; Hamilton's unpublished figure reproduced<br />
in M'Clelland, 1839: pl. 53 fig. 3; adjective, -us, -a, -um)<br />
? Shistura [sic] yenjitee Tekriwal & Rao, 1999: 93 (nomen<br />
nudum; locality: Bhutan: Phuntsholing area)<br />
10.34.158 Schistura scaturigina M'Clelland, 1839<br />
Schistura scaturigina M'Clelland, 1839: 308, 443, pl. 53<br />
fig. 6 (type locality: no data [probably India: below Dar-<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
117<br />
jeeling Himalayas; Hora, 1935a: 65]; types: lost, based<br />
on an unpublished drawing by Hamilton; adjective, -us,<br />
-a, -um)<br />
Schistura subfusca M'Clelland, 1839: 308, 443, pl. 53 fig. 5<br />
(type locality: India: Upper Assam; types: LU; simultaneous<br />
subjective synonym <strong>of</strong> Schistura scaturigina<br />
M'Clelland, 1839: 308, 443, first reviser [apparently<br />
Hora, 1935a: 64] gave precedence to S. scaturigina; adjective,<br />
-us, -a, -um)<br />
Taxonomic notes. Synonymy partly based on Hora (1935a:<br />
64) and Menon (1987: 86). Confirmation seems needed.<br />
Menon also included S. shebbearei and S. zonata in the synonymy<br />
<strong>of</strong> S. scaturigina but they are retained as valid here.<br />
The status <strong>of</strong> all these nominal species should be critically<br />
re-examined.<br />
10.34.159 Schistura schultzi (Smith, 1945)<br />
Noemacheilus schultzi Smith, 1945: 317, fig. 69 (type locality:<br />
Thailand: Chiang Rai Province: Huey Melao on Doi<br />
Hua Mot, Mekong drainage; holotype: USNM 107953;<br />
noun in genitive, indeclinable)<br />
10.34.160 Schistura semiarmata (Day, 1867)<br />
Nemacheilus semiarmatus Day, 1867a: 286 (type locality:<br />
India: Nilgherries: Bowany [Bhavani] and Seegoor Rivers,<br />
Billicul Lake; syntypes: BMNH 1867.7.24.25–27 [3],<br />
AMS B.7740 [1], Whitehead & Talwar, 1976: 157; adjective,<br />
-us, -a, -um)<br />
10.34.161 Schistura sertata Kottelat, 2000<br />
Schistura sertata Kottelat, 2000: 71, fig. 58 (type locality:<br />
Laos: Louangphabang Province: Mekong drainage, Nam<br />
Xi below Kuang Xi waterfall, upstream <strong>of</strong> Ban Thapen;<br />
19°45'10"N 102°00'10"E; holotype: ZRC 45379; adjective,<br />
-us, -a, -um)<br />
10.34.162 Schistura sexcauda (Fowler, 1937)<br />
Nemacheilus sexcauda Fowler, 1937: 156, figs. 75–76 (type<br />
locality: Thailand: Me Poon [Mae Phun, a stream entering<br />
Mae Nam Yom at 17°40'N 99°42'E, Chao Phraya<br />
drainage, de Schauensee, 1946: 5; see Kottelat, 1990a:<br />
207 for discussion <strong>of</strong> locality]; holotype: ANSP 68007;<br />
compound noun, indeclinable)<br />
Noemacheilus fowlerianus Smith, 1945: 306 (type locality:<br />
Thailand: Me Poon [Mae Phun, a stream entering Mae<br />
Nam Yom at 17°40'N 99°42'E, Chao Phraya drainage,<br />
de Schauensee, 1946: 5; see Kottelat, 1990a: 207 for discussion<br />
<strong>of</strong> locality]; lectotype: ANSP 108189, designated<br />
by Kottelat, 1990a: 207; adjective, -us, -a, -um)<br />
10.34.163 Schistura shadiwalensis Mirza & Nalbant, in<br />
Mirza, Nalbant & Banarescu, 1981<br />
Schistura shadiwalensis Mirza & Nalbant, in Mirza, Nalbant<br />
& Bănărescu, 1981: 125, figs. 51–55 (type locality:<br />
Pakistan: Punjab: Gujrat District: Shadiwal [32°37'13"N<br />
74°14'49"E] on Chenab River, right-side tributary <strong>of</strong><br />
Sutlej River; holotype: ISBB 3369; adjective, -is, -is, -e)<br />
Taxonomic notes. Menon (1987: 74) treated this species as<br />
synonym <strong>of</strong> S. horai, without discussion. The figures in the<br />
original descriptions <strong>of</strong> both species show great differences
10.34.174 Schistura spekuli, IRSNB 823, 57.2 mm SL, holotype;<br />
Vietnam: Lai Chau Province: cave in Chi Chu Chai.<br />
10.34.179 Schistura suber, CMK 22522, 44.8 mm SL; Laos:<br />
Mekong drainage: Nam Ngum.<br />
in body shape and colour pattern and without further information<br />
I can only retain the species as valid.<br />
10.34.164 Schistura sharavathiensis Sreekantha, Gururaja,<br />
Remadevi, Indra & Ramachandra, 2006<br />
Schistura sharavathiensis Sreekantha, Gururaja, Remadevi,<br />
Indra & Ramachandra, 2006: 2213, web suppl. fig. 3 (type<br />
locality: India: Karnataka: Shimoga: Sharavati River<br />
(14°08'15"N 74°44'30"E), Kalkatte tributary, 1 km upriver<br />
to Dabbe falls; holotype: ZSI/SRS F-7597; adjective,<br />
-is, -is, -e)<br />
10.34.165 Schistura shebbearei (Hora, 1935)<br />
Nemachilus shebbearei Hora, 1935a: 52, pl. 3 figs. 1–2 (type<br />
locality: India: Northern Bengal [probably rivers and<br />
streams below Darjeeling; Hora, 1935a: 53]; holotype:<br />
ZSI F 11720/1; noun in genitive, indeclinable)<br />
Taxonomic notes. Listed without explanation as a synonym<br />
<strong>of</strong> S. scaturigina by Menon (1987: 86), which I do not follow<br />
judging from the great differences between the two species<br />
as described and figured by Hora (1935a).<br />
Nomenclatural notes. Hora gave the catalogue number <strong>of</strong><br />
the holotype as ZSI F 11720/1, Menon & Yazdani (1968:<br />
125) and Eschmeyer & Fricke (2010) listed it as ZSI<br />
F 11719/1.<br />
10.34.166 Schistura shuangjiangensis (Zhu & Wang, 1985)<br />
Noemacheilus shuangjiangensis Zhu & Wang, 1985: 215,<br />
figs. 18–23 (type locality: China: Yunnan: Shuangjiang<br />
County: Xiaoheijiang River, tributary <strong>of</strong> Lancangjiang<br />
River [Mekong], 23°20'N 99°50'E; holotype: NGI<br />
810443; adjective, -is, -is, -e)<br />
10.34.167 Schistura sigillata Kottelat, 2000<br />
Schistura sigillata Kottelat, 2000: 72, fig. 59 (type locality:<br />
Laos: Saisomboun Special Zone: Houay Sala Yai, a tributary<br />
<strong>of</strong> Nam San, Mekong drainage; 18°35'17"N 103°05'<br />
00"E; holotype: ZRC 45381; adjective, -us, -a, -um)<br />
10.34.168 Schistura sijuensis (Menon, 1987)<br />
Noemacheilus sijuensis Menon, 1987: 175, pl. 6 fig. 2 (type<br />
locality: India: Meghalaya: Garo Hills: Siju Cave, 100<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
118<br />
10.34.185 Schistura tizardi, CMK 22361, 55.9 mm SL; Laos:<br />
Mekong drainage: Bolaven Plateau.<br />
10.34.187 Schistura udomritthiruji, CMK 22042, 42.4 mm SL;<br />
Thailand: Ranong Province: Kapoe.<br />
feet from entrance [25°21'02"N 90°41'04"E]; holotype:<br />
ZSI/SRS F 570; adjective, -is, -is, -e)<br />
10.34.169 Schistura sikmaiensis (Hora, 1921)<br />
Nemachilus sikmaiensis Hora, 1921b: 201, pl. 9 fig. 4, pl. 10<br />
figs. 1–1a (type locality: Burma: Putao, Irrawaddy drainage<br />
[original type locality: India: Manipur: Sikmai stream<br />
near Palel]; neotype: NRM 10740, designated by Kottelat,<br />
1990a: 210; adjective, -is, -is, -e)<br />
Nemacheilus putaoensis Rendahl, 1948: 27, figs. 11–12 (type<br />
locality: Burma: Putao, Irrawaddy drainage; holotype:<br />
NRM 10740; junior objective synonym <strong>of</strong> Nemachilus<br />
sikmaiensis Hora, 1921b: 201; adjective, -is, -is, -e)<br />
10.34.170 Schistura similis Kottelat, 1990<br />
Schistura similis Kottelat, 1990a: 210, fig. 156 (type locality:<br />
Thailand: Tak Province: Amphoe Tha Song Yang: Huei<br />
Ki Teu, Tambon Kanei Jiu, Salween drainage; holotype:<br />
ZRC 38478 [ex ZSM 27483]; adjective, -is, -is, -e)<br />
10.34.171 Schistura singhi (Menon, 1987)<br />
Noemacheilus singhi Menon, 1987: 119, pl. 16 fig. 1 (type<br />
locality: India: Nagaland: Kiphire [25°53'57"N<br />
94°46'51"E]; holotype: ZSI/SRS F 580; noun in genitive,<br />
indeclinable)<br />
10.34.172 Schistura sokolovi Freyh<strong>of</strong> & Serov, 2001<br />
Schistura sokolovi Freyh<strong>of</strong> & Serov, 2001: 172, figs. 40–41<br />
(type locality: Vietnam: Gia Lai Province: Azun River<br />
about 30 km east <strong>of</strong> Pleiku, a tributary to Ba River;<br />
14°02.34'N 108°21.07'E; holotype: ZFMK 27116; noun<br />
in genitive, indeclinable)<br />
10.34.173 Schistura sombooni Kottelat, 1998<br />
Schistura sombooni Kottelat, 1998a: 88, fig. 125 (type locality:<br />
Laos: Khammouan Province: Nam Phao 2 km upriver<br />
<strong>of</strong> Ban Lak Song (5 km upriver <strong>of</strong> Ban Nape), tributary<br />
<strong>of</strong> Nam Theun, Mekong drainage; approx.<br />
18°20'40"N 105°07'30"E; holotype: ZRC 41806; noun<br />
in genitive, indeclinable)
10.35.1 Sectoria atriceps, CMK 16049, 38.3 mm SL; Thailand:<br />
Chao Phraya drainage: Nan River.<br />
10.35.1 Sectoria atriceps, CMK 16059, 78.7 mm SL.<br />
10.34.174 Schistura spekuli Kottelat, 2004<br />
Schistura spekuli Kottelat, 2004b: 187, fig. 1 (type locality:<br />
Vietnam: Lai Chau Province: cave in centre <strong>of</strong> Chin Chu<br />
Chai village [10 km south <strong>of</strong> Tam Duong]; about 22°24'N<br />
103°26'E; holotype: IRSNB 823; noun in genitive, indeclinable)<br />
10.34.175 Schistura spiesi Vidthayanon & Kottelat, 2003<br />
Schistura spiesi Vidthayanon & Kottelat, 2003: 165, fig. 4<br />
(type locality: Thailand: Phitsanulok Province: Thung<br />
Salaeng Luang National Park: upstream section <strong>of</strong> Tham<br />
Phra Wang Daeng cave, 100–200 m from entrance;<br />
16°40'41"N 100°41'24"E; holotype: NIFI 3150; noun in<br />
genitive, indeclinable)<br />
10.34.176 Schistura spiloptera (Valenciennes, in Cuvier<br />
& Valenciennes, 1846)<br />
Cobitis spiloptera Valenciennes, in Cuvier & Valenciennes,<br />
1846: 27, pl. 522 (type locality: Vietnam: Cochinchine;<br />
syntypes: MNHN 3434 [19], Bertin & Estève, 1948: 96;<br />
compound adjective, -us, -a, -um)<br />
Nemacheilus pellegrini Rendahl, 1944: 26, fig. 13 (type<br />
locality: Vietnam: Annam: Thua Luu, 50 km southeast <strong>of</strong><br />
Hué; holotype: NHMG; noun in genitive, indeclinable)<br />
10.34.177 Schistura spilota (Fowler, 1934)<br />
Nemacheilus spilotus Fowler, 1934: 105, fig. 57 (type locality:<br />
Thailand: Chiang Mai Province: Metang River [Nam<br />
Mae Taeng], 35 miles north <strong>of</strong> Chiang Mai, Chao Phraya<br />
drainage; holotype: ANSP 56528; adjective, -us, -a, -um)<br />
10.34.178 Schistura striata (Day, 1867)<br />
Nemacheilus striatus Day, 1867b: 347 (type locality: India:<br />
Wynaad at 3000 feet; syntypes [?]: ZSI A.2683 [lost],<br />
BMNH 1889.2.1.1661–1664 [4], AMS B.7487 [1],<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
119<br />
10.36.2 Seminemacheilus lendlii, CMK 20141, 66.5 mm SL; Turkey:<br />
Beysehir.<br />
10.37.1 Speonectes tiomanensis, CMK 15963, 41.1 mm SL; Malaysia:<br />
Tioman Island.<br />
10.38.1 Sphaerophysa dianchiensis, IHB 63V888, 69.5 mm SL,<br />
holotype; China: Yunnan: Lake Dianchi. (Photograph by Zhang E).<br />
RMNH 2675 [1], Whitehead & Talwar, 1976: 157; adjective,<br />
-us, -a, -um)<br />
10.34.179 Schistura suber Kottelat, 2000<br />
Schistura suber Kottelat, 2000: 72, fig. 60 (type locality:<br />
Laos: Vientiane Province: unnamed small forest stream<br />
along road from Thad Leuk to Nam Leuk dam site,<br />
Mekong drainage; 18°27'05"N 103°04'06"E; holotype:<br />
ZRC 45383; noun in apposition, indeclinable)<br />
10.34.180 Schistura susannae Freyh<strong>of</strong> & Serov, 2001<br />
Schistura susannae Freyh<strong>of</strong> & Serov, 2001: 179, figs. 46–<br />
47 (type locality: Vietnam: Quang Nam Danang Province:<br />
stream Mong Mo south <strong>of</strong> Hai Van Pass at national<br />
highway Nr. 1; 16°09.67'N 108°07.73'E; holotype:<br />
ZFMK 28382; noun in genitive, indeclinable)<br />
10.34.181 Schistura tenura Kottelat, 2000<br />
Schistura tenura Kottelat, 2000: 73, fig. 61 (type locality:<br />
Laos: Vientiane Province: Nam Leuk about 1 km downstream<br />
<strong>of</strong> dam site, Mekong drainage; 18°26'10"N 102°<br />
57'01"E; holotype: ZRC 45384; compound adjective but<br />
proposed as a noun in apposition, indeclinable)<br />
10.34.182 Schistura thanho Freyh<strong>of</strong> & Serov, 2001<br />
Schistura thanho Freyh<strong>of</strong> & Serov, 2001: 182, figs. 49–50<br />
(type locality: Vietnam: Binh Dinh Province: Vinh Thanh<br />
River, about 65 km north-west Quy Nhon; 14°16.19'N<br />
108°45.69'E; holotype: ZFMK 21047; noun in apposition,<br />
indeclinable)<br />
10.34.183 Schistura tigrina Vishwanath & Nebeshwar<br />
Sharma, 2005<br />
Schistura tigrinum Vishwanath & Nebeshwar Sharma, 2005:<br />
79, fig. 1 (type locality: India: Manipur: Tamenglong
District: Tamei Subdivision: Barak River at Khunpung,<br />
Brahmaputra drainage; MUMF 4105; adjective, -us, -a,<br />
-um)<br />
10.34.184 Schistura tirapensis Kottelat, 1990<br />
Noemacheilus arunachalensis Menon, 1987: 129, pl. 16<br />
fig. 10 (type locality: India: Arunachal Pradesh: Tirap District:<br />
Riwa River at Nampong; holotype: ZSI/SRS F 572;<br />
junior primary homonym <strong>of</strong> N. arunachalensis Datta &<br />
Barman, 1984: 275; adjective, -is, -is, -e)<br />
Schistura tirapensis Kottelat, 1990a: 118 (replacement name<br />
for Noemacheilus arunachalensis Menon, 1987: 129;<br />
adjective, -is, -is, -e)<br />
Noemacheilus dattai Menon, 1999: 170 (replacement name<br />
for Noemacheilus arunachalensis Menon, 1987: 129;<br />
noun in genitive, indeclinable)<br />
10.34.185 Schistura tizardi Kottelat, 2000<br />
Schistura tizardi Kottelat, 2000: 73, fig. 62 (type locality:<br />
Laos: Attapu Province: Xe Kaman at Muang Xaisettha,<br />
Mekong drainage; 14°48'27"N 106°55'52"E; holotype:<br />
ZRC 45386; noun in genitive, indeclinable)<br />
10.34.186 Schistura tubulinaris Kottelat, 1998<br />
Schistura tubulinaris Kottelat, 1998a: 89, fig. 127 (type locality:<br />
Laos: Khammouan Province: Nam Theun, from Ban<br />
Signo to about 6 km upriver, Mekong drainage;<br />
17°50'50"N 105°03'00"E; holotype: ZRC 41808; compound<br />
noun, indeclinable)<br />
10.34.187 Schistura udomritthiruji Bohlen & Šlechtová,<br />
2010<br />
Schistura udomritthiruji Bohlen & Šlechtová, 2010: 320,<br />
figs. 1–2 (type locality: Thailand: Ranong Province: small<br />
stream draining into Andaman Sea upstream <strong>of</strong> Kapoe;<br />
9°34'14"N 98º41'40"E; holotype: ZRC 51724; noun in<br />
genitive, indeclinable)<br />
10.34.188 Schistura vinciguerrae (Hora, 1935)<br />
Nemachilus vinciguerrae Hora, 1935a: 62, pl. 3 fig. 12 (type<br />
locality: Burma: Meekalan, Salween drainage, Salween<br />
drainage; holotype: ZSI F 11754/1; Vinciguerra is treated<br />
as a Latinized name and vinciguerrae is correct original<br />
spelling [Code art. 31.1.1 and Example]; noun in<br />
genitive, indeclinable)<br />
10.34.189 Schistura waltoni (Fowler, 1937)<br />
Nemacheilus waltoni Fowler, 1937: 157, figs. 77–79 (type<br />
locality: Thailand: Me Poon [Mae Phun, a stream entering<br />
Mae Nam Yom at 17°40'N 99°42'E, Chao Phraya<br />
drainage, de Schauensee, 1946: 5; see Kottelat, 1990a:<br />
207 for discussion <strong>of</strong> locality]; holotype: ANSP 68008;<br />
noun in genitive, indeclinable)<br />
Noemacheilus obscurus Smith, 1945: 316, fig. 68 (type locality:<br />
Thailand: Chiang Mai Province: Doi Angka [Doi Inthanon],<br />
Chao Phraya drainage; holotype: MCZ 35520;<br />
junior secondary homonym <strong>of</strong> Cobites obscura Swainson,<br />
1839: 310, if both are treated as valid species in<br />
Schistura; adjective, -us, -a, -um)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
120<br />
10.34.190 Schistura xhatensis Kottelat, 2000<br />
Schistura xhatensis Kottelat, 2000: 74, fig. 63 (type locality:<br />
Laos: Louangphabang Province: Nam Xhat, upstream<br />
<strong>of</strong> Ban Nam Sa, Mekong drainage; 20°06'43"N 103°19'<br />
56"E; holotype: ZRC 45388; adjective, -is, -is, -e)<br />
10.34.191 Schistura yersini Freyh<strong>of</strong> & Serov, 2001<br />
Schistura yersini Freyh<strong>of</strong> & Serov, 2001: 186, figs. 52–53<br />
(type locality: Vietnam: Lam Dong Province: small stream<br />
running to Da Dung River south <strong>of</strong> Da Lat; 11°46.86'N<br />
108°25.04'E; holotype: ZFMK 23579; noun in genitive,<br />
indeclinable)<br />
10.34.192 Schistura yingjiangensis (Zhu, 1982)<br />
Nemachilus yingjiangensis Zhu, 1982a: 107, fig. 4 (type<br />
locality: China: Yunnan: Yingjiang County: Daying River,<br />
a tributary <strong>of</strong> Irrawaddy River; holotype: NPIB<br />
784147; adjective, -is, -is, -e)<br />
10.34.193 Schistura zonata M'Clelland, 1839<br />
Schistura zonata M'Clelland, 1839: 308, 441, pl. 53 fig. 1<br />
(type locality: India: Upper Assam: Muttuc District; types:<br />
LU; adjective, -us, -a, -um)<br />
Nemacheilus mugah Day, 1869: 382 (type locality: India:<br />
Cossye River [Kasai] at Midnapore [22°15'N 87°39'E];<br />
types: ZSI A.961 [lost], Whitehead & Talwar, 1976: 157;<br />
noun in apposition, indeclinable)<br />
Taxonomic notes. Nemacheilus mugah is listed as a synonym<br />
<strong>of</strong> S. zonata by Day (1878a: 618) and Menon (1974:<br />
56) (1987: 86) and S. zonata is listed as a synonym <strong>of</strong><br />
S. scaturigina by Menon (1987: 86). The later, unexplained,<br />
synonymy is not followed. The identity <strong>of</strong> all these nominal<br />
species remains unsolved.<br />
10.35 Sectoria Kottelat, 1990<br />
Sectoria Kottelat, 1990a: 229 (type species: Noemacheilus<br />
atriceps Smith, 1945: 312, by original designation). Gender<br />
feminine.<br />
10.35.1 Sectoria atriceps (Smith, 1945)<br />
Noemacheilus atriceps Smith, 1945: 312, fig. 66 (type locality:<br />
Thailand: Nan Province: Menam Kon, tributary <strong>of</strong><br />
Menam Nan, at Ban Khana, Chao Phraya drainage; holotype:<br />
USNM 117750; compound adjective, indeclinable)<br />
10.35.2 Sectoria heterognathos (Chen, 1999)<br />
Schistura heterognathos Chen, 1999: 301, fig. 1 (type locality:<br />
China: Yunnan: Xishuangbanna: Mengla County:<br />
Nanla River, Nazhao (21°40'55"N 101°35'20"E), about<br />
25 km from Mengla, Mekong drainage; holotype: IHB<br />
8810240; compound noun, indeclinable)<br />
Sectoria megastoma Kottelat, 2000: 74, fig. 64 (type locality:<br />
China: Yunnan: Xishuangbanna: market in Mengla,<br />
Mekong drainage; holotype: NRM 33232; compound<br />
noun, proposed as a noun in apposition, indeclinable)
10.36 Seminemacheilus Banarescu & Nalbant, 1995<br />
Heminoemacheilus Bănărescu, 1977: 44 (nomen nudum)<br />
Seminemacheilus Bănărescu & Nalbant, 1995: 449 (type<br />
species: Nemacheilus lendlii Hankó, 1924: 155, by original<br />
designation). Gender masculine.<br />
10.36.1 Seminemacheilus ispartensis Erk'akan, Nalbant<br />
& Özeren, 2007<br />
Seminemacheilus ispartensis Erk'akan, Nalbant & Özeren,<br />
2007: 76, fig. 7 (type locality: Turkey: Isparta Creek,<br />
Egirdir-Isparta Road, 1 st railway pass, 37°38'N 30°31'E;<br />
holotype: HUIC AKD-1; adjective, -is, -is, -e)<br />
Taxonomic notes. Coordinates possibly erroneous; they are<br />
identical to those <strong>of</strong> the type locality <strong>of</strong> Barbatula mediterraneus,<br />
which is from a different locality.<br />
10.36.2 Seminemacheilus lendlii (Hankó, 1924)<br />
Nemachilus lendlii Hankó, 1924: 155, pl. 3 fig. 9 (type locality:<br />
Turkey: Eski-Chehir; syntypes: MNH [10, lost]; noun<br />
in genitive, indeclinable)<br />
10.37 Speonectes Kottelat, 2012<br />
Speonectes Kottelat, 2012: 139 [appendix <strong>of</strong> present work]<br />
(type species: Sundoreonectes tiomanensis Kottelat,<br />
1990b: 52, by original designation). Gender masculine.<br />
10.37.1 Speonectes tiomanensis (Kottelat, 1990)<br />
Sundoreonectes tiomanensis Kottelat, 1990b: 52, pl. 1 fig. 2<br />
(type locality: Malaysia: Pulau Tioman: cave in Gunung<br />
[Mount] Kajang, at 3400 feet; holotype: BMNH 1989.2.<br />
23.1; adjective, -is, -is, -e)<br />
10.38 Sphaerophysa Cao & Zhu, 1988<br />
Sphaerophysa Cao & Zhu, 1988b: 405 (type species:<br />
Sphaerophysa dianchiensis Cao & Zhu, 1988: 405, by<br />
original designation). Gender feminine.<br />
10.38.1 Sphaerophysa dianchiensis Cao & Zhu, 1988<br />
Sphaerophysa dianchiensis Cao & Zhu, 1988b: 405, fig. 1<br />
(type locality: China: Yunnan: Dianchi Lake near Kunming;<br />
holotype: IHB 63V888; adjective, -is, -is, -e)<br />
10.39 Sundoreonectes Kottelat, 1990<br />
Sundoreonectes Kottelat, 1990b: 52 (type species: Nemacheilus<br />
obesus Vaillant, 1902: 134, by original designation).<br />
Gender masculine.<br />
10.39.1 Sundoreonectes obesus (Vaillant, 1902)<br />
Nemacheilus obesus Vaillant, 1902: 134, figs. 39–40 (type<br />
locality: Indonesia: Borneo: Kalimantan Timur: Bloeoe<br />
River [Bluu, 0°42'N 114°24'E]; lectotype: RMNH 7780,<br />
designated by Kottelat, 1984b: 232; adjective, -us, -a,<br />
-um)<br />
10.39.2 Sundoreonectes sabanus (Chin, 1990)<br />
Elxis sabanus Chin, 1990: 74, fig. 3 (type locality: Malaysia:<br />
Borneo: Sabah: Sipitang District: mountain stream<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
121<br />
in Mendolong area; holotype: FMNH 98130; adjective,<br />
-us, -a, -um)<br />
10.40 Tarimichthys Prok<strong>of</strong>iev, 2010<br />
Tarimichthys Prok<strong>of</strong>iev, 2010: 896 (subgenus <strong>of</strong> Triplophysa<br />
Rendahl, 1933: 21; type species: Nemachilus bombifrons<br />
Herzenstein, 1888: 67, by original designation).<br />
Gender masculine.<br />
10.40.1 Tarimichthys bombifrons (Herzenstein, 1888)<br />
Nemachilus bombifrons Herzenstein, 1888: 67, pl. 2 fig. 2,<br />
pl. 4 fig. 2 (type locality: China: Xinjiang: Chotan River<br />
[Khotan, Hotan He; confluence with Yarkand at<br />
40°22'21"N 80°56'18"E]; lectotype: ZISP 7359, designated<br />
by Prok<strong>of</strong>iev, 2002b: 429, fig. 3; compound noun,<br />
indeclinable)<br />
10.40.2 Tarimichthys edsinicus (Prok<strong>of</strong>iev, 2003)<br />
Triplophysa edsinica Prok<strong>of</strong>iev, 2003b: 66, fig. 29 (type<br />
locality: China: "Hexi region": Edsin-Gol [Ejin, Ejina<br />
He, Eiin, Hei-He, Ruo-Shui; a river flowing north to endorheic<br />
Gaxun Nuur (Juyan Lake), 42°25'20"N<br />
100°40'07"E]; holotype: ZISP 17294; adjective, -us, -a,<br />
-um)<br />
10.40.3 Tarimichthys incipiens (Herzenstein, 1888)<br />
Nemachilus bombifrons incipiens Herzenstein, 1888: 73<br />
(type locality: China: Xinjiang: "Keria Mts." [Keriya,<br />
36°53'00"N 81°36'00"E, about 160 km east <strong>of</strong> Khotan];<br />
lectotype: ZISP 7365/1, designated by Prok<strong>of</strong>iev, 2002b:<br />
428, fig. 1; participle, indeclinable)<br />
10.41 Traccatichthys Freyh<strong>of</strong> & Serov, 2001<br />
Traccatichthys Freyh<strong>of</strong> & Serov, 2001: 188 (type species:<br />
Nemacheilus taeniatus Pellegrin & Chevey, 1936: 229,<br />
by original designation). Gender masculine.<br />
10.41.1 Traccatichthys pulcher (Nichols & Pope, 1925)<br />
Nemacheilus pulcher Nichols & Pope, in Nichols, 1925b: 1<br />
(type locality: China: Hainan: Nodoa; holotype: AMNH<br />
8364 [apparently based on single specimen listed by<br />
Nichols & Pope, 1927: 339]; adjective, -er, -ra, -rum)<br />
10.41.2 Traccatichthys taeniatus (Pellegrin & Chevey,<br />
1936)<br />
Nemacheilus pulcher var. taeniata Pellegrin & Chevey, 1936:<br />
229, fig. 6 (type locality: Vietnam: Tonkin: Thiong Khé,<br />
Phu Tho, Red River drainage; holotype: MNHN<br />
1935.348; adjective, -us, -a, -um)<br />
? Micronemacheilus zispi Prok<strong>of</strong>iev, 2004b: 158, fig. 2<br />
[p. 195 in translation] (type locality: China: Hainan: Sanya,<br />
a mountain stream 110 km from Duao-Lyu-Shan village;<br />
holotype: ZISP 46740; noun in genitive, indeclinable)<br />
? Micronemacheilus bacmeensis Nguyen & Vo, in Nguyen,<br />
2005: 562, fig. 10 (type locality: Vietnam: Ha Giang Province:<br />
Bac Me district: Minh Son stream, Red River drainage;<br />
holotype: NCNTTSI; adjective, -is, -is, -e)
10.39.2 Sundoreonectes sabanus, FMNH 99436, 54.1 mm SL;<br />
Malaysia: Borneo: Sabah: Crocker Range.<br />
10.40.1 Tarimichthys bombifrons, ZISP 7341, 203 mm SL; China:<br />
Xinjiang: Lop Nor basin, Quarqan River. (From Herzenstein,<br />
1888: pl. 2 fig. 2).<br />
10.42 Triplophysa Rendahl, 1874<br />
Diplophysa Kessler, 1874: 57 (type species: Diplophysa<br />
strauchii Kessler, 1874: 58, by subsequent designation<br />
by Berg, 1916: 348; junior homonym <strong>of</strong> Diplophysa<br />
Gegenbaur, 1850: 291 in Coelenterata). Gender feminine.<br />
Triplophysa Rendahl, 1933: 21 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus hutjertjuensis<br />
Rendahl, 1933: 28, by original designation). Gender<br />
feminine.<br />
Tauphysa Rendahl, 1933: 22 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Diplophysa kungessana<br />
Kessler, 1879: 286, by original designation). Gender<br />
feminine.<br />
Deuterophysa Rendahl, 1933: 23 (replacement name for<br />
Diplophysa Kessler, 1874: 57; junior homonym <strong>of</strong> Deuterophysa<br />
Warren, 1889: 272 in Lepidoptera). Gender<br />
feminine.<br />
Didymophysa Whitley, 1950: 44 (replacement name for<br />
Diplophysa Kessler, 1874: 57 and Deuterophysa<br />
Rendahl, 1933: 23). Gender feminine.<br />
Diplophysoides Fowler, 1958: 13 (replacement name for<br />
Diplophysa Kessler, 1874: 57 and Deuterophysa<br />
Rendahl, 1933: 23). Gender masculine.<br />
Linemacheilus Anonym, 2005: 67 (nomen nudum)<br />
Taxonomic notes. Triplophysa is an artificial assemblage<br />
<strong>of</strong> species. Among them some species have been grouped in<br />
a few diagnosable lineages and have been named (Hedinichthys,<br />
Qinghaichthys, Labiatophysa, Indotriplophysa,<br />
Tarimichthys) and treated as subgenera <strong>of</strong> Triplophysa, while<br />
the relationships <strong>of</strong> the majority are still unresolved. At this<br />
stage, these lineages seem well supported but their interrelationships<br />
are still not established. In such a situation, there<br />
does not seem to be reason or advantage in using a subgeneric<br />
rank, which implies that some relationship has been<br />
worked out. They are therefore treated as distinct, valid genera.<br />
Nomen nudum<br />
Triplophysa malloryi Bănărescu & Nalbant, 1995: figs. 24b,<br />
25e (nomen nudum)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
122<br />
10.40.1 Tarimichthys bombifrons, ASIZB 50187, 167 mm SL.<br />
10.42.1 ? Triplophysa akhtari (Vijayalakshmanan, 1950)<br />
Nemachilus akhtari Vijayalakshmanan, 1950: 219, fig. 1<br />
(type locality: Afghanistan: Farakhollum, about 10 miles<br />
south <strong>of</strong> Gardan Diwar, Helmund River; syntypes [at least<br />
6]: ZSI F 244/2 [1]; noun in genitive, indeclinable)<br />
Taxonomic notes. Treated as a synonym <strong>of</strong> T. stolickai by<br />
Prok<strong>of</strong>iev (2007b: 4), which geography suggests should be<br />
re-examined.<br />
Nomenclatural notes. Nemacheilus akhtari was described<br />
on the basis <strong>of</strong> an unspecified number <strong>of</strong> specimens; morphometric<br />
data were given for five. No holotype was indicated.<br />
Menon & Yazdani (1968: 121) list only a 'holotype'<br />
(ZSI F 244/2). This specimen is at best a syntype. The fate<br />
<strong>of</strong> the remaining specimens is not known.<br />
10.42.2 Triplophysa alexandrae Prok<strong>of</strong>iev, 2001<br />
Triplophysa alexandrae Prok<strong>of</strong>iev, 2001a: 203, fig. 13 [not<br />
19] (type locality: China: Sichuan: Min River basin, Dadu<br />
He River east <strong>of</strong> Kangding [30°03'N 101°58'E; Yangtze<br />
drainage]; holotype: ZISP 16335; noun in genitive, indeclinable)
10.41.2 Traccatichthys taeniatus, CMK 14885, 66.6 mm SL; Vietnam:<br />
Quang Ninh.<br />
10.42.3 Triplophysa aliensis Zhu & Wu, in Wu & Zhu, 1979<br />
Triplophysa aliensis Zhu & Wu, in Wu & Zhu, 1979: 29, fig.<br />
9 (type locality: China: Xizang [Tibet]: Ali, Xiang-quan<br />
and Shi-quan Rivers, Indus River drainage; syntypes:<br />
NPIB (?) 74.201–244, 255 [45]; adjective, -is, -is, -e)<br />
10.42.4 Triplophysa altipinnis Prok<strong>of</strong>iev, 2003<br />
Triplophysa altipinnis Prok<strong>of</strong>iev, 2003b: 63, fig. 25 (type<br />
locality: China: eastern Qaidam: Dzukha; holotype: ZISP<br />
12492; compound noun, indeclinable)<br />
10.42.5 Triplophysa aluensis Li & Zhu, 2000<br />
Triplophysa aluensis Li & Zhu, 2000: 396, fig. 1 (type locality:<br />
China: Yunnan: Luxi County: Yusun River in cave<br />
Alugudong, 24°17'N 103°48'E [24°33'N 103°45'E;<br />
Romero et al., 2009: 259]; holotype: HRAS 2000062001;<br />
adjective, -is, -is, -e)<br />
10.42.6 Triplophysa angeli (Fang, 1941)<br />
Nemacheilus Angeli Fang, 1941: 256 (type locality: China:<br />
western Sse-chuan [Sichuan] [probably Pingwu County:<br />
Long'an; on Fu Jiang river, Jialing Jiang system, Yangtze<br />
drainage; 32°24'31"N 104°31'36"E; see below]; holotype:<br />
MNHN 6287, Bertin & Estève, 1948: 97, fig. 3; noun in<br />
genitive, indeclinable)<br />
Taxonomic notes. Herzenstein (1888: 21), Rendahl (1933:<br />
46) and Prok<strong>of</strong>iev (2003a: 703) discussed ZISP 4471 received<br />
from MNHN as N. nudus from Sichuan. Fang (1941:<br />
253) explained that this specimen had been taken from a jar<br />
labelled "Nemacheilus nudus Blkr.; Sse-chuan Occid.;<br />
R. P. A. David, 1870–58". The remaining 11 specimens from<br />
this jar (MNHN 6287) are the types <strong>of</strong> Nemacheilus angeli.<br />
In Sichuan, David visited Chengdu, then travelled to Muping.<br />
Muping is now in Western Sichuan, but at David's time<br />
it was an independant princedom and David's map shows it<br />
in Tibet (see Oreias dabryi). After returning to Chengdu,<br />
David wanted to visit Kokonoor [now China: Qinghai]. He<br />
reached to Lon-gan [Sichuan: Pingwu County; also as Lunyan,<br />
Lung-yan, Lungan; 32°24'31"N 104°31'36"E], explored<br />
that area, returned to Chengdu and Shanghai (David, 1871:<br />
97). This makes Pingwu the most likely type locality for<br />
N. angeli. Unfortunately, David did not published a detailed<br />
account <strong>of</strong> this part <strong>of</strong> his second travel. His map (1875)<br />
shows that he made a number <strong>of</strong> small trips around Pingwu.<br />
10.42.7 Triplophysa anterodorsalis Zhu & Cao, in Zhu,<br />
1989<br />
Triplophysa anterodorsalis Zhu & Cao, in Zhu, 1989: 106,<br />
fig. 73 (type locality: China: Sichuan: a branch <strong>of</strong> Jin-<br />
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10.42.9 Triplophysa arnoldii, CMK 19546, 71.0 mm SL; Mongolia:<br />
Lake Ayrag basin.<br />
shajiang River [Yangtze] in Huidongxian County<br />
[26°38'04"N 102°34'41"E]; holotype: NGI ? 60VIII0084;<br />
compound noun, indeclinable)<br />
10.42.8 Triplophysa aquaecaeruleae Prok<strong>of</strong>iev, 2001<br />
Triplophysa aquaecaeruleae Prok<strong>of</strong>iev, 2001a: 195, fig. 7<br />
[not fig. 1] (type locality: China: Qinghai: Ba-tshu River<br />
(? Ba Qu), a tributary <strong>of</strong> Blue River (Changjiang<br />
[Yangtze]) (near Yushu, 33°N 92°E) [Yushu town, Gyêgu<br />
Prefecture, 33°00'11"N 97°00'36"E; Jerku on Kozlov's<br />
1902a map]; holotype: ZISP 12494, He et al., 2008: 48,<br />
fig. 5b; noun in genitive, indeclinable)<br />
Taxonomic notes. The coordinates for the type locality<br />
(Yushu) are given in the original description as about 33°N<br />
92°E. This seems erroneous since this is not within the area<br />
visited by Kozlov (the collector) in 1900 (maps in Kozlov,<br />
1948). There is Yushu in Gyêgu Prefecture, Qinghai, at<br />
33°00'11"N 97°00'36"E. The 'Ba Tschu' <strong>of</strong> Kozlov's (1948)<br />
map <strong>of</strong> Kam enters a Gou-tschou [? difficult to decipher on<br />
map] at about 32°52'33"N 97°04'15"E, based on Google<br />
Earth images.<br />
10.42.9 Triplophysa arnoldii Prok<strong>of</strong>iev, 2006<br />
Triplophysa arnoldii Prok<strong>of</strong>iev, 2006b: 237, figs. 1–2 (type<br />
locality: Mongolia: stream connecting Lakes Ayrag-Nuur<br />
and Khirgis-Nuur [Khyar-gas], Kobdo basin [Khovd Gol;<br />
48°56'20"N 93°20'42"E]; holotype: ZISP 29054; noun<br />
in genitive, indeclinable)<br />
10.42.10 Triplophysa bashanensis Xu & Wang, 2009<br />
Triplophysa bashanensis Xu & Wang, 2009: 381, figs. 2–4<br />
(type locality: China: Shaanxi: Xixiang County (32°35'N<br />
107°27'E): Dahe Town: Huangjiabian stream in Loufang<br />
Village, a tributary <strong>of</strong> Jialingjiang River, 712 masl; holotype:<br />
SIZ 0608008; adjective, -is, -is, -e)<br />
Triplophysa longchiensis Xu & Wang, 2009: 384 (name apparently<br />
erroneously used instead <strong>of</strong> T. bashanensis Xu<br />
& Wang, 2009: 381 in English translation <strong>of</strong> text; formally<br />
a simultaneous objective synonym; as first reviser I<br />
give precedence to T. bashanensis; adjective, -is, -is, -e)<br />
10.42.11 Triplophysa bellibarus (Tchang, Yueh & Hwang,<br />
1963)<br />
Nemachilus bellibarus Tchang, Yueh & Hwang, 1963: 625,<br />
figs. 4–7 (type locality: China: Southern Tibet: Tso-Hwui-<br />
La, upper reaches <strong>of</strong> Nyang-Chu [a tributary <strong>of</strong> Brahmaputra];<br />
holotype: ASIZB TS-409; compound noun,<br />
indeclinable ['Latinized' English nouns belly and bar])<br />
Taxonomic notes. Treated as a synonym <strong>of</strong> T. microps by<br />
Wu & Wu (1992: 196); not listed among synonyms <strong>of</strong> T.<br />
microps by Prok<strong>of</strong>iev (2010: 895).
10.42.12 Triplophysa bleekeri (Sauvage & Dabry de Thiersant,<br />
1874)<br />
Nemachilus Bleekeri Sauvage & Dabry de Thiersant, 1874:<br />
15 (type locality: China: southern Shen-si [Shaanxi]:<br />
Yenkiatsoun [or Yen-kia-tsong; David, 1875: map; Baoji<br />
Prefecture, apparently Mei County [34°17'N 107°45'E];<br />
in Tsang-Yu torrent, Wei He system, Huang He drainage;<br />
see below]; syntypes: MNHN 7890 [1], B.2642 [7], Bertin<br />
& Estève, 1948: 98, fig. 2; noun in genitive, indeclinable)<br />
Taxonomic notes. The holotype is 25.0 mm SL and the original<br />
description does not include usable information for identification.<br />
The name is presently applied for a Chinese species<br />
known from the Han Jiang, Jialing Jiang and Min Jiang<br />
in the Yangtze drainage in Shaanxi and Sichuan provinces<br />
(He et al., 2008: 49). In fact, the type locality is in the Huang<br />
He drainage and the identity <strong>of</strong> the species identified as<br />
T. bleekeri should be re-examined.<br />
There has been speculations about the location <strong>of</strong> the<br />
type locality, which could not be identified. This information<br />
is given by the collector in the description <strong>of</strong> his travel<br />
(David, 1875) and on the accompanying map. Yenkiatsoun<br />
[or Yen-kia-tsong on map] is (or is near) present Mei County<br />
in Baoji Prefecture, Shaanxi, near Wei He river, in Huang<br />
He drainage. David (1875: vol. 1: 267) collected "a Gobius<br />
and another small fish" in the torrent Tsang-yu in Yenkiatsoun<br />
on 31 January 1873. Six small "ablettes" (probably a<br />
small cyprinid since ablette is the French word for Alburnus,<br />
a genus <strong>of</strong> silvery European cyprinids) were caught later<br />
(p. 268).<br />
10.42.13 Triplophysa brachyptera (Herzenstein, 1888)<br />
Nemachilus robustus brachypterus Herzenstein, 1888: 40,<br />
pl. 5 fig. 1 (type locality: China: Gan-ssu [Gansu]; syntypes:<br />
ZISP 7320 [2]; compound adjective, -us, -a, -um)<br />
? Nemachilus minxianensis Wang & Zhu, 1979: 129, fig. 1<br />
(type locality: China: Gansu: upper drainage <strong>of</strong> Tau-He<br />
River, branch <strong>of</strong> Huang He; syntypes: IHB 771465 [1],<br />
771372 [1]; adjective, -is, -is, -e)<br />
Taxonomic notes. Tentative synonymy follows Prok<strong>of</strong>iev<br />
(2007b: 4, 2010: 895).<br />
10.42.14 Triplophysa brahui (Zugmayer, 1912)<br />
Nemachilus brahui Zugmayer, 1912: 598 (type locality: Pakistan:<br />
Balochistan: Kelat [Kalat, Qalat; 29°01'33"N<br />
66°35'24"E; apparently in Pishin Lora River endorheic<br />
basin]; syntypes: ZSM [24, lost; Kottelat, 1988a, pers.<br />
obs.]; noun in genitive, indeclinable)<br />
10.42.15 Triplophysa brevibarba Ding, 1993<br />
Triplophysa brevibarba Ding, 1993: 249, fig. 5 (type locality:<br />
China: Sichuan: Mianning County, 28°34'N 102°10'E;<br />
holotype: MSINR 900264; compound noun, indeclinable)<br />
Triplophysa dingi Prok<strong>of</strong>iev, 2010: 842, 895 (unnecessary<br />
replacement name for Triplophysa brevibarba Ding,<br />
1993: 249; noun in genitive, indeclinable)<br />
Taxonomic notes. Prok<strong>of</strong>iev (2010: 895) considered that T.<br />
brevibarba Ding, 1993 is a secondary junior homonym <strong>of</strong><br />
Nemachilus yarkandensis brevibarbus Herzenstein, 1888:<br />
78 [now in Hedinichthys] and proposed T. dingi as new re-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
124<br />
placement name. However, to be secondary homonyms the<br />
two names should have been used in combination with the<br />
same generic name (Code art. 57.3.1). This means that it is<br />
not enough that N. y. brevibarbus has been placed in Triplophysa<br />
as a synonym <strong>of</strong> T. yarkandensis. It is a senior synonym<br />
only if the combination Triplophysa brevibarba<br />
(Herzenstein) or T. yarkandensis brevibarba (Herzenstein)<br />
has been used. To my knowledge, N. y. brevibarbus Herzenstein<br />
has never been treated as a valid species in Triplophysa<br />
and the combination has never been used. Therefore<br />
T. dingi is an unnecessary replacement name.<br />
Note that brevibarbus is an adjective and declinable and<br />
brevibarba is a compound noun and indeclinable.<br />
10.42.16 Triplophysa brevicauda (Herzenstein, 1888)<br />
Nemachilus Stoliczkae brevicauda Herzenstein, 1888: 23,<br />
pl. 5 fig. 4 (type locality: "Dabsun-Gobi" [apparently<br />
China: Qinghai: Dabsan Nur Lake, 56°58'N 95°00'E];<br />
lectotype: ZISP 7370, designated by Prok<strong>of</strong>iev, 2007b:<br />
12, fig. 5a; compound noun, indeclinable)<br />
10.42.17 Triplophysa cakaensis Cao & Zhu, 1988<br />
Triplophysa cakaensis Cao & Zhu, 1988a: 201, figs. 1–3<br />
(type locality: China: Qinghai: Ulanxian County: stream<br />
in Cakayanhu Lake [36°42'00"N 99°06'00"E] endorheic<br />
basin near Caka, "36°N 98°E"; holotype: IHB 56IX0034;<br />
adjective, -is, -is, -e)<br />
10.42.18 Triplophysa chondrostoma (Herzenstein, 1888)<br />
Nemachilus chondrostoma Herzenstein, 1888: 36, pl. 3 fig. 7<br />
(type locality: China: Qinghai: Zaidam [Quaidam basin]:<br />
Bajan-gol River [37°18'N 96°50'E]; holotype: ZISP 7219,<br />
Prok<strong>of</strong>iev, 2003b: 53, fig. 1; compound noun, indeclinable)<br />
Taxonomic notes. Placed in Hedinichthys by Wu & Wu<br />
(1992: 263).<br />
10.42.19 Triplophysa coniptera (Turdakov, 1954)<br />
Nemachilus conipterus Turdakov, 1954: 117 (type locality:<br />
Kyrgyzstan: Ters River, Talas basin; syntypes [? 34]: LU;<br />
compound adjective, -us, -a, -um)<br />
Nemachilus conipterus salari Turdakov, 1954: 122 (type<br />
locality: Uzbekistan: Salar canal in Tachkent [41°16'N<br />
69° 13'E; Syr Darya drainage]; types: LU; noun in genitive,<br />
indeclinable)<br />
10.42.20 Triplophysa crassilabris Ding, 1994<br />
Triplophysa crassilabris Ding, 1994: 92, fig. 56 (type locality:<br />
China: Sichuan: Xia Man Lake, 3400 masl; holotype:<br />
MSINR 830578; compound noun, indeclinable)<br />
10.42.21 Triplophysa cuneicephala (Shaw & Tchang,<br />
1931)<br />
Barbatula cuneicephalus Shaw & Tchang, 1931: 81, fig. 10<br />
(type locality: China: San-kai-tien, near Peiping [Beijing];<br />
holotype: ZMFIB 6237; compound adjective, -us, -a, -um)<br />
10.42.22 Triplophysa dalaica (Kessler, 1876)<br />
Diplophysa dalaica Kessler, 1876: 24, pl. 3 fig. 1 (type locality:<br />
China: Nei Mongol: Lake Dalai Nor [43°18'N
116°40'E]; lectotype: ZISP 2474, designated by Prok<strong>of</strong>iev,<br />
2003b: 55, fig. 5; adjective, -us, -a, -um)<br />
Nemachilus djaggasteensis Rendahl, 1922: 1 (type locality:<br />
China: Nei Mongol: Djaggaste [Tjaggan Obo, Obor Jalag,<br />
Wu-pu-lin-cha-la-ka, 46.33 120.75] [about 120 km on<br />
road from Kalgan [Zhangjiakou] to Urga [Ulaan Baatar],<br />
41.7039°N 113.839°E; NRM catalogue]; syntypes:<br />
NRM 10338 [18], 20338 [1], 18946 [1], 27321 [5],<br />
MNHN 1933.116 [1], Bertin & Estève, 1948-98; adjective,<br />
-is, -is, -e)<br />
Barbatula toni posteroventralis Nichols, 1925b: 4 (type<br />
locality: China: Shansi: Chin Ssu, 16 miles southwest <strong>of</strong><br />
Taiyuanfu; holotype: AMNH 8410; compound noun, indeclinable)<br />
Taxonomic notes. Nemachilus djaggasteensis and B. posteroventralis<br />
are listed as synonyms <strong>of</strong> T. dalaica by Zhu<br />
(1989: 82). They are not mentioned by Prok<strong>of</strong>iev (2003b,<br />
2010: 895).<br />
10.42.23 Triplophysa daqiaoensis Ding, 1993<br />
Triplophysa daqiaoensis Ding, 1993: 247, fig. 1 (type locality:<br />
China: Sichuan: Minnin County: Daqiao, 28°41'N 102°<br />
12'E; holotype: MSINR 900267; adjective, -is, -is, -e)<br />
10.42.24 Triplophysa dorsalis (Kessler, 1872)<br />
Cobitis dorsalis Kessler, 1872: 67, pl. 11 figs. 34–35 (type<br />
locality: Kazakhstan: Yani-Kurgan [Yanykurgan, Zhanakorgan,<br />
43°54'54"N 67°14'53"E; on Syr-Darya]; syntypes<br />
[8]: ZISP 2079 [2], 3702 [according to Herzenstein, 1888:<br />
20]; noun in apposition, indeclinable)<br />
? N[emacheilus] (Deuterophysa) dorsalis valerii Mitr<strong>of</strong>anov,<br />
2002: 124 (not available: no fixation <strong>of</strong> holotype or syntypes,<br />
Code art. 16.4.1, no intention to establish a new<br />
taxon, Code art. 16.1; locality: Kazakhstan: Ili drainage,<br />
Bolshaya Almatinka River [43°34'01"N 76°54'03"E];<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. The type locality <strong>of</strong> C. dorsalis given by<br />
Kessler is Yani-Kurgan. Berg (1949: 858) gave it as "Yanykurgan,<br />
in the Chimkent or the Dzhizak District ?". Chimket<br />
is in Kazakhstan [Shymkent, 42°19'00"N 69°35'45"E] and<br />
Dzhizak is in Uzbekistan [Djizak, Jizzax, Jizzakh, 40°06'<br />
57"N 67°50'32"E].<br />
10.42.25 Triplophysa dorsonotata (Kessler, 1879)<br />
Nemachilus dorsonotatus Kessler, 1879: 285 (type locality:<br />
China: Xinjiang: Kungès River [Künes He], a tributary<br />
<strong>of</strong> Ili River, alt. 4000 ft.; lectotype: ZISP 4175 [1 <strong>of</strong> 2],<br />
listed by Prok<strong>of</strong>iev, 2007b: 4, 6, 8, but not formally designated,<br />
this specimen is here designated as lectotype;<br />
reprinted as Kessler, 1880: 271; adjective, -us, -a, -um)<br />
? Nemachilus cuëljuensis Anikin, 1905: 6, 18 [<strong>of</strong> reprint]<br />
(type locality: Kyrgyzstan: Issyk Kul Province: Ak-Suu<br />
District: Kuelyu River [Kuilyu; 42°12'16"N 78°57'37"E],<br />
tributary <strong>of</strong> Sarydzhas [Sary-Dzhaz, Sarychat], Tarim<br />
basin [flows to Aksu-He and Yarkand in Xinjiang, China];<br />
syntypes [3]: LU; adjective, -is, -is, -e)<br />
10.42.26 Triplophysa drassensis (Tilak, 1990)<br />
Noemacheilus drassensis Tilak, 1990: 136, fig. 1 (type locality:<br />
India: Jammu and Kashmir: Ladakh: stream near<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
125<br />
Drass [34°25'46"N 75°45'07"E]; holotype: ZSI/NRS<br />
1206; paper dated 1990 but possibly not published until<br />
1997; adjective, -is, -is, -e)<br />
Nomenclatural notes. See Triplophysa shehensis for publication<br />
date.<br />
10.42.27 ? Triplophysa elegans (Kessler, 1874)<br />
Cobitis elegans Kessler, 1874: 43 (type locality: Uzbekistan:<br />
Salar stream near Tachkent [41°16'N 69° 13'E; Syr<br />
Darya drainage]; syntypes: ZISP 10931 [1; also listed<br />
under C. uranoscopus], ZMMU P2542 [1], P-8002 [3],<br />
Svetovidova, 1978: 259, Vasilieva et al., in Pavlinov &<br />
Borissenko, 2001: 28; adjective, indeclinable)<br />
Taxonomic notes. Listed as synonym <strong>of</strong> T. stolickai by Prok<strong>of</strong>iev<br />
(2007b: 4), which geography suggests should be reexamined.<br />
10.42.28 Triplophysa farwelli (Hora, 1935)<br />
Nemachilus farwelli Hora, 1935b: 799, fig. 4d, pl. 1 fig. 2<br />
(type locality: Afghanistan: Helmand River; holotype: ZSI<br />
F 11515/1; noun in genitive, indeclinable)<br />
10.42.29 Triplophysa flavicorpus Yang, Chen & Lan, 2004<br />
Triplophysa flavicorpus Yang, Chen & Lan, 2004: 113, fig. 3<br />
(type locality: China: Guangxi: Du'an County: Hongshuihe<br />
River, a tributary <strong>of</strong> Xijiang; holotype: KIZ<br />
995004; compound noun, indeclinable)<br />
10.42.30 Triplophysa furva Zhu, 1992<br />
Triplophysa furva Zhu, 1992: 242, figs. 3–6 (type locality:<br />
China: Xinjiang: Urümqi River, near Hongyanchi Reservoir,<br />
Urümqi City, 43°47'N 87°37'E; holotype: NGI<br />
880943; adjective, -us, -a, -um)<br />
10.42.31 Triplophysa fuxianensis Yang & Chu, 1990<br />
Triplophysa fuxianensis Yang & Chu, 1990b: 377, fig. 2 (type<br />
locality: China: Yunnan: Fuxian Lake [24°32'N<br />
103°55'E]; holotype: KIZ 873126; adjective, -is, -is, -e)<br />
10.42.32 Triplophysa gejiuensis (Chu & Chen, 1979)<br />
Noemacheilus gejiuensis Chu & Chen, 1979: 285, fig. 1 (type<br />
locality: China: Yunnan: Gejiu County: Bajianjing near<br />
Qiafanf [Bajiaojing 4 km from Kafang, 23°16'N<br />
103°09'E; Romero et al., 2009: 260]; holotype: KIZ<br />
7803001; adjective, -is, -is, -e)<br />
10.42.33 Triplophysa gerzeensis Cao & Zhu, 1988<br />
Triplophysa gerzeensis Cao & Zhu, 1988: 202, figs. 4–5 (type<br />
locality: China: Qinghai–Xizang, stream in Chaco Lake<br />
drainage near Gerze [32°18'15"N 84°03'30"E], 4370<br />
masl; holotype: IHB 76IV0200; adjective, -is, -is, -e)<br />
10.42.34 Triplophysa gracilis (Day, 1877)<br />
Nemacheilus gracilis Day, 1877b: 798 (type locality: India:<br />
Ladakh: Basgo, on headwaters <strong>of</strong> Indus River [34°13'N<br />
77°17'E; downstream <strong>of</strong> Leh]; syntypes: ZSI 1469,<br />
BMNH 1889.2.1.1726 [1], NMW 48413 [1], Whitehead<br />
& Talwar, 1976: 157; also in Day, 1878b: 16, pl. 4 fig. 5;<br />
adjective, -is, -is, -e)<br />
? Nemachilus deTerrai Hora, 1936: 311, figs. 4–5, pl. 12
10.42.37 Triplophysa gundriseri, CMK 19548, 96.4 mm SL; Mongolia:<br />
Lake Uvs basin.<br />
fig. 5–6 (type locality: India: Kashmir: Ladakh: Man<br />
Lagoon; syntypes [9]: ZSI F 12200/1 [6], Menon &<br />
Yazdani, 1968: 122; noun in genitive, indeclinable)<br />
? Triplophysa trewavasae Mirza & Ahmad, 1990: 317, fig. 1<br />
(type locality: Pakistan: Northern Area: near rest house<br />
<strong>of</strong> Yasin, Yasin Valley [Yasin: 36°22'N 73°20'E]; holotype:<br />
GCMNH uncat.; noun in genitive, indeclinable)<br />
Taxonomic notes. Tentative synonymy, following Prok<strong>of</strong>iev<br />
(2010: 895).<br />
10.42.35 Triplophysa grahami (Regan, 1906)<br />
Nemachilus grahami Regan, 1906b: 333 (type locality: China:<br />
Yunnan: Yunnan-Fu [Kunming]; syntypes: BMNH<br />
1905.10.28.16–17 [2]; noun in genitive, indeclinable)<br />
10.42.36 Triplophysa griffithii (Günther, 1868)<br />
Nemachilus griffithii Günther, 1868: 360 (type locality: "Assam<br />
?" [error, see Hora, 1928: 481; correct locality is:<br />
Afghanistan: Arghandab, a rapid and considerable sized<br />
tributary <strong>of</strong> Helmand River, running within two or three<br />
miles <strong>of</strong> Kandahar]; syntypes: BMNH 1860.3.19.93–94<br />
[2], lost ?, Kullander et al., 1999: 142; noun in genitive,<br />
indeclinable)<br />
Noemacheilus naziri Ahmad & Mirza, 1963: 76, figs. 1–2<br />
(type locality: Pakistan: Swat State: Madyan stream near<br />
Madyan [35°08'25"N 72°32'14"E]; syntypes [9]: ISBB<br />
1234 [3], Bănărescu & Nalbant, 1966a: 159; noun in<br />
genitive, indeclinable)<br />
Nemacheilus griffithi hazaraensis Omer & Mirza, 1975: 203,<br />
figs. 1–2 (type locality: Pakistan: Hazara District [now<br />
Abbottabad]: Ilyasi Masjid springs [34°10'16"N<br />
73°15'30"E; in Nawanshehr]; holotype: GCMNH F 12;<br />
adjective, -is, -is, -e)<br />
Taxonomic notes. Günther (1868: 360) indicated the type<br />
locality <strong>of</strong> T. griffithii as "Assam ?". Hora (1928: 481) showed<br />
that this is not correct and that the type locality is near Kandahar,<br />
Afghanistan.<br />
10.42.37 Triplophysa gundriseri Prok<strong>of</strong>iev, 2002<br />
Nemacheilus dorsalis humilis Gundriser, 1962: 253, fig. 4<br />
(type locality: Russia: Tuva Republic: Tes-Khem River,<br />
25 km northwest <strong>of</strong> Erzin, 50°27'N 95°01'E [original type<br />
locality: Russia: Tuva: Erzin and Tes-Khem Rivers]; neotype:<br />
ZISP 52053, designated by Prok<strong>of</strong>iev, 2002a: S49,<br />
fig. 4a; junior primary homonym <strong>of</strong> Nemachilus humilis<br />
Lin, 1932c: 515; adjective, -is, -is, -e)<br />
Triplophysa gundriseri Prok<strong>of</strong>iev, 2002a: S47 (replacement<br />
name for Nemacheilus dorsalis humilis Gundriser, 1962:<br />
253; noun in genitive, indeclinable)<br />
Triplophysa gundriseri chandagaitensis Prok<strong>of</strong>iev, 2002a:<br />
S55, fig. 4a (type locality: Russia: Tuva Republic: Chan-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
126<br />
dagaity River in Chandagaity village, 50°44'N 92°08'E;<br />
holotype: ZISP 52199; adjective, -is, -is, -e)<br />
10.42.38 Triplophysa hexiensis (Zhao & Wang, 1988)<br />
Nemachilus dorsonotatus hexiensis Zhao & Wang, 1988:<br />
113, fig. 2 (type locality: China: Gansu: Hexi area, Ruoshui<br />
and Shiyang He rivers [Ruo Shui: Edsin, Ejin, Ejina<br />
He, Eiin, Hei-He; a river flowing north to endorheic<br />
Gaxun Nuur (Juyan Lake), 42°25'20"N 100°40'07"E;<br />
Shiyang He: an endorheic river disappearing in Gobi<br />
desert about 39°00'N 103°35'E]; syntypes [28]: DBLU<br />
77-0940 [?], 0946, 0953, 0955, 80-0344, 0357, 0362,<br />
0373, 0375, 0383, 0386, 0390, 0394, 0406, 0420, 0435,<br />
0461, 0468, 0563, 0803, 1192–1195, 1197, 1198, 1135<br />
[27]; adjective, -is, -is, -e)<br />
? Triplophysa rossoperegrinatorum Prok<strong>of</strong>iev, 2001a: 200,<br />
fig. 19 [not fig. 13] (type locality: China: Gansu: mountains<br />
north <strong>of</strong> Xining & Lanzhou; holotype: ZISP 3719;<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. Synonymy follows Prok<strong>of</strong>iev (2010: 895).<br />
10.42.39 Triplophysa heyangensis Zhu, 1992<br />
Triplophysa heyangensis Zhu, 1992: 244, figs. 7–10 (type<br />
locality: China: Shaanxi: Heyangxian County: Hanshutaucun<br />
village, 35°11'N 110°05'E, Huang He drainage;<br />
holotype: NGI 820027; adjective, -is, -is, -e)<br />
10.42.40 Triplophysa hialmari Prok<strong>of</strong>iev, 2001<br />
Triplophysa hialmari Prok<strong>of</strong>iev, 2001a: 198, fig. 1 [not 7]<br />
(type locality: China: Qinghai: Ba-tshu River (? Ba Qu),<br />
a tributary <strong>of</strong> Blue River (Changjiang [Yangtze]) (near<br />
Yushu, 33°N 92°E) [Yushu town, Gyêgu Prefecture,<br />
33°00'11"N 97°00'36"E]; holotype: ZISP 26245; noun<br />
in genitive, indeclinable)<br />
Taxonomic notes. See discussion <strong>of</strong> type locality under<br />
T. aquaecaeruleae.<br />
10.42.41 Triplophysa hsutschouensis (Rendahl, 1933)<br />
Nemacheilus hsutschouensis Rendahl, 1933: 41 (type locality:<br />
China: Nei Mongol: Hsutschou, in a stream flowing<br />
to Etsin Gol [Edsin, Ejin, Ejina He, Eiin, Hei-He, Ruo-<br />
Shui; a river flowing north to endorheic Gaxun Nuur (Juyan<br />
Lake), 42°25'20"N 100°40'07"E]; syntypes: NRM<br />
10206 [1], 12339 [7]; adjective, -is, -is, -e)<br />
10.42.42 Triplophysa huanglongensis Gao, in Anonym,<br />
1992<br />
Triplophysa huanglongensis Gao, in Anonym, 1992: 83, fig.<br />
3-87 (type locality: China: Shaanxi: Huang Long County:<br />
Baima village, upper reaches <strong>of</strong> Ju He; holotype: Fisheries<br />
Institute <strong>of</strong> Shaanxi Province, Xian; adjective, -is,<br />
-is, -e)<br />
10.42.43 Triplophysa huanjiangensis Yang, Wu & Lan,<br />
2011<br />
Triplophysa huanjiangensis Yang, Wu & Lan, 2011: 567,<br />
figs. 1, 3 (type locality: China: Guangxi: Huanjiang County<br />
[Nandan County on p. 569]: Chuanshan Town, cave in<br />
Leyi village (25°06'02.6"N 108°00'01"E); holotype: GIF<br />
07040316; spelt huangjiangensis p. 566, an obvious er-
or <strong>of</strong> inadvertence, thus incorrect original spelling [Code<br />
art. 32.5.1]; adjective, -is, -is, -e)<br />
10.42.44 Triplophysa huapingensis Zheng, Yang & Chen,<br />
2012<br />
Triplophysa huapingensis Zheng, Yang & Chen, 2012a: 832,<br />
fig. 1 (type locality: China: Guangxi: Leye County: Huaping<br />
town, Hongshuihe River, a tributary <strong>of</strong> Xijiang; holotype:<br />
KIZ 2008007607; adjective, -is, -is, -e)<br />
10.42.45 Triplophysa hutjertjuensis (Rendahl, 1933)<br />
Nemacheilus hutjertjuensis Rendahl, 1933: 28 (type locality:<br />
China: Nei Mongol: stream close to Hutjentju Göl;<br />
syntypes [12]: NRM 10202 [10], MNHN 1933.118 [1],<br />
Bertin & Estève, 1948: 98; adjective, -is, -is, -e)<br />
10.42.46 Triplophysa intermedia (Kessler, 1876)<br />
Diplophysa intermedia Kessler, 1876: 28, pl. 3 fig. 4 [not 3]<br />
(type locality: China: Nei Mongol: Lake Dalaï-Nor [43°<br />
18'N 116°40'E; not Hu-Lun Lake, which is another Dalai<br />
Nur]; holotype: ZISP 2476; adjective, -us, -a, -um)<br />
10.42.47 Triplophysa jianchuanensis Zheng, Du, Chen &<br />
Yang, 2010<br />
Triplophysa jianchuanensis Zheng, Du, Chen & Yang, 2010:<br />
22, fig. 1 (type locality: China: Yunnan: Lancangjiang<br />
drainage [Mekong]: Dali Prefecture: Jianchuan County:<br />
Jiangkou spring, 26°31'22.6"N 99°57'30.4"E; holotype:<br />
KIZ 2008004424; adjective, -is, -is, -e)<br />
10.42.48 Triplophysa jiarongensis Lin, Li & Song, 2012<br />
Triplophysa jiarongensis Lin, Li & Song, 2012: 640, figs. 2–<br />
4 (type locality: China: Guizhou: Libo County: Jiarong<br />
Town, Shuijingwan Cave, 616 masl, 25°28'N 108° 06'E;<br />
holotype: SOU 10111801; adjective, -is, -is, -e)<br />
10.42.49 Triplophysa kafirnigani (Turdakov, 1947)<br />
Nemachilus dorsalis kafirnigani Turdakov, 1947a: 58 (type<br />
locality: Tajikistan: Kafirnigan River near Stalinabad<br />
[Dushanbe], Amu Darya drainage; syntypes [16]: ZMMU<br />
P-3647 [1], P-3651 [3], Vasilieva et al., in Pavlinov &<br />
Borissenko, 2001: 28; noun in genitive, indeclinable)<br />
10.42.50 Triplophysa kullmanni Banarescu & Nalbant,<br />
in Ladiges, 1975<br />
Triplophysa kullmanni Bănărescu & Nalbant, in Ladiges,<br />
1975: 40 (type locality: Central Afghanistan: Ab-e-Nawar<br />
[Dasht-e-Nawar, a lake west <strong>of</strong> Ghazni; 33°31'43"N<br />
67°48'06"E]; holotype: ZMH H 4778; noun in genitive,<br />
indeclinable)<br />
Nomenclatural notes. The original description <strong>of</strong> this species<br />
by Bănărescu & Nalbant (1975: 246, pl. 5 fig. 5) appeared<br />
in November 1975, but a popular account by Ladiges<br />
based on Bănărescu & Nalbant's manuscript appeared in<br />
February 1975. As this account includes illustrations and a<br />
short description, it makes the name available.<br />
10.42.51 Triplophysa kungessana (Kessler, 1879)<br />
Diplophysa kungessana Kessler, 1879: 286 (type locality:<br />
China: Xinjiang: Kungès River [Künes He, tributary <strong>of</strong><br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
127<br />
Ili River], alt. 4000 ft.; holotype: ZISP 4176; reprinted<br />
as Kessler, 1880: 238; adjective, -us, -a, -um)<br />
10.42.52 Triplophysa lacusnigri (Berg, 1931)<br />
Nemachilus stoliczkai var. lacus-nigri Berg, 1931a: 25, pls.<br />
3–5 (type locality: Tajikistan: Lake Karakul and Karaart<br />
River; syntypes [131+]: ZISP 26367 [6+], 26368 [6+],<br />
26369 [6+], Eschmeyer & Fricke, 2010, Prok<strong>of</strong>iev,<br />
2007b: 10, fig. 4f; noun in genitive, indeclinable)<br />
10.42.53 Triplophysa lacustris Yang & Chu, 1990<br />
Triplophysa lacustris Yang & Chu, 1990b: 380, fig. 3 (type<br />
locality: China: Yunnan: Xinyun Lake [24°20'N 102°<br />
46'E], Nanpanjiang drainage; holotype: KIZ 0.1399; adjective,<br />
-is, -is, -e)<br />
10.42.54 Triplophysa laterimaculata Li, Liu & Yang, 2007<br />
Triplophysa laterimaculata Li, Liu & Yang, 2007: 48, fig. 2<br />
(type locality: China: Xinjiang: Kashigar City [Kashgar]:<br />
Shufu County [39°21'00"N 75°51'03"E]: Kezile River, a<br />
tributary <strong>of</strong> Tarim River [39°N 76°E], Huangdi Town;<br />
holotype: KIZ 200408220489; adjective, -us, -a, -um)<br />
10.42.55 Triplophysa laticeps Zhou & Cui, 1997<br />
Triplophysa laticeps Zhou & Cui, 1997: 179, figs. 2–3 (type<br />
locality: China: Yunnan: Lufeng County: Luzhijiang, tributary<br />
<strong>of</strong> Yuanjiang River [upper Red River], near Lufeng<br />
city [25°15'N 102°03'E, 1600 masl]; holotype: KIZ<br />
914077; compound adjective, indeclinable)<br />
10.42.56 Triplophysa lingyunensis (Liao & Luo, in Liao,<br />
Wang & Luo, 1997)<br />
Schistura lingyunensis Liao & Luo, in Liao, Wang & Luo,<br />
1997: 4, fig. 1 (type locality: China: Guangxi: Lingyun<br />
County: cave in Guancang village, 24°25'N 106°30'E;<br />
holotype: IHB 92vi003; adjective, -is, -is, -e)<br />
10.42.57 Triplophysa lixianensis He, Song & Zhang, 2008<br />
Triplophysa lixianensis He, Song & Zhang, 2008: 43, fig. 1<br />
(type locality: China: Sichuan: Aba Prefecture: Lixian<br />
County: upper Yangtze drainage: Zagunao River, a tributary<br />
on western bank <strong>of</strong> Min Jiang, at Jiujiapeng village<br />
(31°33"N 102°52"E), 2450 masl; holotype: SCUM<br />
20070717008; adjective, -is, -is, -e)<br />
10.42.58 Triplophysa longianguis Wu & Wu, 1984<br />
Triplophysa longianguis Wu & Wu, 1984: 327, figs. 1–5<br />
(type locality: China: Qinghai: Jigzhi County: Lake Sunm<br />
Cuo; holotype NPIB 8370294; compound noun, indeclinable)<br />
10.42.59 Triplophysa longibarbata (Chen, Yang, Sket &<br />
Aljancic, 1998)<br />
Paracobitis longibarbata Chen, Yang, Sket & Aljancic, 1998:<br />
59, fig. 1 (type locality: China: Guizhou: Libo County:<br />
cave approx. 13 km northeast <strong>of</strong> Libo town [25°15'N<br />
108°00'E; Romero et al., 2009: 260]; holotype: KIZ<br />
953001; adjective, -us, -a, -um)<br />
Nemacheilus liboensis Ran, 2000: 329, 332 (nomen nudum)<br />
Nemacheilus liboensis Chen, 2000: 41 (nomen nudum)
Paracobitis posterodorsalus Li, Ran & Chen, in Ran, Li &<br />
Chen, 2006: 81, fig. 1 (type locality: China: Guangxi:<br />
Nandan County: cave at 25°02'N 107°36'E [24°59'N<br />
107°32'E; Romero et al., 2009: 256]; holotype: HRAS<br />
041005001; also spelt posterodarsalus p. 80, posterodorsalus<br />
p. 81; as first reviser I select posterodorsalus<br />
as the correct original spelling; compound adjective,<br />
-us, -a, -um)<br />
? Paracobitis maolanensis Li, Ran & Chen, 2006: 1, fig. 1<br />
(type locality: China: Guizhou: Libo County: cave in<br />
Maolan Karst Forestry Natural Reserve, 25°25'04"N<br />
107°52'E [25°23'N 108°04'E; Romero et al., 2009: 255];<br />
holotype: HRAS 050103001; adjective, -is, -is, -e)<br />
Taxonomic notes. Synonymy partly derived from Du et al.<br />
(2008: 33) and Romero et al. (2009), but Yang, Wu & Lan<br />
(2011: 570) treat P. posterodorsalus and P. maolanensis as<br />
valid species <strong>of</strong> Triplophysa.<br />
10.42.60 Triplophysa longipectoralis Zheng, Du, Chen &<br />
Yang, 2009<br />
Triplophysa longipectoralis Zheng, Du, Chen & Yang, 2009:<br />
222, figs. 1–2 (type locality: China: Guangxi: Liujiang<br />
basin: Huanjiang County: cave in Xunle town, Hechi City;<br />
holotype: KIZ 2001004573; compound noun, indeclinable)<br />
10.42.61 Triplophysa macrocephala Yang, Wu & Yang, 2012<br />
Triplophysa macrocephala Yang, Wu & Yang, 2012: 170,<br />
1–2 (type locality: China: Guangxi: Nandan County: Lihu<br />
Town: Renguang village, 25°10'15"N 107°43'50"E<br />
[Xijiang drainage, Pearl River]; holotype: KIZ<br />
2010003078; compound adjective, -us, -a, -um)<br />
10.42.62 Triplophysa macromaculata Yang, in Chu &<br />
Chen, 1990<br />
Triplophysa macromaculata Yang, in Chu & Chen, 1990:<br />
58, fig. 53 (type locality: China: Yunnan: Jiuxiang <strong>of</strong> Yiliang<br />
County, 25°06'N 103°23'E; holotype: KIZ 874021;<br />
compound adjective, -us, -a, -um)<br />
10.42.63 Triplophysa macrophthalma Zhu & Guo, 1985<br />
Triplophysa macrophthalma Zhu & Guo, 1985: 323, fig. 2<br />
(type locality: China: Yunnan: Goujie, Yiliang County<br />
(27°30'N 104°00'E), Nanpanjiang drainage; holotype:<br />
NGI 810019, He et al., 2008: 48, fig. 5e; compound adjective,<br />
-us, -a, -um)<br />
10.42.64 Triplophysa markehenensis (Zhu & Wu, 1981)<br />
Nemachilus pseudoscleropterus markehenensis Zhu & Wu,<br />
1981: 223, fig. 4 (type locality: China: Qinghai: Jigzhi<br />
County [33°26'N 101°26'E]: Marke He, Baiyu; holotype:<br />
NPIB 71005; adjective, -is, -is, -e)<br />
10.42.65 Triplophysa marmorata (Heckel, 1838)<br />
Cobitis marmorata Heckel, 1838: 76, pl. 12 figs. 1–2 (type<br />
locality: India: Kashmir; syntypes [at least 3]: NHMW<br />
48561 [3], Eschmeyer & Fricke, 2010; adjective, -us, -a,<br />
-um)<br />
Cobitis vittata Heckel, 1838: 80, pl. 12 figs. 3–4 (type locality:<br />
India: Kashmir; syntypes: NHMW 48451 [3], Esch-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
128<br />
meyer & Fricke, 2010; simultaneous subjective synonym<br />
<strong>of</strong> Cobitis marmorata Heckel, 1838: 76; first reviser<br />
[Günther, 1868: 356] gave precedence to C. marmorata;<br />
adjective, -us, -a, -um)<br />
? Nemachilus griffithii var. afghana Hora, 1935b: 799, fig. 4c<br />
(type locality: Afghanistan: springs at Car-i-Chashma,<br />
sources <strong>of</strong> Kabul River [Sar-i Chashma, Chashma-i Shir,<br />
Surkh Kotal, 36°02'60"N 68°34'00"'E] [locality <strong>of</strong> holotype;<br />
Menon & Yazdani, 1968: 122]; holotype: ZSI F<br />
11525/1; adjective, -us, -a, -um)<br />
Taxonomic notes. Synonymy follows Kullander et al. (1999:<br />
142).<br />
10.42.66 Triplophysa microphysus (Fang, 1935)<br />
Nemacheilus microphysus Fang, 1935c: 753, figs. 3–5 (type<br />
locality: Kazakhstan: Lepsa [46°20'49"N 78°19'55"E],<br />
Bulenka River, Lake Balkash basin; holotype: ZMB<br />
10277; compound noun [incorrect Latinization <strong>of</strong> physa],<br />
indeclinable)<br />
10.42.67 Triplophysa microphthalma (Liao & Wang, in<br />
Liao, Wang & Luo, 1997)<br />
Schistura dabryi microphthalmus Liao & Wang, in Liao,<br />
Wang & Luo, 1997: 4, fig. 2 (type locality: China:<br />
Guizhou: Weng'an County: Laoying Cave, 26°53'N<br />
107°02'E [Hawk Cave, 26°53'N 107°02'E; Romero et al.,<br />
2009: 258]; holotype: IHB 91x005; secondary junior<br />
homonym <strong>of</strong> Diplophysa microphthalma Kessler, 1879:<br />
308 when both were placed in Triplophysa; never replaced<br />
and not presently treated as congeneric, therefore<br />
replacement not needed [Code art. 59.2]; compound adjective,<br />
-us, -a, -um)<br />
10.42.68 Triplophysa microps (Steindachner, 1866)<br />
Cobitis microps Steindachner, 1866: 794, pl. 13 figs. 3–3a<br />
(type locality: India: Ladakh: Rupshu: Leh and Phirse<br />
stream near Manechan, alt. about 16000 ft.; syntypes [10]:<br />
NMW 48562 [11 ?], Eschmeyer & Fricke, 2010; compound<br />
noun, indeclinable)<br />
10.42.69 Triplophysa moquensis Ding, 1994<br />
Triplophysa moquensis Ding, 1994: 69, fig. 35 (type locality:<br />
China: Sichuan: Xia Man Lake; holotype: MSINR<br />
830833; adjective, -is, -is, -e)<br />
10.42.70 Triplophysa nandanensis Lan, Yang & Chen,<br />
1995<br />
Triplophysa nandanensis Lan, Yang & Chen, 1995: 368,<br />
fig. 2 (type locality: China: Guangxi: Nandan County:<br />
outlet <strong>of</strong> underground river [25°10'N 107°10'E] at Mayang<br />
village, Liuzhai Town [25°18'N 107°24'E; Romero<br />
et al., 2009: 261]; holotype: KIZ 9110011; adjective, -is,<br />
-is, -e)<br />
10.42.71 Triplophysa nanpanjiangensis (Zhu & Cao,<br />
1988)<br />
Oreias dabryi nanpanjiangensis Zhu & Cao, 1988: 98, figs.<br />
7–8 (type locality: China: Yunnan: Zhanyi County: outlet<br />
<strong>of</strong> subterranean waters near Haijiashao; holotype: NGI<br />
818512; adjective, -is, -is, -e)
10.42.72 Triplophysa nasobarbatula Wang & Li, 2001<br />
Triplophysa nasobarbatula Wang & Li, 2001: 98, fig. 1 (type<br />
locality: China: Guizhou: Libo County: Weng'an town,<br />
107°54'E 25°14'N; holotype: ZMCZ 840207; adjective,<br />
-us, -a, -um)<br />
10.42.73 Triplophysa ninglangensis Wu & Wu, 1988<br />
Triplophysa ninglangensis Wu & Wu, 1988a: 19, fig. 3 (type<br />
locality: China: Yunnan: Ninglang County [27°17'02"N<br />
100°51'36"E]: Ninglang River; holotype: NPIB 8609464;<br />
adjective, -is, -is, -e)<br />
10.42.74 Triplophysa nujiangensa Chen, Cui & Yang,<br />
2004<br />
Triplophysa nujiangensa Chen, Cui & Yang, 2004b: 505,<br />
fig. 1 (type locality: China: Yunnan: about 20 km south<br />
<strong>of</strong> Liuku, left bank tributaries <strong>of</strong> Nujiang River [Salween]<br />
in north <strong>of</strong> Shangjiang Xiang, 818 masl; 25°40'59"N<br />
98°52'27"E; holotype: KIZ 20007496; adjective, -us,<br />
-a, -um)<br />
Nomenclatural notes. The name nujiangensa is an incorrect<br />
Latinization but cannot be corrected (Code art. 32.5.1).<br />
It should have been nujiangensis.<br />
10.42.75 Triplophysa obscura Wang, in Chen, Xu, Fang,<br />
Song & Wang, 1987<br />
Triplophysa obscura Wang, in Chen, Xu, Fang, Song &<br />
Wang, 1987: 28, fig. 15 (type locality: China: Gansu:<br />
upper Bailungjiang (tributary <strong>of</strong> Jialingjiang) in Diebu<br />
Xian [Tèwo, 34°00'N 103°33'E]; holotype: SIZX 822270;<br />
adjective, -us, -a, -um)<br />
10.42.76 Triplophysa obtusirostra Wu & Wu, 1988<br />
Triplophysa obtusirostra Wu & Wu, 1988b: 197, figs. 1–5<br />
(type locality: China: small lake at source <strong>of</strong> Kar Qu in<br />
sources <strong>of</strong> Huanghe River, 38°48'N 96°24'E [erroneous,<br />
Kar Qu is at 35°02'N 96°50'E]; holotype: NPIB 856031;<br />
compound noun [incorrect spelling <strong>of</strong> rostrum], indeclinable)<br />
10.42.77 Triplophysa orientalis (Herzenstein, 1888)<br />
Nemachilus kungessanus orientalis Herzenstein, 1888: 44,<br />
pl. 6 fig. 2 (type locality: China: Qinghai: Zaidam [Qaidam<br />
basin]; syntypes: ZISP 7271 [more than 6] "and others";<br />
adjective, -is, -is, -e)<br />
Nemachilus kungessanus elongatus Herzenstein, 1888: 44,<br />
pl. 5 fig. 3 (type locality: China: Qinghai: Qaidam basin:<br />
Nomochun-gol [Nomhon He River; Nuomuhong County:<br />
36°25'N 96°27'E]; syntypes: ZISP 7386/1 [2]; simultaneous<br />
subjective synonym <strong>of</strong> Nemachilus kungessanus<br />
orientalis Herzenstein, 1888: 44; first reviser [apparently<br />
Zhu, 1989: 84] gave precedence to N. k. orientalis;<br />
adjective, -us, -a, -um)<br />
10.42.78 Triplophysa papillosolabiata (Kessler, 1879)<br />
Diplophysa papilloso-labiata Kessler, 1879: 299 (type locality:<br />
China: Xinjiang: Balgantaigol [Balguntay stream;<br />
Baiguntay city: 42°45'49"N 86°19'13"E] [localities <strong>of</strong><br />
paralectotypes: Great Juldus [Bol'shoi Yulduz; Tung<br />
Hung, Hai Tu], Smaller Juldus [Malyi Yulduz] [Yulduz<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
129<br />
10.42.97 Triplophysa siluroides, ZISP 7244, 148 mm SL, syntype;<br />
China: Qinghai: Huang He River. (From Herzenstein, 1888:<br />
pl. 7 fig. 1).<br />
10.42.97 Triplophysa siluroides, ZISP 7244, 148 mm SL. (From<br />
Herzenstein, 1888: pl. 7 fig. 1).<br />
valley is valley <strong>of</strong> Khaidyk-gol (43°N 83–86°E)]]; lectotype<br />
ZISP 4191, designated by Prok<strong>of</strong>iev, 2003b: 58,<br />
fig. 11; must be emended as papillosolabiata, Code art.<br />
32.5.2.3; reprinted as Kessler, 1880: 257; adjective, -us,<br />
-a, -um)<br />
Taxonomic notes. Validity follows Prok<strong>of</strong>iev (2003b).<br />
10.42.79 Triplophysa pappenheimi (Fang, 1935)<br />
Nemacheilus pappenheimi Fang, 1935c: 761, figs. 8–9 (type<br />
locality: China: Qinghai: Si-ning-fu [Xining, 36°37'00"N<br />
101°46'00"E]; holotype: ZMB 16462; noun in genitive,<br />
indeclinable)<br />
10.42.80 Triplophysa paradoxa (Turdakov, 1955)<br />
Nemacheilus paradoxus Turdakov, 1955: 153 (type locality:<br />
Kyrgyzstan: Karakol [river running through Issyk Kul<br />
lake at 2°36'N 78°21'E], in Talas basin / Kyrgyzstan &<br />
Kazakhstan: Talas River basin; syntypes [25]: ZMMU<br />
P-8797 [1], Vasilieva et al., in Pavlinov & Borissenko,<br />
2001: 26; treated as adjective, -us, -a, -um)<br />
10.42.81 Triplophysa parva Chen, Li & Yang, 2009<br />
Triplophysa parvus Chen, Li & Yang, 2009: 86, figs. 2, 3a<br />
(type locality: China: Yunnan: Yiliang County: Jialonghe<br />
River, 24°57'N 103°07'E, altitute 1770 m, flowing from<br />
Yangzonghai Lake to Nanpangjiang; holotype: KIZ<br />
750413007; adjective, -us, -a, -um)<br />
10.42.82 Triplophysa "pedaschenkoi" (Berg, 1931)<br />
Diplophysa strauchi ulacholica var. pedaschenkoi Berg,<br />
1931b: 312, fig. 2 (an infrasubspecific name, not available,<br />
Code arts. 1.3.4, 45.5; locality: Kyrgyzstan: Lake<br />
Issyk-kul: Karasu Bay near Karakol harbour, Akterek Bay
and at mouth <strong>of</strong> Ulachol stream; material: ZISP)<br />
Nomenclatural notes. Because it is infrasubspecific, the<br />
name D. s. u. var. pedaschenkoi is excluded from zoological<br />
nomenclature (Code arts. 1.3.4, 45.5) and is not available<br />
from Berg (1931b). If the name has been used later as the<br />
valid name <strong>of</strong> a species or subspecies, then it is available<br />
from that next usage (Code art. 45.5.1). I found only one<br />
usage <strong>of</strong> the name as a valid species, by Prok<strong>of</strong>iev (2006b:<br />
257), but there might be earlier uses. To be available from a<br />
next usage as a species-group name, Code art. 45.5.1 requires<br />
that it satisfies arts. 11–18. Article 16 requires that<br />
for names published after 1999 to be available, the author<br />
must explicitly indicate it is intentionally new. This was not<br />
the case in Prok<strong>of</strong>iev (2006b). If there were an earlier use<br />
before 2000, it does not need to be indicated and the name is<br />
automatically made available.<br />
10.42.83 Triplophysa polyfasciata Ding, in Ding, Fang &<br />
Fang, 1996<br />
Triplophysa polyfasciata Ding, in Ding, Fang & Fang, 1996:<br />
11, fig. 1 (type locality: China: Sichuan: Wenchuan County<br />
(31°N 103°E): Sanjiang village, in a tributary <strong>of</strong> upper<br />
Minjian River; holotype: MSINR 910093; also spelt<br />
polpyfasciata p. 11, an obvious inadvertent error, thus<br />
incorrect original spelling [Code art. 32.5.1]; adjective,<br />
-us, -a, -um)<br />
10.42.84 Triplophysa pseudoscleroptera (Zhu & Wu,<br />
1981)<br />
Nemachilus pseudoscleropterus Zhu & Wu, 1981: 221, figs.<br />
1–3 (type locality: China: Qinghai: a tributary <strong>of</strong> Golmud<br />
He River, in Qaidam Pendi [Qaidam depression;<br />
Golmud town: 36°24'N 94°55'E]; holotype: NPIB 75066;<br />
compound adjective, -us, -a, -um)<br />
10.42.85 Triplophysa qiubeiensis Li & Yang, in Li, Yang,<br />
Chen, Tao, Qi & Han, 2008<br />
Triplophysa qiubeiensis Li & Yang, in Li, Yang, Chen, Tao, Qi<br />
& Han, 2008: 674, fig. 1 (type locality: China: Yunnan:<br />
Qiubei County (24°05'N 104°01'E): cave near Nijiao village;<br />
holotype: KIZ 20060409001; adjective, -is, -is, -e)<br />
10.42.86 ? Triplophysa retropinnis (Herzenstein, 1888)<br />
Nemachilus dorsonotatus retropinnis Herzenstein, 1888: 33,<br />
pl. 3 fig. 5a–b (type locality: China: Qinghai: Chami<br />
[Hami, Xinjiang; 42°48'00"N 93°27'00"E]; syntypes:<br />
ZISP 4206 [2] [ZISP 7208, according to Prok<strong>of</strong>iev,<br />
2007b: 4]; compound noun, indeclinable)<br />
Nemachilus dorsonotatus plagiognathus Herzenstein, 1888:<br />
33, pl. 5 fig. 5, pl. 7 fig. 2 (type locality: China: Qinghai:<br />
Lake Kuku-nor [Qinghai Hu] / Eastern Zaidam [Tsaidam;<br />
Qinghai: Qaidam basin] / spring at Galmyk / Ganssu<br />
/ Dabsun-gobi [apparently Caka Yanhu 36°42'00"N<br />
99°06'00"E, Dalai Dabassu on map in Przewal'skii,<br />
1876] / Chami [Hami, Xinjiang; 42°48'00"N 93°27'<br />
00"E] / Lake Alak-nor [Alag Hu] / Chuan-che River near<br />
Gomi [Huang He near Balekhun Gomi, approx. 30°10'N<br />
100°45'E; now flooded in Longyangxia Reservoir]; syntypes:<br />
ZISP 7252 [1], 7306 [2], 7312 [5], 7315 [2], 7319<br />
[more than 6], 7371 [1], 7853 [2], 7854 [1], 7855 [1];<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
130<br />
simultaneous subjective synonym <strong>of</strong> Nemachilus dorsonotatus<br />
retropinnis Herzenstein, 1888: 33; first reviser<br />
[apparently Prok<strong>of</strong>iev, 2007b: 9] gave precedence to<br />
N. d. retropinnis; compound adjective, -us, -a, -um)<br />
? Nemacheilus Bertini Fang, 1941: 253 (type locality: China:<br />
"high valleys <strong>of</strong> Mongolia" [apparently China: Hebei:<br />
Chongli County: Siwantse [Xiwanzi], 40°58'25"N<br />
115°16'22"E]; holotype: MNHN 3800, Bertin & Estève,<br />
1948: 97, fig. 1; noun in genitive, indeclinable)<br />
Nemachilus carassus Nikolski, in Prok<strong>of</strong>iev, 2007b: 4 (unpublished<br />
name on a label, not available; based on ZISP<br />
12576; locality: China: Qinghai: Lake Alyk Nor [Alag<br />
Hu, 35°34'N 97°06'E], southern slope <strong>of</strong> Burkhan Buddha<br />
Range [erroneously identified as a lake in Mongolia<br />
by Kottelat, 2006: 55; but see map in Kozlov, 1902a])<br />
Taxonomic notes. Treated as subspecies <strong>of</strong> T. stolickai by<br />
Prok<strong>of</strong>iev (2007b: 4), which geography suggests should be<br />
re-examined. The types <strong>of</strong> N. bertini were collected by<br />
A. David, apparently together with the holotype <strong>of</strong> Barbatula<br />
nuda, west <strong>of</strong> Beijing (see discussion under that species)<br />
and I doubt it is conspecific with T. retropinnis.<br />
10.42.87 Triplophysa robusta (Kessler, 1876)<br />
Nemachilus robustus Kessler, 1876: 32 (type locality: China:<br />
Gansu; holotype: ZISP 2486, Eschmeyer & Fricke,<br />
2010; adjective, -us, -a, -um)<br />
10.42.88 Triplophysa rosa Chen & Yang, 2005<br />
Triplophysa rosa Chen & Yang, 2005: 601, fig. 2 (type locality:<br />
China: Chongqing City: Wulong County: Dongba<br />
Cave [entrance 29°13'40.44"N 107°55'16.46"E], Wu<br />
Jiang drainage, Tianxing Town, Jiangkou Town; holotype:<br />
KIZ 200211001; noun in apposition, indeclinable)<br />
Linemacheilus wulong Anonym, 2005: 67 (nomen nudum)<br />
Heminoemacheilus wulongensis Wu, He, Zhen & Li, 2007:<br />
90 (nomen nudum)<br />
10.42.89 Triplophysa scapanognatha Prok<strong>of</strong>iev, 2007<br />
Triplophysa scapanognatha Prok<strong>of</strong>iev, 2007b: 9, fig. 4a (type<br />
locality: China: Qinghai: Quaidam [Qaidam]: Datzan<br />
Sume, near Donkyr [Donkür, Huangyuan, 36°41'N<br />
101°16'E; Huang He drainage]; holotype: ZISP 12495;<br />
compound adjective, -us, -a, -um)<br />
10.42.90 Triplophysa scleroptera (Herzenstein, 1888)<br />
Nemachilus scleropterus Herzenstein, 1888: 54, pl. 6 fig. 1<br />
(type locality: China: Qinghai: Chuan-Che River near<br />
Gomi [Huang He near Balekhun Gomi, approx. 30°10'N<br />
100°45'E; now flooded in Longyangxia Reservoir]; lectotype:<br />
ZISP 7231, designated by Prok<strong>of</strong>iev, 2003b: 61,<br />
fig. 20; compound adjective, -us, -a, -um)<br />
10.42.91 Triplophysa sellaefer (Nichols, 1925)<br />
Barbatula yarkandensis sellaefer Nichols, 1925b: 4 (type<br />
locality: China: Shansi: Chin Ssu, Taiyuanfu [Qingxu,<br />
37°36'N 112°21'E]; holotype: AMNH 8411; noun in apposition,<br />
indeclinable, Code art. 31.2.2)<br />
10.42.92 Triplophysa sewerzowi (Nikolski, 1938)<br />
Nemachilus sewerzowi Nikolski, 1938: 319, figs. 1–3 (type
locality: Kazakhstan: Alma-Atinskaya oblast: Kaskelen<br />
River, 2 km before it enters Ili River; holotype: ZMMU<br />
P-3638; noun in genitive, indeclinable)<br />
10.42.93 Triplophysa shaanxiensis Chen, in Chen, Xu,<br />
Fang, Song & Wang, 1987<br />
Triplophysa shaanxiensis Chen, in Chen, Xu, Fang, Song &<br />
Wang, 1987: 21, fig. 8 (type locality: China: Shaanxi:<br />
Tongchuan Xian; holotype: SIZX 801811; adjective, -is,<br />
-is, -e)<br />
10.42.94 Triplophysa shehensis Menon, 1987<br />
Triplophysa shehensis Menon, 1987: 212, pl. 16 fig. 9 (type<br />
locality: India: Kashmir: Ladakh: Sheh, irrigation canal,<br />
nearly 12 km east <strong>of</strong> Leh; holotype: ZSI/NRS uncat.;<br />
adjective, -is, -is, -e)<br />
Noemacheilus shehensis Tilak, 1990: 138, figs. 2–3 (type<br />
locality: India: Kashmir: Ladakh: irrigation canal at Sheh,<br />
nearly 12 km east <strong>of</strong> Leh; holotype: ZSI/NRS 1208; junior<br />
secondary homonym <strong>of</strong> Triplophysa shehensis Menon,<br />
1987: 212; paper dated 1990 but possibly not published<br />
until 1997; adjective, -is, -is, -e)<br />
Nomenclatural notes. Menon (1987) described this species<br />
and indicated the author as Tilak, then in press. Tilak's paper<br />
only appeared in 1990 (date on the cover, actual publication<br />
possibly much later, the paper is first listed in Zoological<br />
Record for 1997). The description in Menon is based on a<br />
single <strong>of</strong> Tilak's specimens and is not extracted from Tilak's<br />
text. Thus Menon is author <strong>of</strong> the name and the single specimen<br />
he examined is the holotype. The locality data listed<br />
by Menon is only "irrigation canal, Sheh" but as all <strong>of</strong> Tilak's<br />
(1997) specimens are from a single locality, it can be<br />
completed as above. Tilak's N. shehensis, being based on a<br />
separate description, without reference to Menon's description,<br />
is a distinct nominal species and a junior secondary<br />
homonym <strong>of</strong> T. shehensis when transferred to Triplophysa.<br />
It is not known whether the holotype <strong>of</strong> the two nominal<br />
species is the same specimen or not, which would make them<br />
objective synonyms.<br />
10.42.95 Triplophysa shilinensis Chen & Yang, in Chen,<br />
Yang & Xu, 1992<br />
Triplophysa shilinensis Chen & Yang, in Chen, Yang & Xu,<br />
1992: 17, fig. 1 (type locality: China: Yunnan: Lunan<br />
County: cave at Weiboyi village near Shi Lin Stone Forest<br />
[24°47'N 103°22'E; Romero et al., 2009: 263]; holotype:<br />
KIZ 913001; adjective, -is, -is, -e)<br />
10.42.96 Triplophysa shiyangensis (Zhao & Wang, 1983)<br />
Nemachilus shiyangensis Zhao & Wang, 1983: 97, fig. 1<br />
(type locality: China: Gansu: Leitai He stream near Wuwei<br />
[37°55'N 102°38'E], Shiyang He River system, "Huang<br />
He drainage" [in fact, Shiyang is an endorheic river<br />
ending in Gobi desert about 39°00'N 103°35'E]; holotype:<br />
DBLU 770582; adjective, -is, -is, -e)<br />
10.42.97 Triplophysa siluroides (Herzenstein, 1888)<br />
Nemachilus siluroides Herzenstein, 1888: 62, pl. 7 fig. 1,<br />
pl. 8 fig. 10 (type locality: China: Qinghai: Chuan-che<br />
River near Gomi [Huang He River near Balekhun Gomi,<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
131<br />
approx. 30°10'N 100°45'E; now flooded in Longyangxia<br />
Reservoir]; syntypes: ZISP 7243 [1], 7244 [1], 7851 [1])<br />
10.42.98 Triplophysa stenura (Herzenstein, 1888)<br />
Nemachilus stenurus Herzenstein, 1888: 64, pl. 1 fig. 1 (type<br />
locality: China: Tibet: Dy-Tschu [Tongtian He, in upper<br />
Yangtze basin; He et al., 2008: 48]; lectotype ZISP 7355,<br />
designated by Prok<strong>of</strong>iev, 2009a: 699; compound adjective,<br />
-us, -a, -um)<br />
? Nemachilus lhasae Regan, 1905b: 301 (type locality: Tibet:<br />
Lhasa; holotype: BMNH 1905.2.8.16; noun in genitive,<br />
indeclinable)<br />
Taxonomic notes. Triplophysa stenura is presently reported<br />
from the upper Yangtze, Mekong, Salween and Brahmaputra<br />
drainages, which suggests that several species are confused<br />
under this name.<br />
10.42.99 Triplophysa stewarti (Hora, 1922)<br />
Diplophysa stewarti Hora, 1922: 70, fig. 2c–d (type locality:<br />
China: Xizang: Rham-tso [Hram Tso, Bam Tso;<br />
28°08'52"N 89°20'02"E]; holotype: ZSI F 2894/1; noun<br />
in genitive, indeclinable)<br />
Nemachilus panguri Hora, 1936: 318, figs. 8–9, pl. 12 figs.<br />
3–4 (type locality: India: Jammu & Kashmir: Ladakh:<br />
Pangur Tso [Spanggur Tso, 33°32'N 78°55'E] / China:<br />
Xizang: Tso Nyak [apparently at 33°29'28"N 80°18'11"E]<br />
[both in Pangong Tso endorheic basin]; syntypes ["several"]:<br />
ZSI F 12204/1 [2], Menon & Yazdani, 1968: 123;<br />
noun in genitive, indeclinable)<br />
Nemachilus longianalis Ren & Wu, 1982: 82, fig. 2 (type<br />
locality: China: Northern Xizang: Nam Cuo Lake [? Nam<br />
Co, Nam Tso, 30°42'N 90°33'E]; holotype: HFRI 80-<br />
1669; compound noun, indeclinable)<br />
Nemachilus longianalis tangtianhensis Wu & Wu, 1988a:<br />
21, fig. 4 (type locality: China: Qinghai: mouth <strong>of</strong> Dangqu,<br />
upper reaches <strong>of</strong> Tangtianhe River; holotype: NPIB<br />
8606088; adjective, -is, -is, -e)<br />
10.42.100 Triplophysa stolickai (Steindachner, 1866)<br />
Cobitis Stoličkai Steindachner, 1866: 793, pl. 14 fig. 2 (type<br />
locality: India: Ladakh: Rupshu: rivulets in vicinity <strong>of</strong><br />
Lake Tsumureri [Tso Morari] [contra Eschmeyer &<br />
Fricke, 2010, "Umgebung" is not the name <strong>of</strong> a river but<br />
the German word meaning vicinity]; syntypes [12]: NMW<br />
48436 [5], 48439 [1], 50477 [4], Eschmeyer & Fricke,<br />
2010; the č in original spelling must be emended in c,<br />
Code art. 32.5.2.1; noun in genitive, indeclinable)<br />
Taxonomic notes. Prok<strong>of</strong>iev (2007b: 4) recognised T. stolickai<br />
with a number <strong>of</strong> subspecies. I do not recognise these<br />
subspecies because the Evolutionary Species Concept used<br />
here does not accomodate subspecies, and because these 'subspecies'<br />
are diagnosable lineages occupying very distinct and<br />
isolated areas and fulfill the criteria to be recognised as species.<br />
See T. cakaensis, T. dorsonotata, T. hexiensis, T. obtusirostra,<br />
T. retropinnis and T. uranoscopus.<br />
10.42.101 Triplophysa strauchii (Kessler, 1874)<br />
Diplophysa Strauchii Kessler, 1874: 58, pl. 8 fig. 40 (type<br />
locality: Kazakhstan: Ili River, tributary <strong>of</strong> Lake Balkash /<br />
Ich-Balya River; syntypes: ZISP 2328 [2], 2329 [1],
10.42.98 Triplophysa stenura, CMK 13062, 60.8 mm SL; China:<br />
Sichuan: Yangtze drainage: Yalong.<br />
Herzenstein, 1888: 51; noun in genitive, indeclinable)<br />
? Nemachilus strauchi transiens Herzenstein, 1888: 50 (type<br />
locality: China: Xinjiang: Tschertschen-darja [Charchan-Darya,<br />
Qarqan He; Qarqan town or Qiemo County:<br />
38°08'09"N 85°31'48"E] [Qarqan He, sometimes enters<br />
Lop Nor from SW]; syntypes: ZISP 7349 [5]; participle,<br />
indeclinable)<br />
? Nemachilus strauchi zaisanicus Menshikov, 1937: 441<br />
(type locality: Kazakhstan: Tarbagatai District: Karasu<br />
River at Akdzhar, basin <strong>of</strong> Zaisan Lake, 40 km south <strong>of</strong><br />
Lake Zaisan, between "47 and 48°N 53–54°E" [Aksuat,<br />
47°45'30"N 82°48'20"E; river maybe is Kargyba]; syntypes<br />
[27]: ZISP 26863 [18]; adjective, -us, -a, -um)<br />
? Nemachilus ruzskyi Nekrashevich, 1948: 121 (type locality:<br />
Kazakhstan: Lake Alakol [46°10'N 81°35'E; endorheic<br />
basin East <strong>of</strong> Lake Balkash]; types: LU; noun in genitive,<br />
indeclinable)<br />
? N[emacheilus] (Deuterophysa) strauchi jimbeyi Mitr<strong>of</strong>anov,<br />
2002: 125 (not available: no fixation <strong>of</strong> a holotype<br />
or syntypes, Code art. 16.4.1, no intention to establish<br />
a new taxon, Code art. 16.1; locality: Uzbekistan and<br />
Kyrgyzstan: Naryn River [40°54'N 71°45'E for confluence<br />
with Kara Darya; Karyn town: 41°26'N 76°00'E];<br />
noun in genitive, indeclinable)<br />
Taxonomic notes. Geography suggests that multiple species<br />
are confused under T. strauchii. Mitr<strong>of</strong>anov (2002: 125)<br />
considered N. s. zaisanicus and N. ruzskyi as synonyms <strong>of</strong><br />
T. strauchii.<br />
10.42.102 Triplophysa tanggulaensis (Zhu, 1982)<br />
Nemachilus tanggulaensis Zhu, 1982b: 223, figs. 1–4 (type<br />
locality: China: Qinghai: hot spring <strong>of</strong> Wenquan [33°<br />
14'02"N 91°51'06"E], between Yanshiping [33°35'51"<br />
92°04'07"E] and Tanggula Shankou [pass, 32°53'07"N<br />
91°55'04"]; Tanggula town 34°13'00"N 92°26'30"E];<br />
holotype: NPIB 75061; adjective, -is, -is, -e)<br />
10.42.103 Triplophysa tianeensis Chen, Cui & Yang, 2004<br />
Triplophysa tianeensis Chen, Cui & Yang, 2004a: 228, fig. 1<br />
(type locality: China: Guangxi: Tian'e County: Bala town:<br />
Hongshuihe underground river [24°58'N 107°02'E;<br />
Romero et al., 2009: 264]; holotype: KIZ 200301003;<br />
adjective, -is, -is, -e)<br />
10.42.104 Triplophysa tibetana (Regan, 1905)<br />
Nemachilus tibetanus Regan, 1905a: 187 (type locality:<br />
China: Tibet: Lhasa; holotype: BMNH 1904.12.28.86;<br />
adjective, -us, -a, -um)<br />
10.42.105 Triplophysa turpanensis Wu & Wu, 1992<br />
Triplophysa turpanensis Wu & Wu, 1992: 175, fig. 35 (type<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
132<br />
10.43.1 Troglocobitis starostini, ZISP 46159, 49.5 mm SL, holotype;<br />
Turkmenistan: Lebap Province: cave near Garlyk. (Photograph<br />
by Zakhar Zhidkov).<br />
locality: China: Xinjiang: Turpan City [42°59'N 89°11'E];<br />
holotype: NPIB 8706076; adjective, -is, -is, -e)<br />
10.42.106 Triplophysa ulacholica (Anikin, 1905)<br />
Nemachilus ulacholicus Anikin, 1905: 3, 18 [<strong>of</strong> reprint] (type<br />
locality: Kyrgyzstan: Lake Issyk-kul at mouth <strong>of</strong> Ulakhol<br />
River; syntypes [4]: LU, Vasil'eva, pers. comm., Eschmeyer<br />
& Fricke, 2010; adjective, -us, -a, -um)<br />
Nemacheilus strauchi dorsaloides Turdakov, 1947b: 155<br />
(type locality: Kyrgyzstan: Tyupsky Bay <strong>of</strong> Lake Issykkul;<br />
syntypes [28]: LU)<br />
Nemachilus strauchi reuniens Turdakov, 1952: 57 (type<br />
locality: Kyrgyzstan: Irisu River, tributary <strong>of</strong> Karkara<br />
River [tributary <strong>of</strong> Charyn River, Lake Balkash basin];<br />
syntypes [10]: LU; participle, indeclinable)<br />
10.42.107 Triplophysa uranoscopus (Kessler, 1872)<br />
Cobitis uranoscopus Kessler, 1872: 66, pl. 11 figs. 32–33<br />
(type locality: Tajikistan: Khodjaduk [Khodzhauk, Upper<br />
Amu Darya; Zeravshan, Berg, 1949: 862; apparently<br />
includes syntypes from Lake Iskanderkul, 39°04'30"N<br />
68°22'00"E]; syntypes [16]: ZISP 10931, ZMMU P2248,<br />
P2262, Svetovidova, 1978: 259, 262, Vasilieva et al., in<br />
Pavlinov & Borissenko, 2001: 28; compound noun, indeclinable)<br />
Nemachilus fedtschenkoae Nikolski, 1903: 93 (type locality:<br />
Transcaspia: Murgab River [Tajikistan: Murgab River,<br />
upper reaches <strong>of</strong> Amu-Darya drainage]; holotype: ZISP<br />
12710; noun in genitive, indeclinable)<br />
10.42.108 Triplophysa venusta Zhu & Cao, 1988<br />
Triplophysa venusta Zhu & Cao, 1988: 96, figs. 3–6 (type<br />
locality: China: Yunnan: Lijiang County: Heilongtan pond<br />
near Lijiang [26°53'N 100°14'E]; holotype: IHB<br />
60IV0303; adjective, -us, -a, -um)<br />
10.42.109 Triplophysa waisihani Cao & Zhang, 2008<br />
Triplophysa waisihani Cao & Zhang, 2008: 35, fig. 1 (type<br />
locality: China: Xinjiang: Yining County [Gulja; 43°55'N<br />
81°19E]: Kax River, a tributary <strong>of</strong> Ili River drainage at<br />
Dunmaza Town, "43°82'E 81°53'E, 1143 masl" [Dongmazar,<br />
Ma-char-erh, 43°49'20"N 81°52'10"E]; holotype:<br />
IHB 890064; noun in genitive, indeclinable)<br />
10.42.110 Triplophysa wuweiensis (Li & Chang, 1974)<br />
Nemachilus wuweiensis Li & Chang, 1974: 415, fig. 2 (type<br />
locality: China: Gansu: Wuwei County [37°55'N<br />
102°38'E]; holotype: ASIZB G65-243; adjective, -is,<br />
-is, -e)
10.44.1 Tuberoschistura baenzigeri, CMK 5144, 41.9 and 41.6<br />
mm SL; Thailand: Sai Buri.<br />
10.42.111 Triplophysa xiangshuingensis Li, 2004<br />
Triplophysa xiangshuingensis Li, 2004: 93, fig. 1 (type locality:<br />
China: Yunnan: Shilin County: underground river in<br />
Xiangshuig [24°52'N 103°21'E; Romero et al., 2009:<br />
264]; holotype: HRAS 200001128001; also spelt Xiang<br />
shuiqingensis on p. 96, an obvious inadvertent error, thus<br />
incorrect original spelling [Code art. 32.5.1]; adjective,<br />
-is, -is, -e)<br />
10.42.112 Triplophysa xiangxiensis (Yang, Yuan & Liao,<br />
1986)<br />
Noemacheilus xiangxiensis Yang, Yuan & Liao, 1986: 219,<br />
fig. 1 (type locality: China: Hunan: Xianxi Prefecture:<br />
Longshan County: Huoyan Town: tributary <strong>of</strong> Yuan River<br />
[Feihu Cave, 27°27'N 109°26'E; Romero et al., 2009:<br />
265]; holotype: HACW 8401; adjective, -is, -is, -e)<br />
10.42.113 Triplophysa xichangensis Zhu & Cao, in Zhu,<br />
1989<br />
Triplophysa xichangensis Zhu & Cao, in Zhu, 1989: 79, fig.<br />
53 (type locality: China: Sichuan: Xichangxian County<br />
[27°53'57"N 102°16'12"E]: Anninghe River; holotype:<br />
NGI or IHB 65V0714; adjective, -is, -is, -e)<br />
10.42.114 Triplophysa xingshanensis (Yang & Xie, 1983)<br />
Nemachilus xingshanensis Yang & Xie, 1983: 314, fig. 1<br />
(type locality: China: Hubei: Xingshan County [31°21'N<br />
110°45'E]: Xiangqi River, a tributary <strong>of</strong> upper Changjiang<br />
River [Yangtze]; holotype: HACW 81.8.09; adjective,<br />
-is, -is, -e)<br />
10.42.115 Triplophysa xiqiensis Ding & Lai, 1996<br />
Triplophysa xiqiensis Ding & Lai, 1996: 374, fig. 1 (type<br />
locality: China: Sichuan: Zhaojuexian County: Sikai village,<br />
27°55'N 102°45'E; holotype: MSINR 930013; adjective,<br />
-is, -is, -e)<br />
10.42.116 Triplophysa yaopeizhii Xu, Zhang & Cai, in<br />
Zhang, Cai & Xu, 1995<br />
Triplophysa yaopeizhii Xu, Zhang & Cai, in Zhang, Cai &<br />
Xu, 1995: 57, fig. 11-1 (type locality: China: Xizang:<br />
Jiangda, Mangkang and Gongjue Counties (30.9–31.5°N<br />
80.0–98.2°E), in Jinshajiang drainage [upper Yangtze]<br />
[see also map, fig. 11–2] [Jomda County: 31°29'58"N<br />
98°13'06"E; Markham, Markam: 29°41'40"N 98°35'<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
133<br />
10.45.2 Turcinoemacheilus kosswigi, CMK 22390, 57.8 mm SL;<br />
Turkey: Tigris drainage.<br />
44"E; Gongjo County: 30°5135"N 98°16'15"E]; syntypes<br />
[34]: probably includes material listed by Xu & Zhang<br />
(1996): ASIZB 92801262, 0696, 0778, 0877, 1258, 1316,<br />
0920 [7], IZSX 928-0694, 0885, 0856, 0734, 0715, 0876,<br />
0733, 0744, 0757, 0853, 0722, 0831, 1373, 1434, 1430,<br />
1444, 1427, 1465, 1315, 0949, 1317 [22]; noun in genitive,<br />
indeclinable)<br />
Nomenclatural notes. The name Triplophysa yaopeizhii first<br />
appeared in Zhang et al. (1995: 57) who neither explicitly<br />
listed it as a new name nor gave any bibliographic reference<br />
to another description. Another description appeared in Xu<br />
& Zhang (1996: 377), but it is not valid because it appeared<br />
later. The material listed in the 1995 description are syntypes<br />
and the material listed in the 1996 paper as holotype<br />
and paratypes (apparently from a different locality) have no<br />
type status.<br />
10.42.117 Triplophysa yunnanensis Yang, 1990<br />
Triplophysa yunnanensis Yang, in Chu & Chen, 1990: 56,<br />
fig. 51 (type locality: China: Yunnan: Yiliang County:<br />
Jiuxiang, 25°06'N 103°23'E; holotype: KIZ 874200; adjective,<br />
-is, -is, -e)<br />
10.42.118 Triplophysa zhaoi Prok<strong>of</strong>iev, 2006<br />
Nemachilus turfanensis Herzenstein, 1899: 414 (nomen nudum;<br />
locality: China: Xinjiang: Turpan depression: Lemdjin,<br />
Lükqün)<br />
Triplophysa zhaoi Prok<strong>of</strong>iev, 2006b: 244, figs. 40–42 (type<br />
locality: China: Xinjiang: swampy creeks <strong>of</strong> Lukchun<br />
oasis [Lükqün, 42°43'50"N 89°42'02"E], Turfan depression<br />
[Turpan]; holotype: ZISP 10548; noun in genitive,<br />
indeclinable)<br />
Nomenclatural notes. Nemacheilus turfanensis is mentioned<br />
in a list <strong>of</strong> fishes collected by the expedition <strong>of</strong><br />
Grumm-Grzhimaylo (1896, 1899), prepaired by Herzenstein<br />
before his death. The description was never completed or<br />
published.<br />
10.42.119 Triplophysa zhenfengensis Wang & Li, 2001<br />
Triplophysa zhenfengensis Wang & Li, 2001: 99, fig. 2 (type<br />
locality: China: Guizhou: Zhenfeng County: Longchang<br />
town, 25°32'N 105°12'E [25°27'47"N 105°29'50"E];<br />
holotype: ZMCZ 9050268; adjective, -is, -is, -e)<br />
10.43 Troglocobitis Parin, 1983<br />
Troglocobitis Parin, 1983: 83 (subgenus <strong>of</strong> Nemacheilus,<br />
Bleeker, 1863a: 37; type species: Nemacheilus starostini<br />
Parin, 1983: 83, by original designation). Gender feminine.<br />
Taxonomic notes. Related or possibly a synonym <strong>of</strong> Para-
cobitis, from which it differs by the loss <strong>of</strong> the bony capsule<br />
<strong>of</strong> the gas bladder (see Prok<strong>of</strong>iev, 2009b: 889). The loss <strong>of</strong><br />
structures alone is not a reliable criterion to recognised cave<br />
fishes as distinct genera. Identical losses may occur in parallel<br />
in many cave species.<br />
10.43.1 Troglocobitis starostini (Parin, 1983)<br />
Noemacheilus starostini Parin, 1983: 83, fig. (type locality:<br />
Turkmenistan: Lebap Province: Chardzhouskaya oblast:<br />
unnamed karst depresion [maybe 37°37'06"N<br />
66°21'09"E] at foothills <strong>of</strong> Kugitangtau Range [Köýtendag]<br />
east <strong>of</strong> Karlyuk village [Garlyk, 37°35'25"N<br />
66°20'35"E] [Vasilieva et al., in Pavlinov & Borissenko,<br />
2001: 29]; holotype: ZISP 46159; noun in genitive, indeclinable)<br />
10.44 Tuberoschistura Kottelat, 1990<br />
Tuberoschistura Kottelat, 1990a: 232 (type species: Nemacheilus<br />
baenzigeri Kottelat, 1983: 151, by original designation).<br />
Gender feminine.<br />
10.44.1 Tuberoschistura baenzigeri (Kottelat, 1983)<br />
Noemacheilus baenzigeri Kottelat, 1983: 151, figs. 1–2 (type<br />
locality: Thailand: Chiang Mai Province: Mae Nam Taeng<br />
at Mae Taeng, Chao Phraya drainage, 19°07'N 98°56'E;<br />
holotype: MHNG 2081.32; noun in genitive, indeclinable)<br />
10.44.2 Tuberoschistura cambodgiensis Kottelat, 1990<br />
Tuberoschistura cambodgiensis Kottelat, 1990a: 237, fig.<br />
178 (type locality: Cambodia: Stung Chihreng, a northern<br />
tributary <strong>of</strong> Grand Lac, at "Pont Khmer" on road from<br />
Siem Reap to Kompong Thom, Mekong drainage; holotype:<br />
MNHN 1988.99; adjective, -is, -is, -e)<br />
10.45 Turcinoemacheilus Banarescu & Nalbant, 1964<br />
Turcinoemacheilus Bănărescu & Nalbant, 1964: 178 (type<br />
species: Turcinoemacheilus kosswigi Bănărescu & Nalbant,<br />
1964: 178, by original designation). Gender masculine.<br />
10.45.1 Turcinoemacheilus himalaya Conway, Edds,<br />
Shrestha & Mayden, 2011<br />
Turcinoemacheilus himalaya Conway, Edds, Shrestha &<br />
Mayden, 2011: 1748, figs. 1, 2a (type locality: Nepal:<br />
Bagmati Zone: Sindhupalchok District: Indrawati River<br />
at Melamchi Township, 27°49'42.5"N 85°34'37.1"E;<br />
holotype: KU 40558; noun in apposition, indeclinable)<br />
10.45.2 Turcinoemacheilus kosswigi Banarescu & Nalbant,<br />
1964<br />
Turcinoemacheilus kosswigi Bănărescu & Nalbant, 1964:<br />
178, pl. 8 fig. 14 (type locality: Turkey: Hakkari<br />
[37°34'40"N 43°44'10"E]: Kapodiz Kadun, Tigris drainage;<br />
holotype: ZMH H 1884, Wilkens, 1977: 159; noun<br />
in genitive, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
134<br />
10.46 Yunnanilus Nichols, 1925<br />
Yunnanilus Nichols, 1925b: 1 (subgenus <strong>of</strong> Nemacheilus<br />
Bleeker, 1863a: 37; type species: Nemacheilus pleurotaenia<br />
Regan, 1904: 192, by subsequent designation by<br />
Norman, 1926: 31). Gender masculine.<br />
Taxonomic notes. The genus Yunnanilus is very heterogenous.<br />
Part <strong>of</strong> the species that had been placed in a Y. nigromaculatus<br />
group (e.g. by Chen et al., 2012) are now Eonemachilus.<br />
Other species <strong>of</strong> that group are still listed under<br />
Yunnanilus but may belong to Heminoemacheilus or some<br />
unnamed genus or genera. Yunnanilus brevis and some unnamed<br />
species from Myanmar are placed in Petruichthys.<br />
Nomen nudum<br />
Yunnanilus macroiscalus Li & Duan, 1999: 255<br />
10.46.1 Yunnanilus altus Kottelat & Chu, 1988<br />
Yunnanilus altus Kottelat & Chu, 1988a: 72, fig. 6 (type locality:<br />
China: Yunnan: Zhanyi County: Xiangshuiho at<br />
Haijiashao, 25°40'N 103°57'E; holotype: KIZ 774630;<br />
adjective, -us, -a, -um)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Heminoemacheilus.<br />
10.46.2 Yunnanilus analis Yang, in Chu & Chen, 1990<br />
Yunnanilus analis Yang, in Chu & Chen, 1990: 19, fig. 14<br />
(type locality: China: Yunnan: Xingyun Lake, 24°17–23'N<br />
102°45–48'E; holotype: KIZ 01402; treated as noun in<br />
apposition, indeclinable)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Heminoemacheilus.<br />
10.46.3 Yunnanilus bajiangensis Li, 2004<br />
Yunnanilus bajiangensis Li, 2004: 94, fig. 2 (type locality:<br />
China: Yunnan: Shilin County: Heilongtan Reservoir, Ba<br />
Jiang drainage [24°46'N 103°16'E; Romero et al., 2009:<br />
267]; holotype: HRAS 9504001; spelt bajiangensis (2x)<br />
and bajingensis (2x); bajiangensis assumed to be correct<br />
spelling as the name is based on the name <strong>of</strong> a river<br />
[jiang]; adjective, -is, -is, -e)<br />
10.46.4 Yunnanilus beipanjiangensis Li, Mao & Sun, in<br />
Li, Mao, Sun & Lu, 1994<br />
Yunnanilus beipanjiangensis Li, Mao & Sun, in Li, Mao,<br />
Sun & Lu, 1994: 370, fig. 1 (type locality: China: Yunnan:<br />
Zhangyi County [Zhanyi]: Xintun, 26.02°N<br />
104.01°E; holotype: FACQR 9107016; adjective, -is,<br />
-is, -e)<br />
10.46.5 Yunnanilus caohaiensis Ding, 1992<br />
Yunnanilus caohaiensis Ding, 1992: 489, fig. 1 (type locality:<br />
China: Guizhou: Weining County: Caohai Lake,<br />
26°49'–26°53'N 104°12'–104°18'E; holotype: MSINR<br />
900077; adjective, -is, -is, -e)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Heminoemacheilus.<br />
10.46.6 Yunnanilus chui Yang, 1991<br />
Yunnanilus chui Yang, 1991: 199, fig. 3 (type locality: China:<br />
Yunnan: Fuxian Lake at Haikou, 24°29'N 102°56'E;<br />
holotype: KIZ 893095; noun in genitive, indeclinable)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Eonemachilus.
10.46.7 Yunnanilus discoloris Zhou & He, 1989<br />
Yunnanilus discoloris Zhou & He, 1989: 381, fig. 1 (type<br />
locality: China: Yunnan: Chengdong County: White Dragon<br />
Spring [24°54'N 102°48'E; Romero et al., 2009: 268,<br />
but real locality is in cement factory at 24°52'36.8"N<br />
102°51'47.2"E, pers. obs.]; holotype: YU 875001; adjective<br />
in genitive [not nominative, Code art. 31.2], treated<br />
as noun in apposition, indeclinable)<br />
10.46.8 Yunnanilus elakatis Cao & Zhu, in Zheng, 1989<br />
Yunnanilus pleurotaenia elakatis Cao & Zhu, in Zheng, 1989:<br />
43, fig. 21 (type locality: China: Yunnan: Yiliang County;<br />
holotype: IHB 64V036; noun in apposition, indeclinable)<br />
10.46.9 Yunnanilus ganheensis An, Liu & Li, 2009<br />
Yunnanilus ganheensis An, Liu & Li, 2009: 635, fig. 11 (type<br />
locality: China: Yunnan: Xundan County: Ganhe, 25°30'N<br />
103°40'E [Yangtze drainage]; holotype: HRAS<br />
920217004; adjective, -is, -is, -e)<br />
10.46.10 Yunnanilus jinxiensis Zhu, Du & Chen, in Zhu,<br />
Du, Chen & Yang, 2009<br />
Yunnanilus jinxiensis Zhu, Du & Chen, in Zhu, Du, Chen &<br />
Yang, 2009: 196, fig. 1 (type locality: China: Guangxi:<br />
Jingxi County: Ludon village, Pearl River drainage; holotype:<br />
KIZ 0800018; adjective, -is, -is, -e)<br />
10.46.11 Yunnanilus longibarbatus Gan, Chen & Yang,<br />
2007<br />
Yunnanilus longibarbatus Gan, Chen & Yang, 2007: 322,<br />
fig. 1 (type locality: China: Guangxi: Du-An County:<br />
Hongshui River drainage in Gaoling township; holotype:<br />
KIZ 2003050255; adjective, -us, -a, -um)<br />
10.46.12 Yunnanilus longibulla Yang, in Chu & Chen,<br />
1990<br />
Yunnanilus longibulla Yang, in Chu & Chen, 1990: 21, fig. 16<br />
(type locality: China: Yunnan: Chenghai Lake, 26°27–<br />
38'N 100°38–41'E; holotype: KIZ 818500; compound<br />
noun, indeclinable)<br />
10.46.13 Yunnanilus macrogaster Kottelat & Chu, 1988<br />
Yunnanilus macrogaster Kottelat & Chu, 1988a: 81, fig. 17<br />
(type locality: China: Yunnan: Luoping County: Tatantze<br />
[Datangzi], 24°52'N 104°18'E; holotype: KIZ 806062;<br />
compound noun, indeclinable)<br />
10.46.14 Yunnanilus macrositanus Li, in Li, Wu, Xu, Gao,<br />
Chen, Wu & Wang, 1999<br />
Yunnanilus macrositanus Li, in Li, Wu, Xu, Gao, Chen, Wu<br />
& Wang, 1999: 4, fig. 1 (type locality: China: Yunnan:<br />
Lunan County: Heilongtan, 24°45'42"N 103°15'29"E;<br />
holotype: HRAS 6502041; spelt macrositanus pp. 3, 4, 6,<br />
macroistainus p. 4, macroistanus pp. 5, 6; as first reviser<br />
I select macrositanus as correct original spelling; etymology<br />
unknown, treated as noun in apposition, indeclinable)<br />
? Yunnanilus forkicaudalis Li, in Li, Wu, Xu, Gao, Chen,<br />
Wu & Wang, 1999: 4, fig. 2 (type locality: China: Yunnan:<br />
Lunan County: Heilongtan, 24°45'42"N 103°15'<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
135<br />
29"E; holotype: HRAS 6509047; spelt forkicaudalis<br />
pp. 3, 4, forkeicaudalis p. 5; as first reviser I select forkicaudalis<br />
as correct original spelling; compound noun,<br />
indeclinable; apparently a simultaneous subjective synonym<br />
<strong>of</strong> Yunnanilus macrositanus Li, in Li, Wu, Xu, Gao,<br />
Chen, Wu & Wang, 1999: 4; as first reviser I give precedence<br />
to Y. macrositanus)<br />
10.46.15 Yunnanilus nanpanjiangensis Li, Mao & Lu, in<br />
Li, Mao, Sun & Lu, 1994<br />
Yunnanilus nanpanjiangensis Li, Mao & Lu, in Li, Mao,<br />
Sun & Lu, 1994: 371, fig. 2 (type locality: China: Yunnan:<br />
Luoping County: Agang town, 25°04'N 104°07'E<br />
[She'Ao village; 25°05'N 104°08'E; Romero et al., 2009:<br />
271]; holotype: FACQR 9191119; also spelt nanpanjingensis<br />
p. 374, an obvious inadvertent error, thus incorrect<br />
original spelling [Code art. 32.5.1]; adjective, -is,<br />
-is, -e)<br />
10.46.16 Yunnanilus niger Kottelat & Chu, 1988<br />
Yunnanilus niger Kottelat & Chu, 1988a: 73, fig. 8 (type<br />
locality: China: Yunnan: Luoping County: Tatantze [Datangzi],<br />
24°52'N 104°18'E; holotype: KIZ 806075; adjective,<br />
-er, -ra, -rum)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Heminoemacheilus.<br />
10.46.17 Yunnanilus niulanensis Chen, Yang & Yang,<br />
2012<br />
Yunnanilus niulanensis Chen, Yang & Yang, 2012: 58, fig. 1<br />
(type locality: China; Yunnan: Songming County: Yanglinhe<br />
River, 25°14'34"N 103°03'53.3"E, 1909 masl,<br />
upper reaches <strong>of</strong> Niulanjiang River, in Yanglin town, Jinsha<br />
Jiang drainage [Yangtze]; holotype: KIZ 20060285;<br />
adjective, -is, -is, -e)<br />
10.46.18 Yunnanilus obtusirostris Yang, in Yang & Chen,<br />
1995<br />
Yunnanilus obtusirostris Yang, in Yang & Chen, 1995: 21,<br />
fig. 6 (type locality: China: Yunnan: Chengjiang County:<br />
West Dragon Spring, flowing into Fuxian Lake [24°30'N<br />
102°53'E; Romero et al., 2009: 272]; holotype: KIZ<br />
878052; compound noun, indeclinable)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Heminoemacheilus.<br />
10.46.19 Yunnanilus pachycephalus Kottelat & Chu, 1988<br />
Yunnanilus pachycephalus Kottelat & Chu, 1988a: 74, fig. 10<br />
(type locality: China: Yunnan: Xuanwei County:<br />
Weizhangho at Yangliu, 26°38'N 104°15'E; holotype: KIZ<br />
82100392; compound adjective, -us, -a, -um)<br />
10.46.20 Yunnanilus paludosus Kottelat & Chu, 1988<br />
Yunnanilus paludosus Kottelat & Chu, 1988a: 76, fig. 12<br />
(type locality: China: Yunnan: Luoping County: Tatantze<br />
[Datangzi], 24°52'N 104°18'E; holotype: KIZ 806059;<br />
adjective, -us, -a, -um)<br />
Yunnanilus macrolepis Li, Tao & Mao, in Li, Tao, Mao & Lu,<br />
2000: 349, fig. 1 (type locality: China: Yunnan: Luoping<br />
County: Xuetian Longan, 24°50'N 104°20'E [24°53'N<br />
104°18'E; Romero et al., 2009: 270]; holotype: HRAS<br />
or ASIZB 9307001; compound noun, indeclinable)
10.46.22 Yunnanilus pleurotaenia, CMK 13100, 45.5 mm SL;<br />
China: Yunnan: Lake Dianchi basin.<br />
10.46.21 Yunnanilus parvus Kottelat & Chu, 1988<br />
Yunnanilus parvus Kottelat & Chu, 1988a: 77, fig. 13 (type<br />
locality: China: Yunnan: Kaiyuan County: Nan Tong cave,<br />
23°39'N 103°17'E; holotype: KIZ 847174; adjective,<br />
-us, -a, -um)<br />
10.46.22 Yunnanilus pleurotaenia (Regan, 1904)<br />
Nemachilus pleurotaenia Regan, 1904: 192 (type locality:<br />
China: Yunnan: Sea <strong>of</strong> Tien at Yunnan Fu [Lake Dianchi,<br />
Kunming]; lectotype: BMNH 1904.1.26.36, designated<br />
by Kottelat & Chu, 1988a: 82; compound noun, indeclinable)<br />
? Yunnanilus tigeriveinus Li & Duan, 1999: 254, fig. 1 (type<br />
locality: China: Yunnan: suburbs <strong>of</strong> Kunming, in an<br />
"opening <strong>of</strong> underground water", 25°10'N 102°55'E; holotype:<br />
HRAS 9901004; also spelt tigerivinus p. 256, as<br />
tigeriveinus is used five times and tigerivinus once, tigerivinus<br />
is treated as an inadvertent error, thus incorrect<br />
original spelling [Code art. 32.5.1]; etymology unknown,<br />
treated as noun in apposition, indeclinable)<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
136<br />
10.46.23 Yunnanilus pulcherrimus Yang, Chen & Lan,<br />
2004<br />
Yunnanilus pulcherrimus Yang, Chen & Lan, 2004: 112,<br />
fig. 1 (type locality: China: Guangxi: Du'an County:<br />
Hongshuihe River, a tributary <strong>of</strong> Xijiang [23°56'N<br />
108°05'E; Romero et al., 2009: 274]; holotype: KIZ<br />
995001; also in Lan, 2004; adjective, -us, -a, -um)<br />
Yunnanilus zebrinus Lan & Zhang, in Zhou & Zhang, 2006:<br />
87 (nomen nudum; bibliographic reference make it clear<br />
that it is an inadvertent error for Yunnanilus pulcherrimus<br />
Yang, Chen & Lan, 2004: 112; fig. II-19 is fig. 1 <strong>of</strong><br />
Yang et al., 2004: 112)<br />
Taxonomic notes. Possibly a species <strong>of</strong> Micronemacheilus.<br />
10.46.24 Yunnanilus sichuanensis Ding, 1995<br />
Yunnanilus sichuanensis Ding, 1995: 253, fig. 1 (type locality:<br />
China: Sichuan: Mianning County: Anning River,<br />
Shuyalong Jiang River, 2020 masl, 28°41'N 102°12'E;<br />
holotype: MSINR 930026; adjective, -is, -is, -e)<br />
10.46.25 Yunnanilus spanisbripes An, Liu & Li, 2009<br />
Yunnanilus spanisbripes An, Liu & Li, 2009: 631, fig. 1 (type<br />
locality: China: Yunnan: Zhanyi County: Niulanjiang<br />
River, 25°59'N 103°36'E [Yangtze drainage]; holotype:<br />
HRAS 9206064; also spelt spanitripes, p. 637; as first<br />
reviser I select spanisbripes as correct original spelling;<br />
etymology unknown, noun in apposition, indeclinable)
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
APPENDIX: NEW GENUS-GROUP AND FAMILY-GROUP NAMES<br />
Ambastaia, new genus<br />
Type species. — Botia nigrolineata Kottelat & Chu, 1987.<br />
Diagnosis. — Ambastaia is distinguished from all other genera<br />
<strong>of</strong> the family Botiidae by its unique colour pattern in<br />
adults consisting in a whitish-yellowish background with a<br />
black midlateral stripe on the flank and another, middorsal<br />
stripe, with vertical bars connecting the two stripes and extending<br />
on the lower half <strong>of</strong> the body. In A. nigrolineata only<br />
the two stripes are present in juveniles and the bars appear at<br />
about 40–50 mm SL. In A. sidthimunki, the mid-dorsal stripe<br />
is <strong>of</strong>ten longitudinally divided by a whitish median area, or<br />
a row <strong>of</strong> whitish blotches, and the midlateral stripe may appear<br />
as a row <strong>of</strong> closely connected blotches; the juveniles<br />
have not yet been described.<br />
Etymology. — Named for Ambastai (or Ambastus in Latin),<br />
a river in Ptolemy’s (ca. 90–168) Γεωγραφκ γφγησις<br />
(Geographikê Hyphêgêsis, Geography). This river has been<br />
identified as the Mekong (van der Meulen, 1974, 1975).<br />
Gender feminine.<br />
Remarks. — The colour pattern <strong>of</strong> A. nigrolineata and A.<br />
sidthimunki and its ontogeny are unique within Botiidae. Both<br />
were originally described in the genus Botia and they were<br />
later transferred to Yasuhikotakia by Nalbant (2002). Still<br />
the colour pattern distinguishes them from that <strong>of</strong> all species<br />
<strong>of</strong> Yasuhikotakia, which includes a large blackish blotch at<br />
the base <strong>of</strong> the caudal fin and, in juveniles, a number <strong>of</strong> narrow<br />
bars on the flank (see figures in Kottelat, 2001b: 87). In<br />
most individuals, these bars become indistinct with age. Other<br />
elements <strong>of</strong> the colour pattern (e.g. middorsal stripes, black<br />
spots and subdistal margin on fins) may be present. This<br />
contrasts with Ambastaia in which the stripes appear first,<br />
followed by the vertical elements. Those species <strong>of</strong> Yasuhikotakia<br />
that have been studied osteologically have a long<br />
process <strong>of</strong> the frontal along the anterior margin <strong>of</strong> the orbit<br />
(called “spinous fringe” by Taki, 1972); it is missing in A.<br />
sidthimunki. Based on this and other characters, Taki (1972:<br />
78) commented that A. sidthimunki “appears to be the remotest<br />
from [...] the rest in the group” [his modesta group,<br />
equivalent to Yasuhikotakia]. This condition <strong>of</strong> the ‘spinous<br />
fringe’ has not yet been checked in all known species and<br />
therefore it is not used here as a diagnostic character.<br />
The molecular phylogeny presented by Šlechtová et al. (2005)<br />
included A. nigrolineata and A. sidthimunki and showed that<br />
they are each other’s closest relative but not closely related<br />
to the other species placed in Yasuhikotakia. On the contrary,<br />
they form the sister-group <strong>of</strong> Sinibotia and together<br />
they are the sister-group <strong>of</strong> Syncrossus. This lineage, in turn,<br />
constitutes the sister group <strong>of</strong> Yasuhikotakia. This clearly<br />
137<br />
shows that A. sidthimunki and A. nigrolineata form a distinct<br />
lineage. With the recognition <strong>of</strong> Ambastaia, Yasuhikotakia<br />
seems now monophyletic.<br />
Admittedly, it is quite weak to diagnose a genus by its colour<br />
pattern; combined with the molecular evidence it seems<br />
however justified to formally recognise this lineage by a name<br />
awaiting for (possible) more detailed anatomical studies.<br />
Assuming that this kind <strong>of</strong> research will still get support and<br />
be tolerated. I wish to point that to me, in botiids, the ontogeny<br />
<strong>of</strong> the colour pattern seems much more informative than<br />
the colour pattern itself. Unfortunately there is no published<br />
information on the colour pattern <strong>of</strong> the juveniles <strong>of</strong> the<br />
Chinese genera (Sinibotia, Leptobotia, Parabotia).<br />
Theriodes, new genus<br />
Type species. — Acanthophthalmus sandakanensis Inger &<br />
Chin, 1962: 120.<br />
Diagnosis. — Theriodes is distinguished from the other genera<br />
<strong>of</strong> the family Cobitidae in Southeast Asia in having the<br />
black marking at the base <strong>of</strong> the caudal-fin made <strong>of</strong> a single<br />
small black spot; it is slightly vertically elongated, at midheight<br />
<strong>of</strong> the caudal-fin base. The gill opening is small, oval,<br />
its length is about equal to the length <strong>of</strong> the base <strong>of</strong> the pectoral<br />
fin and it is located entirely above the base <strong>of</strong> the pectoral<br />
fin. The anterior nostril is at the tip <strong>of</strong> a short conical<br />
tube and the posterior nostril is not immediately adjacent to<br />
the tube, but separated by a distance about equal to its own<br />
diameter (shared with Kottelatlimia katik). The pectoral fin<br />
has 5 branched rays. In the male the first two pectoral rays<br />
are thickened and have a few tubercles along the anterior<br />
edge; the second ray is branched but there is no membrane<br />
between the branches and the posterior branch is much thinner<br />
than the anterior one; superficially the ray may appear<br />
unbranched (as originally described by Inger & Chin, 1962);<br />
the branches are more discernible in females and juveniles.<br />
The other pectoral rays are clearly branched, but the branching<br />
point is close to the tip <strong>of</strong> the rays.<br />
The following characters also help to distinguish the genus,<br />
although none is unique to it: pelvic fin with 1 simple and 4<br />
branched rays; dorsal-fin origin about halfway between pelvic-fin<br />
origin and anal-fin origin; caudal fin rounded; lower<br />
lip interrupted medially, each half with an inner, thickened<br />
mental lobe; membrane connecting mental lobe to corner <strong>of</strong><br />
mouth with two barbel-like projections, one <strong>of</strong> them continuing<br />
the mental lobe; barbels short, broad, somewhat triangular<br />
and flattened; bifid suborbital spine; eye covered by<br />
skin; 25–26 + 10–11 = 35–36 vertebrae (Kottelat & Lim,<br />
1992: 202).
Etymology. — Named for Theriodes Kolpos (or Theriodis<br />
Sinus in Latin, the Bay <strong>of</strong> the Beasts), a place name in Ptolemy’s<br />
(ca. 90–168) Γεωγραφκ γφγησις (Geographikê Hyphêgêsis,<br />
Geography). It has been identified by some as possibly<br />
Borneo (van der Meulen, 1974, 1975). Treated as masculine.<br />
Remarks. — Theriodes sandakanensis was originally described<br />
as a species <strong>of</strong> Acanthophthalmus (a junior synonym<br />
<strong>of</strong> Pangio) (Inger & Chin, 1962: 120). There was no explanation<br />
as to why the species was placed in Pangio but the<br />
description includes comparison only with species placed in<br />
Pangio by earlier authors (e.g. Weber & de Beaufort, 1916;<br />
Smith, 1945). Several <strong>of</strong> these species have a diagnostic<br />
‘barred’ colour pattern and Inger & Chin explicitly discussed<br />
this barred pattern. It is not appropriate to describe the colour<br />
pattern as barred. In the species <strong>of</strong> Pangio with a barred<br />
pattern, the bars are black, regular, with sharp edges and<br />
contrasted on a white to yellowish background. In Theriodes,<br />
the colour pattern is better described as dark brown<br />
marks on a paler, yellowish brown background, made <strong>of</strong> superficial<br />
and inner pigments; the marks are irregular and<br />
somewhat organised in 4–5 transverse bands. Besides, species<br />
<strong>of</strong> Pangio are more elongated, with 45–71 vertebrae (Kottelat<br />
& Lim, 1993), while T. sandakanensis has only 35–36<br />
(Kottelat & Lim, 1992). Roberts (1989: 103) placed T. sandakanensis<br />
in the genus Lepidocephalichthys. It indeed<br />
shares a number <strong>of</strong> characters with Lepidocephalichthys, e.g.,<br />
the rounded caudal fin, the general morphology <strong>of</strong> the lower<br />
lip, and the vertebrae counts (see Kottelat & Lim, 1992).<br />
Theriodes is, however, missing the most obvious character<br />
diagnosing Lepidocephalichthys, the sexually dimorphic<br />
modified pectoral-fin rays in males. Sexual dimorphism provides<br />
very efficient characters to identify lineages in Cobitidae<br />
(Šlechtová et al., 2008). In many genera, males have<br />
thickened, hardened or rigid anterior pectoral rays, <strong>of</strong>ten with<br />
projections or appendages. Lepidocephalichthys is unique<br />
in having no modification on the anterior rays but in having<br />
the last two rays (7 and 8) fused and swollen, <strong>of</strong>ten with a<br />
longitudinally elongated, <strong>of</strong>ten laminar, protuberance along<br />
the dorsal surface (Kottelat & Lim, 1992: 203, fig. 1; Nalbant,<br />
1963, 1994; Havird & Page, 2010). In Pangio, the pectoral-fin<br />
rays <strong>of</strong> the males are longer and more rigid than in<br />
the females, and they have a typical curled appearance (Kottelat<br />
& Lim, 1993). In Kottelatlimia, earlier included in Lepidocephalichthys,<br />
a number <strong>of</strong> segments <strong>of</strong> the dorsal hemitrich<br />
<strong>of</strong> the first branched ray have a posterior laminar projection<br />
forming a kind <strong>of</strong> blade or saw-like structure (Kottelat<br />
& Tan, 2008: 66, fig. 3).<br />
None <strong>of</strong> these structures is present in Theriodes. The pectoral<br />
fin is small, with only 5 branched rays (apparently fewer<br />
than in all other cobitid genera, 6–10). The first two rays are<br />
thickened in the male, and the second ray (homologous to<br />
the first branched ray in the females and juveniles) is described<br />
as “appeared unbranched” by Inger & Chin (1962:<br />
123). In fact it is branched but there is no membrane between<br />
the branches. A similar situation is also observed in<br />
Kottelatlimia and Acantopsis (Kottelat & Tan, 2008).<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
138<br />
Havird & Page (2010: 156) mentioned the sexual dimorphism<br />
in Theriodes and Lepidocephalichthys, but in a confusing<br />
way. The main problem is their misunderstanding <strong>of</strong><br />
the ‘lamina circularis’. They comment that the lamina circularis<br />
is “formed by the first and second pectoral rays in mature<br />
males <strong>of</strong> [T.] sandakanensis (as well as a thickening <strong>of</strong><br />
the first pelvic ray) vs. the seventh and eighth pectoral rays<br />
in mature males <strong>of</strong> Lepidocephalichthys”. Elsewhere (p. 137;<br />
also Havird et al., 2010: 13), “[t]his modification is referred<br />
to as the lamina circularis and is formed by a fusion and<br />
hardening <strong>of</strong> the innermost (seventh and eighth) pectoral rays.<br />
In mature males <strong>of</strong> other cobitid genera, the lamina circularis<br />
is formed by different pectoral rays (usually the second)<br />
or is absent”. To refer to all types <strong>of</strong> sexually dimorphic<br />
modification <strong>of</strong> the pectoral rays as the same structure,<br />
under the same name, obviously is misleading as the compared<br />
structures are not homologous. Next comes the use <strong>of</strong><br />
the wording ‘lamina circularis’ to refer to just any modification<br />
<strong>of</strong> the pectoral rays. This obviously ignores that there is<br />
a very large corpus <strong>of</strong> literature on cobitids and their sexual<br />
dimorphism by authors in Europe, China, Japan and Korea,<br />
spanning over about 140 years. Lamina circularis means circular<br />
lamella. The sexual dimorphism in Cobitis was first<br />
described (imperfectly as a swelling <strong>of</strong> the ray) by Canestrini<br />
(1871) and it later became known as Canestrini scale. As<br />
implied by both names, it refers to a round structure and it<br />
would be strange to use it as a general term meaning modified<br />
pectoral ray. The lamina circularis is a laminar, usually<br />
but not always circular, posterior projection <strong>of</strong> the first (proximal-most)<br />
segment <strong>of</strong> the dorsal hemitrich <strong>of</strong> the first<br />
branched pectoral ray (see, e.g., Nalbant, 1963: 356; Kottelat<br />
& Freyh<strong>of</strong>, 2007: 301, fig. 61). A second lamina circularis<br />
may also be present on the second branched ray in some<br />
species. It is not homologous with the thickened rays 1–2 <strong>of</strong><br />
Theriodes, with the curled rays <strong>of</strong> Pangio, with the fused<br />
rays 7–8 <strong>of</strong> Lepidocephalichthys, or with the blade or ‘saw’<br />
on ray 2 <strong>of</strong> Kottelatlimia. But it is apparently homologous<br />
with the projection on the first segment <strong>of</strong> ray 2 (the proximal-most<br />
‘tooth’ <strong>of</strong> the ‘saw’) in Kottelatlimia (Kottelat &<br />
Tan, 2008: 66, fig. 3).<br />
The black pattern at the base <strong>of</strong> the caudal fin is also useful<br />
to distinguish some lineages in all families <strong>of</strong> Cobitoidea.<br />
Theriodes has a single small black spot at mid-height <strong>of</strong> the<br />
caudal fin, a pattern not seen in any other cobitid (but known<br />
in other loach families). In Lepidocephalichthys and Kottelatlimia,<br />
as in the majority <strong>of</strong> cobitids, if present, the black<br />
spot (or one <strong>of</strong> the black spots) is in the upper third <strong>of</strong> the<br />
caudal-fin base, usually somewhat slanted. There is no spot<br />
in Pangio, but in some species a bar occupies most <strong>of</strong> the<br />
caudal-fin base.<br />
The gill opening seems smaller in Theriodes than in any other<br />
cobitid, but this still requires confirmation. In most cobitids,<br />
the posterior nostril is adjacent to the anterior one (or to<br />
the base <strong>of</strong> the tube at the top <strong>of</strong> which the anterior nostril is<br />
located), but in Theriodes there is some distance between<br />
the tube and the posterior nostril. The diagnostic value <strong>of</strong><br />
this character needs to be checked in more genera.
Material. — Theriodes sandakanensis: ZRC 37645, 16 (<strong>of</strong><br />
17); CMK 11557, 2; 10.5–35.0 mm SL; Borneo: Sabah:<br />
Kinabatangan drainage at SAFODA, near Batu Puteh, K. K.<br />
P. Lim et al., 7–13 Apr. 1994. - ZRC 45473, 2 (<strong>of</strong> 3), 21.8–<br />
22.1 mm SL; Borneo: Sabah: Lungmanis, Goh Y.-Y., 27 Sep.<br />
1998.<br />
Speonectes, new genus<br />
Type species. — Sundoreonectes tiomanensis Kottelat, 1990.<br />
Diagnosis. — Speonectes is distinguished from the other<br />
genera <strong>of</strong> Nemacheilidae by the following combination <strong>of</strong><br />
characters (but none unique to it): anterior and posterior<br />
nostrils adjacent; rim <strong>of</strong> anterior nostril as a short tube with<br />
posterior edge produced in a long filament (‘nasal barbel’);<br />
processus dentiformis present; lower lip with a median notch;<br />
lateral line present, incomplete, reaching vertical through end<br />
<strong>of</strong> anal-fin base; axillary pelvic lobe present, rudimentary;<br />
pelvic-fin origin about at vertical through dorsal-fin origin;<br />
anus closer to base <strong>of</strong> pelvic fin than to anal-fin origin; dorsal<br />
and ventral crests on caudal peduncle, supported by<br />
procurrent rays; posterior chamber <strong>of</strong> gas bladder large and<br />
somewhat globulous, located dorso-posterior to stomach and<br />
connected to encapsulated anterior part by a long slender<br />
duct.<br />
The ‘nasal barbel’ is also known in other nemacheilid species<br />
that had earlier been placed in Oreonectes (e.g. Bănărescu<br />
& Nalbant, 1995: 455) but are now placed in Oreonectes,<br />
Lefua, Indoreonectes and Sundoreonectes (Kottelat,<br />
1990b; Prok<strong>of</strong>iev, 2005). A few species <strong>of</strong> Schistura also<br />
have a ‘nasal barbel’, although less developed; they are immediately<br />
distinguished from Speonectes by their barred<br />
colour pattern, anus closer to base <strong>of</strong> anal-fin origin than to<br />
pelvic fin; caudal fin emarginated or forked; and (for those<br />
species in which this character has been examined) posterior<br />
chamber <strong>of</strong> gas bladder absent or immediately adjacent<br />
to encapsulated anterior part.<br />
Speonectes is distinguished from Oreonectes in having: an<br />
almost complete lateral line, which reaches to a vertical<br />
through the end <strong>of</strong> the base <strong>of</strong> the anal fin (vs. absent or up<br />
to 18 pores, not reaching pelvic fin; Du et al., 2008); a large<br />
posterior chamber <strong>of</strong> the air bladder, located dorso-posterior<br />
to the stomach and connected to the encapsulated anterior<br />
part by a long slender duct (vs. in direct contact with the<br />
bony capsule); the dorsal-fin origin about on a vertical<br />
through pelvic-fin origin (vs. conspicuously behind the base<br />
<strong>of</strong> the pelvic fin); adjacent nostrils (vs. widely separated,<br />
posterior one at short distance <strong>of</strong> anterior margin <strong>of</strong> orbit);<br />
and the anus closer to the base <strong>of</strong> the pelvic fins than to the<br />
anal-fin origin (vs. immediately in front <strong>of</strong> anal-fin origin).<br />
Oreonectes occurs in southeastern China and northern Vietnam.<br />
Speonectes is distinguished from Lefua in having: an almost<br />
complete lateral line, which reaches to a vertical through the<br />
end <strong>of</strong> the base <strong>of</strong> the anal fin (vs. no lateral line); the dorsal-<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
139<br />
fin origin about on a vertical through pelvic-fin origin (vs.<br />
above posterior extremity <strong>of</strong> pelvic-fin base); adjacent nostrils<br />
(vs. widely separated, posterior one at short distance <strong>of</strong><br />
anterior margin <strong>of</strong> orbit); and the anus closer to the base <strong>of</strong><br />
the pelvic fins than to the origin <strong>of</strong> the anal fin (vs. immediately<br />
in front <strong>of</strong> anal-fin origin). Lefua occurs in the Amur<br />
drainage, Korea, northeastern China and Japan.<br />
Speonectes is distinguished from Indoreonectes in having:<br />
an almost complete lateral line, which reaches a vertical<br />
through the end <strong>of</strong> the base <strong>of</strong> the anal fin (vs. reaching at<br />
most to a vertical through middle <strong>of</strong> pectoral fin); a large<br />
posterior chamber <strong>of</strong> the air bladder, located dorso-posterior<br />
to the stomach and connected to the anterior encapsulated<br />
part by a long slender duct (vs. rudimentary and adjacent<br />
to the bony capsule); the dorsal-fin origin about at a vertical<br />
through the pelvic-fin origin (vs. above the posterior extremity<br />
<strong>of</strong> the pelvic-fin base or behind); and the anus closer to<br />
the base <strong>of</strong> the pelvic fin than to the anal-fin origin (vs. immediately<br />
in front <strong>of</strong> anal-fin origin). Indoreonectes is known<br />
only from Peninsular India.<br />
The only known species <strong>of</strong> Speonectes (S. tiomanensis) inhabits<br />
a cave on Tioman Island (<strong>of</strong>f Malay Peninsula) and<br />
was earlier placed in the genus Sundoreonectes, which includes<br />
at least two species from the highlands <strong>of</strong> Borneo.<br />
Besides the features commonly observed in cavefishes (reduced<br />
eyes, white body, etc.), Speonectes is distinguished<br />
from Sundoreonectes in having: a median notch in the lower<br />
lip (vs. no notch, lip continuous along the anterior edge, but<br />
with numerous narrow furrows; postlabial groove almost<br />
continuous, interrupted medially only by two very narrow<br />
frena); an almost complete lateral line, which reaches to a<br />
vertical through the posterior extremity <strong>of</strong> the base <strong>of</strong> the<br />
anal fin (vs. reaching to above pelvic fin); the anus closer to<br />
the base <strong>of</strong> the pelvic fin than to the anal-fin origin (vs. immediately<br />
in front <strong>of</strong> at anal-fin origin); and the caudal fin<br />
slightly emarginate (vs. truncate or slightly rounded). Sundoreonectes<br />
was distinguished from Oreonectes, Lefua and<br />
Indoreonectes by the absence <strong>of</strong> a frontal fontanelle; it is<br />
not possible to dissect the single specimen <strong>of</strong> S. tiomanensis<br />
available to me to check this character, but no fontanelle<br />
could be felt through the skin with a needle.<br />
Etymology. — From the Greek σποζ (speos: a cave, cavern,<br />
grot) and νκτηζ (nectes: a swimmer); a reference to the cave<br />
habitat <strong>of</strong> the only known species <strong>of</strong> the genus. Gender masculine.<br />
Remarks. — Unpublished molecular data (V. Šlechtová and<br />
J. Bohlen, pers. comm.) shows that Speonectes tiomanensis<br />
belongs to a lineage very distinct from Oreonectes, Lefua<br />
and Sundoreonectes.<br />
Material. — Speonectes tiomanensis: CMK 15963, 1; Malaysia:<br />
Tioman Island. Indoreonectes evezardi: CMK 6439,<br />
2; India: Madhya Pradesh. Oreonectes platycephalus: CMK<br />
7207, 3; Hong Kong. - CMK 14892, 5; Vietnam: Quang Ninh.<br />
Sundorectes sabahensis: CMK 6435, 2; 17455, 10; Malaysia:<br />
Sarawak. Lefua costata: CMK 21951, 9; South Korea.
Fig. 1. Barbucca diabolica, ZRC 47747, 29.0 mm SL. Mouth.<br />
lj, lower jaw; llp, barbel-like projection <strong>of</strong> lower lip; ml, median<br />
lobe. (Photograph by Tan Heok Hui, retouched by author).<br />
Additional data from: Bănărescu & Nalbant, 1995; Prok<strong>of</strong>iev,<br />
2005; Du et al., 2008; Kottelat, 1990b.<br />
Barbuccidae, new family<br />
Type genus. — Barbucca Roberts, 1989: 100.<br />
Diagnosis. — Barbuccidae includes the single genus Barbucca,<br />
with one named and several unnamed species. The<br />
most obvious characters distinguishing Barbuccidae from the<br />
other families <strong>of</strong> Cobitoidea are in the structure <strong>of</strong> the mouth<br />
(Fig. 1) and the gas-bladder capsule. Upper lip, fleshy,<br />
smooth, with or without a median notch. Lower lip continuous<br />
with upper lip, fleshy, interrupted medially, with a thick,<br />
fleshy barbel-like projection along its posterior edge, near<br />
lateral extremity. Skin <strong>of</strong> throat between median extremities<br />
<strong>of</strong> lower lip forming a fleshy, triangular lobe, with a s<strong>of</strong>t,<br />
thin, compressed projection orientated perpendicular to body.<br />
Two pairs <strong>of</strong> rostral barbels, one pair <strong>of</strong> maxillary barbels.<br />
All barbels with rings <strong>of</strong> small projections. Lower jaw exposed.<br />
The shape <strong>of</strong> the gas-bladder capsule distinguishes Barbuccidae<br />
from all other lineages <strong>of</strong> Cobitoidea. The capsule<br />
appears as a single globulous structure in Cobitidae, incompletely<br />
formed in Botiidae and Vaillantellidae (Sawada,<br />
1982). In Nemacheilidae, Balitoridae, Gastromyzontidae<br />
(Sawada, 1982), Serpenticobitidae (Nalbant, 2001) and Ellopostomatidae<br />
(Kevin Conway, pers. comm., 2011) the capsule<br />
is made <strong>of</strong> two more or less spherical structures (formed<br />
by the horizontal and the descending processes <strong>of</strong> centrum 2<br />
and the inner and outer arms <strong>of</strong> the os suspensorium) connected<br />
posteriorly to each other to form a canal (called ‘manubrium’<br />
by, e.g., Bănărescu & Nalbant, 1995) around the duct<br />
connecting the left and right halves <strong>of</strong> the anterior chamber<br />
<strong>of</strong> the gas bladder. The manubrium is not a complete tube in<br />
Ellopostoma but a thin lamina <strong>of</strong> bone that rims the ventral<br />
part <strong>of</strong> the connecting duct. In Barbuccidae, the ‘manubrium’<br />
is missing, the two capsule are posteriorly independent<br />
and the duct is exposed (Fig. 2). The coracoid has a large<br />
posterior process, not reported in other families. Posterior<br />
edge <strong>of</strong> basipterygium <strong>of</strong> pelvic girdle with elongate poster-<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
lj<br />
ml<br />
llp<br />
140<br />
pcp<br />
d<br />
gbc<br />
gbp<br />
Fig. 2. Barbucca diabolica, UMMZ 240150, 24.4 mm SL. Ventral<br />
view <strong>of</strong> pectoral girdle. d, duct; gbc, gas-bladder capsule;<br />
gbp, gas-bladder posterior chamber; pcp, posterior coracoid process.<br />
(Photograph by Kevin Conway).<br />
odorsal process, which articulates with 2 nd last rib close to<br />
its base. (Based on photographs and information provided<br />
by Kevin Conway).<br />
Barbuccidae are further distinguished by having: body short,<br />
compact, with a barred colour pattern. Head small, with a<br />
blunt snout, with large, oval eyes orientated dorso-laterally.<br />
Anterior nostril at tip <strong>of</strong> a short tube; posterior nostril in<br />
contact with base <strong>of</strong> tube and adjacent to eye orbit. Caudal<br />
fin emarginate. Dorsal-fin origin slightly in advance <strong>of</strong> pelvic-fin<br />
origin. Pectoral and pelvic fins directed laterally, each<br />
with a single simple ray. Large tubercles disposed irregularly<br />
on lower half <strong>of</strong> flank above anal fin (number and exact<br />
distribution variable and depending <strong>of</strong> species).<br />
Serpenticobitidae, new family<br />
Type genus. — Serpenticobitis Roberts, 1997: 109.<br />
Diagnosis. — Serpenticobitidae includes the single genus<br />
Serpenticobitis, with three named species. It is distinguished<br />
from the other families <strong>of</strong> Cobitoidea in having: body short,<br />
cylindrical, with a barred colour pattern. Head small, snout<br />
rounded and conspicuously overhanging. A large bifid spine<br />
present immediately below eye. Caudal fin slightly forked.<br />
Dorsal-fin origin in advance <strong>of</strong> pelvic-fin origin. Pectoral<br />
fins directed laterally. Pectoral and pelvic fins with a single<br />
simple ray. Mouth small, arched. Upper and lower lips fleshy<br />
and continuous. Lower lip with a few folds and a median<br />
interruption but no mental lobes. Two pairs <strong>of</strong> rostral barbels,<br />
one pair <strong>of</strong> maxillary barbels. Lower jaw partly exposed.<br />
Anterior nostril pierced at front side <strong>of</strong> a flap-like<br />
tube. All fins with conspicuous rows <strong>of</strong> spots. A large and<br />
conspicuous black spot at midheight <strong>of</strong> caudal-fin base. Gasbladder<br />
capsule in two parts connected by a ‘manubrium’<br />
(Nalbant, 2001: 10).<br />
Remarks. — I follow here the molecular phylogeny <strong>of</strong> Šlech-
tová et al. (2007) and Bohlen & Šlechtová (2009). However,<br />
their phylogeny recovers Ellopostomatidae, Nemacheilidae<br />
and Balitoridae as an unresolved trichotomy and their<br />
Balitoridae is quite heterogeneous and includes a number <strong>of</strong><br />
very distinctive groups. In their Balitoridae, they recovered<br />
four lineages corresponding to Gastromyzontidae, ‘Homalopteridae’<br />
<strong>of</strong> earlier authors (e.g., Silas, 1953), Barbucca<br />
and Serpenticobitis. The position <strong>of</strong> these two genera has<br />
been very uncertain since their description, with Barbucca<br />
being placed in Cobitidae (Roberts, 1989) or in Balitoridae<br />
(Kottelat et al., 1993; Šlechtová et al., 2007), and Serpenticobitis<br />
hypothesized to belong to Cobitidae (Roberts, 1997;<br />
Kottelat, 2001b), Nemacheilidae (Nalbant, 2002: pl. 7) or<br />
Balitoridae (Šlechtová et al., 2007). As more characters (morphological<br />
and molecular) are explored and as data on more<br />
species become available, these two genera are likely to continue<br />
their Brownian movement within the phylogeny <strong>of</strong><br />
Cobitoidei. I find it convenient to give these two genera the<br />
It is impossible to mention all those who provided information<br />
and literature over the years <strong>of</strong> gestation <strong>of</strong> this checklist,<br />
but in the final years the following helped in finding<br />
rare or hard-to-get literature and helped with translations,<br />
information and comments: Nina Bogutskaya (ZISP), Jörg<br />
Bohlen (Institute <strong>of</strong> Animal Physiology and Genetics,<br />
Liběchov, Czechia), Ralf Britz (BMNH), Chen I-Shiung<br />
(National Taiwan Ocean University, Keelung, Taiwan), Chen<br />
Xiao-Yong (KIZ), Kevin Conway (Texas A&M University,<br />
College Station, USA), Fang Fang (NRM), Jörg Freyh<strong>of</strong><br />
(Leibniz-Institute <strong>of</strong> Freshwater Ecology and Inland Fisheries,<br />
Berlin, Germany), Alain Hennache (Parc de Clères,<br />
Clères, France), Kim Ik-Soo (CNUC), Katsuma Kubota<br />
(Bangkok, Thailand), Sven Kullander (NRM), Kelvin Lim<br />
(ZRC), Michal Mikšik (Prague), Sonia Fisch-Muller<br />
(MHNG), Ng Heok Hee (ZRC), Rohan Pethiyagoda (AMS),<br />
Graham Proudlove (University <strong>of</strong> Manchester, U. K.), Vendula<br />
Šlechtová (Institute <strong>of</strong> Animal Physiology and Genetics,<br />
Liběchov, Czechia), Tan Heok Hui (ZRC), Tran Anh Duc<br />
(Hanoi University <strong>of</strong> Science, Hanoi, Vietnam), Davut Turan<br />
(Rize University, Rize, Turkey), Ekaterina Vasil’eva<br />
(ZMMU), Claude Weber (MHNG), Yang Jian (Guangxi<br />
Teacher’s Education University, Nanning, China), Zhang E<br />
(IHB). Kevin Conway generously made available a number<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
ACKNOWLEDGEMENTS<br />
141<br />
same rank as the two other lineages (Gastromyzontidae and<br />
Balitoridae s.s.).<br />
Some might prefer recognising a hierarchy <strong>of</strong> families, subfamilies,<br />
tribes etc. and are free to do so. At this stage, I do<br />
not see any advantage at recognising a hierarchy <strong>of</strong> ranks<br />
within the still unstable Nemacheilidae-Balitoridae-Ellopostomatidae<br />
assemblage (see Introduction, Families vs. subfamilies).<br />
The classification with a single rank allows easier<br />
reference for those not concerned by phylogeny or taxonomy.<br />
It is easier (for example for ecologists) to handle units<br />
with some sort <strong>of</strong> internal consistency in morphology, habitat<br />
preferences, etc. with only one level <strong>of</strong> family-group<br />
names, without having to worry about a hierarchy <strong>of</strong> names<br />
or debates on phylogeny. A classification using (naming) few<br />
ranks only is less quickly obsolete than a classification using<br />
a complex hierarchy.<br />
<strong>of</strong> unpublished osteological information and photographs<br />
used in the diagnoses <strong>of</strong> the new taxa. Kevin Conway, Jörg<br />
Bohlen and Jörg Freyh<strong>of</strong> read and commented on the manuscript.<br />
Rohan Pethiyagoda commented on the manuscript and<br />
helped in making it more readable. Hazelina Yeo Hwan Theng<br />
read and checked the manuscript. Jörg Bohlen, Chen Xiao-<br />
Yong, Kevin Conway, Jörg Freyh<strong>of</strong>, Alexander Golubtsov<br />
(Severtsov Institute <strong>of</strong> Ecology and Evolution, Moscow,<br />
Russia), Kazumi Hosoya (Kinki University, Nara, Japan),<br />
Zdenek Lajbner (Institute <strong>of</strong> Animal Physiology and Genetics,<br />
Liběchov, Czechia), Lan Jia-Hu (Fishery Technique Popularization<br />
Station, Du’an, China), Beta Mahatvaraj (Chennai,<br />
India), Artem Prok<strong>of</strong>iev (Severtsov Institute <strong>of</strong> Ecology<br />
and Evolution, Moscow, Russia), Ren Qiu (KIZ), Tan Heok<br />
Hui, Yang Jian, Zhang E, Zhao Ya-Hui (ASIZB) and Zakhar<br />
Zhidkov (Saint Petersburg University, Russia) provided photographs.<br />
Tan Heok Hui designed the cover page.<br />
Over the years, Antoinette Kottelat-Kloetzli contributed to<br />
my work on loaches by assisting in library work, sorting and<br />
shelving collections, supporting my frequent and long absences<br />
while in the field, coining a number <strong>of</strong> names <strong>of</strong> new<br />
taxa (and building a stock <strong>of</strong> names available for future new<br />
taxa), and many other ways.
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
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Zhu, S.-Q., 1982a. [Five new species <strong>of</strong> fishes <strong>of</strong> the genus<br />
Nemachilus from Yunnan Province, China]. Acta Zootaxonomica<br />
Sinica, 7(1): 104–111. [In Chinese, English<br />
summary].<br />
Zhu, S.-Q., 1982b. [A new species <strong>of</strong> the genus Nemachilus<br />
(Pisces: Cobitidae) from Qinghai Province, China]. Acta<br />
Zootaxonomica Sinica, 7(2): 223–224. [In Chinese, English<br />
summary].<br />
Zhu, S.-Q., 1983. [A new genus and species <strong>of</strong> Nemachilinae<br />
(Pisces: Cobitidae) from China]. Acta Zootaxonomica<br />
Sinica, 8(3): 311–313. [In Chinese, English summary].<br />
Zhu, S.-Q., 1989. [The loaches <strong>of</strong> the subfamily Nemacheilinae<br />
in China (Cypriniformes: Cobitidae)]. Jiangsu Sci-<br />
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[In Chinese, English summary].<br />
Zhu, S.-Q., 1992. [Three new species <strong>of</strong> nemacheilinae fishes<br />
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Acta Zootaxonomica Sinica, 12(3): 323–331. [In Chinese,<br />
English summary].<br />
Zhu, S.-Q. & W.-X. Cao., 1988. [Descriptions <strong>of</strong> two new<br />
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Zhu, S.-Q. & Q.-Z. Guo, 1985. [Descriptions <strong>of</strong> a new genus<br />
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Zhu, Y., Y.-J. Lu, J.-X. Yang & S. Zhang, 2008. A new blind<br />
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Annals and Magazine <strong>of</strong> Natural History, Ser. 8, 10(60):<br />
595–599.
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
INDEX<br />
Subspecies and infrasubspecific names are listed under their generic name in alphabetical sequence and with omission <strong>of</strong><br />
intercallated names (e.g. Acanthophthalmus kuhlii malayanus is listed as Acanthophthalmus malayanus). Accidental misspelling<br />
<strong>of</strong> generic names are corrected. Numbers in bold refer to valid names <strong>of</strong> valid species.<br />
A<br />
abbreviata, Homaloptera 7.9.1<br />
abbreviata, Jinshaia 7.9.1<br />
Aborichthys 10.1<br />
— bijulensis 10.1<br />
— elongatus 10.1.1, 10.30.3<br />
— garoensis 10.1.2<br />
— kempi 10.1, 10.1.3<br />
— rosammai 10.34.154<br />
— tikaderi 10.1.4<br />
abyssinicus, Afronemacheilus 10.3.1<br />
abyssinicus, Nemachilus 10.3, 10.3.1<br />
Acanestrinia 4.5<br />
Acanthocobitis 10.2<br />
— botia 2.2.7, 10.2.2, 10.2.3<br />
— longipinnis 10.2, 10.2.1, 10.2.5, 10.21.14<br />
— mandalayensis 10.2.3<br />
— mooreh 10.2.4<br />
— pavonacea 10.2.5<br />
— pictilis 10.2.6<br />
— rubidipinnis 10.2.3, 10.2.7<br />
— urophthalma 10.2.8<br />
— zonalternans 10.2.9<br />
Acanthophthalmus 4.5, 4.16<br />
— agmus 4.16.1<br />
— anguillaris 4.16, 4.16.5<br />
— borneensis 4.16.1<br />
— cuneovirgatus 4.16.9<br />
— fasciatus 4.16, 4.16.16<br />
— goaensis 4.16.14<br />
— javanicus 4.16.24<br />
— longipinnis 4.10.5<br />
— lorentzi 4.10.14<br />
— mariae 4.16.21<br />
— malayanus 4.16.20<br />
— muraeniformis 4.16.22<br />
— myersi 4.16.23<br />
— pahangensis 4.1.1<br />
— robiginosus 4.16.28<br />
— sandakanensis 4.21, 4.21.1<br />
— semicinctus 4.16.29<br />
— shelfordii 4.16.30<br />
— sumatranus 4.16.29<br />
— unistriatus 4.10.10<br />
— vermicularis 4.16.5<br />
Acanthopis berdmorei 4.10.5<br />
— micropogon 4.10.15<br />
Acanthopsides 4<br />
Acanthopsis 4, 4.2, 4.5<br />
Acanthopsis angustus 4.5<br />
— biaculeata 4.2.1<br />
— lachnostoma 4.5.1<br />
— linea 4.5.29<br />
— paludica 4.5<br />
Acanthopsoides 4.1<br />
— delphax 4.1.1<br />
— gracilentus 4.1.2<br />
— gracilis 4.1, 4.1.3<br />
— hapalias 4.1.4<br />
— molobrion 4.1.5<br />
— namromensis 4.1.2<br />
— robertsi 4.1.6<br />
Acantophthalmus 4.5, 4.16<br />
— fasciatus 4.16.16<br />
— javanicus 4.16.24<br />
— superbus 4.16.32<br />
Acantopsis 4, 4.2, 4.5<br />
— arenae 4.5.3<br />
— dialuzona 4.2, 4.2.1<br />
— guttatus 4.5.3<br />
— multistigmatus 4.2.2<br />
— octoactinotos 4.2.2<br />
— spectabilis 4.2.3<br />
— thiemmedhi 4.2.4<br />
Acoura 10.34<br />
Acourus 10.34<br />
aculeata, Schistura 4.10.9<br />
Acura 10.34<br />
acuticauda, Sinohomaloptera 7.5.16<br />
acuticephala, Homatula 10.11.1<br />
acuticephala, Paracobitis 10.11.1<br />
acuticephala, Schistura 10.34.3<br />
acuticephalus, Nemachilus 10.34.3<br />
Adiposia 10, 10.26<br />
Adiposiidae 10<br />
Aenobarbus 4.10<br />
aenobarbus, Allotrius 4.10<br />
aequabilis, Gastromyzon 8.5.1<br />
aeroides, Gastromyzon 8.5.2<br />
afasciata, Schistura 10.34.4, 10.34.13<br />
afghana, Nemachilus 10.42.65<br />
afrenatus, Nemachilus 10.25.16<br />
Afronemacheilus 10.3<br />
— abyssinicus 10.3.1<br />
— kaffa 10.3<br />
agma, Pangio 4.16.1<br />
agmus, Acanthophthalmus 4.16.1<br />
agrensis, Platacanthus 4.10, 4.10.16<br />
176<br />
aizawlensis, Schistura 10.34.5<br />
ajdan, Cobitis 4.13.4<br />
ajmonis, Nemachilus 10.14.7<br />
akhtari, Nemachilus 10.42.1<br />
akhtari, Triplophysa 10.42.1<br />
alasanicus, Nemachilus 10.25.25<br />
albescens, Canthophrys 4.4, 4.4.1<br />
albicoloris, Cobitis 4.20.4<br />
albisuera, Sewellia 8.16.1<br />
alcoides, Pangio 4.16.2<br />
alepidota, Paraschistura 10.28.1<br />
alepidotus, Noemacheilus 10.28.1<br />
alexandrae, Triplophysa 10.42.2<br />
aliensis, Triplophysa 10.42.3<br />
alkaia, Lepidocephalichthys 4.10.1<br />
Allotrius aenobarbus 4.10<br />
almorhae, Botia 2.2, 2.2.1<br />
alta, Schistura 10.34.6<br />
altayensis, Barbatula 10.4.1<br />
alternans, Pangio 4.16.3<br />
alticeps, Nemacheilus 10.33<br />
alticeps, Nemachilus 10.33.1<br />
alticeps, Qinghaichthys 10.33.1<br />
alticrista, Schistura 10.34.7<br />
altipedunculata, Schistura 10.34.8<br />
altipedunculatus, Noemacheilus 10.34.8<br />
altipinnis, Triplophysa 10.42.4<br />
altus, Yunnanilus 10.46.1<br />
aluensis, Triplophysa 10.42.5<br />
Ambastaia 2.1<br />
— nigrolineata 2.1.1<br />
— sidthimunki 2.1.2<br />
ammophila, Pangio 4.16.4<br />
amnicola, Cobitis 4.4.1<br />
amphilekta, Cobitis 4.5.4<br />
amphisquamata, Homaloptera 7.8.1<br />
amphisquamata, Homalopterula 7.8.1<br />
amplizona, Schistura 10.34.9<br />
amudarjensis, Dzihunia 10.7.1<br />
amudarjensis, Nemacheilus 10.7<br />
amudarjensis, Nemachilus 10.7.1<br />
analis, Sewellia 8.16.2<br />
analis, Yunnanilus 10.46.2<br />
anambarensis, Noemacheilus 10.34.10<br />
anambarensis, Schistura 10.34.10<br />
anatoliae, Cobitinula 4.5, 4.5.61<br />
anatolicus, Oxynoemacheilus 10.25.1<br />
andrewsi, Lefua 10.17.1<br />
angeli, Nemacheilus 10.4.8, 10.42.6
angeli, Triplophysa 10.42.6<br />
anglicana, Nemacheilus 10.4.2<br />
angorae, Nemachilus 10.25.2<br />
angorae, Oxynoemacheilus 10.25.2<br />
anguilla, Nemacheilus 10.21.1<br />
anguillaris, Acanthophthalmus 4.16, 4.16.5<br />
anguillaris, Pangio 4.16.5<br />
anguillicaudata, Cobitis 4.13.1<br />
anguillicaudatus, Misgurnus 4.13.1, 4.18.1<br />
anguillioides, Homatula 10.11.2<br />
anguillioides, Paracobitis 10.11.2<br />
angustus, Acanthopsis 4.5<br />
anisura, Platycara 7.1.2<br />
Annamia 8.1<br />
— normani 8.1.1<br />
— thuathienensis 8.1.1<br />
annamitica, Balitora 7, 7.1.1<br />
annandalei, Bhavania 7.3.1<br />
annandalei, Lepidocephalichthys 4.10.2<br />
anophthalmus, Oreonectes 10.24.6<br />
antennata, Schistura 10.34.11<br />
anterodorsalis, Triplophysa 10.42.7<br />
Aperioptus 4.2<br />
— megalomycter 5.1. 5.1.1<br />
— pictorius 4.2, 4.2.1<br />
aphelocheilus, Protomyzon 8.14.1<br />
apoda, Pangio 4.16.6<br />
Apua 4.16<br />
— fusca 4.16, 4.16.13<br />
aquaecaeruleae, Triplophysa 10.42.8,<br />
10.42.40<br />
arachthosensis, Cobitis 4.5.5<br />
aralensis, Cobitis 4.20.1<br />
aramis, Schistura 10.34.12<br />
araxensis, Orthrias 10.25.3<br />
araxensis, Oxynoemacheilus 10.25.3<br />
arenae, Acantopsis 4.5.3<br />
arenae, Cobitis 4.5.2<br />
arenae, Misgurnus 4.5.2<br />
arenata, Cobitis 10.2.2<br />
arenicolus, Nemacheilus 10.21.2<br />
argentata, Cobites 10.2.2<br />
argyrogramma, Cobitis 10.25.4<br />
argyrogramma, Oxynoemacheilus 10.25.4<br />
arifi, Schistura 10.34.4, 10.34.13,<br />
10.34.42, 10.34.97<br />
arnoldii, Triplophysa 10.42.9<br />
arunachalensis, Bhavania 7.1<br />
arunachalensis, Lepidocephalichthys 4.10.3<br />
arunachalensis, Noemacheilus 4.10.3,<br />
10.34.184<br />
assamensis, Neonoemacheilus 10.23.1<br />
assamensis, Noemacheilus 10.23.1<br />
atactos, Pangio 4.16.7<br />
athos, Schistura 10.34.14<br />
atra, Schistura 10.34.15<br />
atriceps, Noemacheilus 10.35, 10.35.1<br />
atriceps, Sectoria 10.35.1<br />
aurantiaca, Schistura 10.34.16<br />
aurata, Cobitis 4.20.1<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
aurata, Sabanejewia 4.20.1<br />
aureus, Nemacheilus 10.2.4<br />
auronigrus, Gastromyzon 8.5.3<br />
australis, Bhavania 7.3, 7.3.1<br />
australis, Platycara 7.3.1<br />
aymonieri, Gyrinocheilus 1.1.1<br />
aymonieri, Psilorhynchus 1.1.1<br />
B<br />
bachmaensis, Schistura 10.34.17<br />
bacmeensis, Micronemacheilus 10.41.2<br />
badamensis, Nemachilus 10.15.1<br />
baenzigeri, Nemacheilus 10.44<br />
baenzigeri, Noemacheilus 10.44.1<br />
baenzigeri, Tuberoschistura 10.44.1<br />
bairdi, Schistura 10.34.18<br />
bajiangensis, Yunnanilus 10.46.3<br />
balcanica, Cobitis 4.20, 4.20.2<br />
balcanica, Sabanejewia 4.20.2<br />
balgara, Cobitis 4.10.9<br />
Balitora 7, 7.1<br />
— annamitica 7, 7.1.1<br />
— brucei 7.1, 7.1.2<br />
— burmanica 7.1.3, 7.1.12<br />
— burmanicus 7.1.3<br />
— eddsi 7.1.4<br />
— elongata 7.5.3<br />
— erythrorhina 7.6.5<br />
— haithanhi 7.1.5<br />
— heteroura 7.1.5<br />
— kwangsiensis 7.1.5<br />
— lancangjiangensis 7.1.6<br />
— laticauda 7.1.7<br />
— lineolata 8.16, 8.16.6<br />
— longibarbata 7.1.9<br />
— ludongensis 7.1.8<br />
— maculata 7.1.2<br />
— melanosoma 7.1.3<br />
— meridionalis 7.1.10<br />
— mysorensis 7.1.11<br />
— nantingensis 7.1.12<br />
— nigrocorpa 7.1.5<br />
— nujiangensis 7.5.9<br />
— ocellata 7.7.6<br />
— pavonina 7.6.5<br />
— pengi 7.5.11<br />
— shimogensis 7.1.7<br />
— tchangi 7.5.15<br />
— vanlani 7.1.5<br />
— vanlongi 7.1.6<br />
Balitoridae 7<br />
Balitoropsis 7.2<br />
— bartschi 7.2, 7.2.10<br />
— batek 7.2.1<br />
— leonardi 7.2.2<br />
— ophiolepis 7.2.3<br />
— sexmaculata 7.2.4<br />
— stephensoni 7.2.5<br />
— tatereganii 7.2.6<br />
— vulgaris 7.2.7<br />
177<br />
Balitoropsis yunnanensis 7.2.8<br />
— yuwonoi 7.2.9<br />
— zollingeri 7.2.10<br />
balteata, Schistura 10.34.19<br />
balteatus, Nemacheilus 10.34.19<br />
baltica, Sabanejewia 4.20.3<br />
baluchiorum, Nemachilus 10.34.20<br />
baluchiorum, Schistura 10.34.20<br />
bamaensis, Paraprotomyzon 8.11.1<br />
bampurenris, Paraschistura 10.28.2<br />
bampurensis, Nemacheilus 10.28.2<br />
bampurensis, Paraschistura 10.34.20<br />
banar, Nemacheilus 10.21.3<br />
banarescui, Leptobotia 2.5.1<br />
banarescui, Orthrias 10.25.5<br />
banarescui, Oxynoemacheilus 10.25.5<br />
banarescui, Parabotia 2.5.1<br />
Bangfaia 9.1<br />
bannaensis, Schistura 10.34.21<br />
baotingensis, Plesiomyzon 8.13, 8.13.1<br />
barapaniensis, Noemacheilus 10.30.3<br />
Barbatula 10.4<br />
— altayensis 10.4.1<br />
— barbatula 10.4.2, 10.4.15<br />
— bergamensis 10.25.40<br />
— caud<strong>of</strong>urca 10.34.33<br />
— cinica 10.25.10<br />
— cobdoensis. 10.4.4<br />
— cobdonensis 10.4.3<br />
— compressirostris 10.4.4<br />
— cuneicephalus 10.42.21<br />
— dgebuadzei 10.4.5<br />
— erdali 10.25.13<br />
— fowleri 10.4.8<br />
— germencica 10.25.18<br />
— golubtsovi 10.4.3, 10.4.4, 10.4.6<br />
— incerta 10.34.74<br />
— kirinensis 10.4.8<br />
— markakulensis 10.4.4, 10.4.7, 10.4.15<br />
— mediterraneus 10.25.27, 10.36.1<br />
— nuda 10.4.8, 10.4.15, 10.42.86<br />
— oreas 10.4.9, 10.4.15<br />
— orthocauda 10.34.126<br />
— paucilepis 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
— phoxinoides 10.25.34<br />
— pindus 10.25.35<br />
— posteroventralis 10.42.22<br />
— potaninorum 10.4.10<br />
— quignardi 10.4.11<br />
— sawadai 10.4.12<br />
— sellaefer 10.42.91<br />
— shansi 10.4.8<br />
— sturanyi 10.4.13<br />
— tigris 10.4.14<br />
— tomiana 10.4.4, 10.4.14, 10.4.15<br />
— tomianus 10.4.15<br />
— toni 10.4.3, 10.4.4, 10.4.8, 10.4.14,<br />
10.4.15<br />
— uniformis 10.34.74
Barbatula zetensis 10.4.16<br />
barbatula, Barbatula 10.4.2, 10.4.15<br />
barbatula, Cobitis 10.4, 10.4.2<br />
barbatuloides, Cobitis 4.10, 4.10.4<br />
barbatuloides, Lepidocephalichthys 4.10.4<br />
Barbucca 6, 6.1<br />
— diabolica 6.1, 6.1.1<br />
Barbuccidae 6<br />
bario, Gastromyzon 8.5.4<br />
bartschi, Balitoropsis 7.2, 7.2.10<br />
bashai, Pangio 4.16<br />
bashanensis, Triplophysa 10.42.10<br />
batek, Balitoropsis 7.2.1<br />
batek, Homaloptera 7.2.1<br />
battalgili, Cobitis 4.5.41<br />
beauforti, Botia 2.7.1<br />
beauforti, Syncrossus 2.7.1<br />
Beaufortia 8, 8.2<br />
— buas 8.2.1<br />
— cyclica 8.2.2, 8.2.4<br />
— daon 8.2.3<br />
— elongata 8.2.4<br />
— fasciolata 8.2.16<br />
— gracilicauda 8.2.7<br />
— huangguoshuensis 8.2.5<br />
— intermedia 8.2.6<br />
— kweichowensis 8.2.7<br />
— leveretti 8.2.8<br />
— liui 8.2.9<br />
— loos 8.2.10<br />
— multiocellata 8.2.16<br />
— niulanensis 8.2.11<br />
— pingi 8.2.12<br />
— polliciformis 8.2.2<br />
— polylepis 8.2.13<br />
— szechuanensis 8.2.14<br />
— triocellata 8.2.16<br />
— yunnanensis 8.2.15<br />
— zebroida 8.2.16<br />
Beaufortini 8<br />
beavani, Nemachilus 10.34.22<br />
beavani, Schistura 10.34.22<br />
beipanjiangensis, Yunnanilus 10.46.4<br />
bella, Schistura 10.34.23<br />
bellibarus, Nemachilus 10.42.11<br />
bellibarus, Triplophysa 10.42.11<br />
berdmorei, Acanthopis 4.10.5<br />
berdmorei, Lepidocephalichthys 4.10.5<br />
berdmorei, Syncrossus 2.7, 2.7.2<br />
berezowskii, Homatula 10.11.3<br />
berezowskii, Nemacheilus 10.11.3<br />
bergamensis, Barbatula 10.25.40<br />
bergi, Nemacheilus 10.25.25<br />
bergianus, Nemachilus 10.25.6<br />
bergianus, Oxynoemacheilus 10.25.6<br />
bertini, Nemacheilus 10.42.86<br />
Beyshehiria 4.5<br />
Bhavania 7.3<br />
— annandalei 7.3.1<br />
— arunachalensis 7.1<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Bhavania australis 7.3.1<br />
bhimachari, Nemachilus 10.34.24<br />
bhimachari, Schistura 10.34.24<br />
biaculeata, Acanthopsis 4.2.1<br />
Bibarba 4.3<br />
— bibarba 4.3, 4.3.1<br />
bibarba, Bibarba 4.3, 4.3.1<br />
Bicanestrinia 4.5<br />
bifurcata, Cobitis 4.13.1<br />
bijulensis, Aborichthys 10.1<br />
bilineata, Cobitis 4.5.6<br />
bilineata, Helgia 7.6<br />
bilineata, Homaloptera 7.6.1<br />
bilseli, Cobitis 4.5, 4.5.7<br />
bilturio, Cobitis 10.2.2<br />
bimaculatus, Parabotia 2.5.2<br />
bimucronata, Cobitis 10.2.2<br />
binotatus, Nemacheilus 10.21.4<br />
bipartitus, Nemacheilus 4.13<br />
bipartitus, Nemachilus 4.13.4<br />
birdi, Botia 2.2.2<br />
birmanicus, Lepidocephalus 4.10.5<br />
bispinosa, Somileptes 4.4, 4.4.1<br />
bitaimac, Pangio 4.16.8<br />
biwae, Cobitis 4.5.8<br />
blackiana, Nemacheilus 10.4.2<br />
bleekeri, Nemachilus 10.42.12<br />
bleekeri, Triplophysa 10.42.12<br />
blythi, Botia 2.2.1<br />
blythii, Nemacheilus 10<br />
bolavenensis, Schistura 10.34.25<br />
bombifrons, Nemachilus 10.40<br />
bombifrons, Tarimichthys 10.40.1<br />
borneensis, Acanthophthalmus 4.16.1<br />
borneensis, Gastromyzon 8.5, 8.5.5<br />
borneensis, Protomyzon 8.14.2<br />
bosniaca, Cobitis 4.20.2<br />
Botia 1, 2.2, 2.7<br />
— almorhae 2.2, 2.2.1<br />
— beauforti 2.7.1<br />
— birdi 2.2.2<br />
— blythi 2.2.1<br />
— caudipunctata 2.8.1<br />
— citrauratea 2.4.1<br />
— compressicauda 2.4.13<br />
— dario 2.2.3<br />
— dayi 2.2.7<br />
— elongata 2.4, 2.4.1, 2.5.5<br />
— eos 2.8.2<br />
— fangi 2.4.10<br />
— formosa 2.7.5<br />
— gigantea 2.6.2<br />
— grandis 2.2.1, 2.7<br />
— helodes 2.7.3<br />
— hexafurca 2.6.4<br />
— histrionica 2.2.4<br />
— horae 2.8.6<br />
— javedi 2.2.2<br />
— kolhapurensis 2.2.8<br />
— kubotai 2.2.5<br />
178<br />
Botia kwangsiensis 2.5.6<br />
— lecontei 2.8.3<br />
— lohachata 2.2.6<br />
— longidorsalis 2.84<br />
— longiventralis 2.6.1<br />
— lucasbahi 2.7.5<br />
— macrolineata 2.2.3<br />
— maculosa 2.5.10<br />
— modesta 2.8, 2.8.5<br />
— morleti 2.8.6<br />
— multifasciata 2.5.6<br />
— nebulosa 2.2.7<br />
— nigrolineata 2.1, 2.1.1<br />
— pratti 2.4.10<br />
— pulchra 2.6.2<br />
— pulchripinnis 2.8.3<br />
— reevesae 2.6.3<br />
— reversa 2.7.6<br />
— robusta 2.6.4<br />
— rostrata 2.2.7<br />
— rubripinnis 2.8.5<br />
— sidthimunki 2.1.2<br />
— splendida 2.8.7<br />
— striata 2.2.8<br />
— superciliaris 2.6, 2.6.5<br />
— tientainensis 2.4.13<br />
— udomritthiruji 2.2.9<br />
— variegata 2.4<br />
— wui 2.5.6<br />
— yunnanensis 2.7.5<br />
— zebra 2.6.6<br />
botia, Acanthocobitis 2.2.7, 10.2.2, 10.2.3<br />
botia, Cobitis 10.2.2<br />
Botiidae 1<br />
boutanensis, Cobitis 10.26.1<br />
boutanensis, Paracobitis 10.26.1<br />
brachyptera, Triplophysa 10.42.13<br />
brachypterus, Nemachilus 10.42.13<br />
brahui, Nemachilus 10.42.14<br />
brahui, Triplophysa 10.42.14<br />
brandtii, Nemachilus 10.25.7<br />
brandtii, Oxynoemacheilus 10.25.4, 10.25.7<br />
brevibarba, Triplophysa 10.42.15<br />
brevibarbus, Nemachilus 10.9.4, 10.42.15<br />
brevibarbus, Triplophysa 10.42.15<br />
brevicauda, Nemachilus 10.42.16<br />
brevicauda, Triplophysa 10.42.16<br />
breviceps, Noemacheilus<br />
breviceps, Schistura 10.34.26<br />
brevifasciata, Kichulchoia 4.7.1<br />
brevifasciata, Niwaella 4.7, 4.7.1<br />
brevirostris, Parabotia 2.5.3<br />
brevis, Nemachilus 10.29, 10.29.2<br />
brevis, Petruichthys 10.29.2<br />
brevis, Sewellia 8.16.8<br />
brevis, Yunnanilus 10.46<br />
breviventralis, Sewellia 8.16.3<br />
brucei, Balitora 7.1, 7.1.2<br />
brunei, Neogastromyzon 8.10.1<br />
brunneana, Physoschistura 10.30.1
unneanus, Nemacheilus 10.30<br />
brunneanus, Nemachilus 10.30.1<br />
buas, Beaufortia 8.2.1<br />
buas, Gastromyzon 8.2.1<br />
bucculenta, Schistura 10.34.27<br />
bucculentus, Noemacheilus 10.34.27<br />
bulgarica, Cobitis 4.20.4<br />
bulgarica, Sabanejewia 4.20.4<br />
buphoensis, Misgurnus 4.13.**<br />
bureschi, Nemacheilus 10.25.8<br />
bureschi, Oxynoemacheilus 10.25.8<br />
burmanica, Balitora 7.1.3, 7.1.12<br />
burmanicus, Balitora 7.1.3<br />
C<br />
cakaensis, Triplophysa 10.42.17, 10.42.100<br />
calderoni, Cobitis 4.5.9<br />
caldwelli, Homaloptera 8.17.5<br />
caldwelli, Vanmanenia 8.17.5<br />
callichroma, Schistura 10.34.28<br />
callichromus, Noemacheilus 10.34.28<br />
callidora, Schistura 10.34.29<br />
cambodgiensis, Tuberoschistura 10.44.2<br />
Canthophrys 4.4<br />
— albescens 4.4, 4.4.1<br />
— flavicauda 2.2.3<br />
— gongota 4.4.1<br />
— rubiginosus 4.16.25<br />
— unispina 10.2.2<br />
— vittatus 4.10.9<br />
— zebra 2.2, 2.2.3<br />
— olivaceus 4.10.9<br />
caobangensis, Vanmanenia 8.17.6<br />
caohaiensis, Yunnanilus 10.46.5<br />
carassus, Nemachilus 10.42.86<br />
carbonaria, Schistura 10.34.30<br />
carletoni, Nemacheilus 10.34.31<br />
carletoni, Schistura 10.34.31<br />
carnaticus, Cobitis 4.10.16<br />
caspia, Cobitis 4.20.5<br />
caspia, Sabanejewia 4.20.5<br />
cataracta, Schistura 10.34.32<br />
cataractus, Lepidocephalus 4.10.5<br />
caucasica, Cobitis 4.20.6<br />
caucasica, Sabanejewia 4.20.6<br />
caucasicus, Nemachilus 10.4.2<br />
caudipunctata, Botia 2.8.1<br />
caudipunctata, Yasuhikotakia 2.8.1<br />
caud<strong>of</strong>urca, Barbatula 10.34.33<br />
caud<strong>of</strong>urca, Schistura 10.34.33<br />
caud<strong>of</strong>urcatus, Lepidocephalus 4.10.8<br />
cestoideus, Misgurnus 4.13.4<br />
ceyhanensis, Oxynoemacheilus 10.25.9<br />
ceyhanensis, Schistura 10.25.15, 10.25.32<br />
ceyhanensis, Schistura 10.25.9, 10.25.38<br />
chandagaitensis, Triplophysa 10.42.37<br />
changtingensis, Pseudogastromyzon 8.15.1<br />
cha-ny, Cobitis 2.4.11<br />
chapaensis, Nemacheilus 10.34.34<br />
chapaensis, Schistura 10.34.34<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
chapaensis, Sinogastromyzon 7.13.1<br />
chekianensis, Homaloptera 8.17.14<br />
cheni, Nemacheilus 2.4<br />
cheni, Pseudogastromyzon 8.15.2<br />
chenyiyui, Crossostoma 8.4.1<br />
chenyiyui, Formosania 8.4.1<br />
chindwinensis, Physoschistura 10.30.2<br />
chindwinica, Schistura 10.34.35<br />
chindwinicus, Nemacheilus 10.34.35<br />
chini, Neogastromyzon 8.10.2<br />
chlorosoma, Cobitis 10.2.9<br />
Choia 4.7<br />
choii, Cobitis 4.5.10, 4.5.27<br />
choirorhynchos, Cobitis 4.2.1<br />
chondrostoma, Nemachilus 10.42.18<br />
chondrostoma, Triplophysa 10.42.18<br />
choprai, Indotriplophysa 10.14.1<br />
choprai, Nemachilus 10.14.1<br />
Chopraia 7.7.7<br />
— rupicola 7.7, 7.7.3<br />
choresmi, Nemachilus 10.7.1<br />
Chromobotia 2.3<br />
— macracanthus 2.3.1<br />
chryseus, Nemacheilus 10.34.47<br />
chrysicristinae, Paraschistura 10.28.3<br />
chrysicristinae, Schistura 10.28.3<br />
chrysolaimos, Cobitis 10.21.5<br />
chrysolaimos, Nemacheilus 10.21.5<br />
chui, Yunnanilus 10.46.6<br />
cincticauda, Cobitis 10.34.36<br />
cincticauda, Schistura 10.34.36<br />
cinerea, Cobites 10.21.7<br />
cingulata, Serpenticobitis 9.1.1<br />
cinica, Barbatula 10.25.10<br />
cinicus, Oxynoemacheilus 10.25.10<br />
cinnamomea, Cobitis 4.16, 4.16.25<br />
cinnamomea, Vaillantella 3.1.1<br />
cirrhifurcata, Cobitis 4.13.1<br />
citrauratea, Botia 2.4.1<br />
Claea 10.5<br />
— dabryi 10.5.1, 10.42.6<br />
— niulanjiangensis 10.5.2<br />
clatrata, Schistura 10.34.37<br />
cleopatra, Nemacheilus 10.21.6<br />
cobdoensis, Barbatula 10.4.4<br />
cobdonensis, Barbatula 10.4.3<br />
cobdonensis, Nemachilus 10.4.3<br />
Cobites 10.4<br />
— argentata 10.2.2<br />
— cinerea 10.21.7<br />
— obscura 10.34.157, 10.34.189<br />
Cobitichthys 4.10<br />
— dichachrous 4.13.1<br />
— enalios 4.13.1<br />
— polynema 4.13.1<br />
Cobitidae 4<br />
Cobitinula 4.5<br />
— anatoliae 4.5, 4.5.61<br />
Cobitis 4, 4.5<br />
— ajdan 4.13.4<br />
179<br />
Cobitis albicoloris 4.20.4<br />
— amnicola 4.4.1<br />
— amphilekta 4.5.4<br />
— anguillicaudata 4.13.1<br />
— arachthosensis 4.5.5<br />
— aralensis 4.20.1<br />
— arenae 4.5.2<br />
— arenata 10.2.2<br />
— argyrogramma 10.25.4<br />
— aurata 4.20.1<br />
— balcanica 4.20, 4.20.2<br />
— balgara 4.10.9<br />
— barbatula 10.4, 10.4.2<br />
— barbatuloides 4.10, 4.10.4<br />
— battalgili 4.5.41<br />
— bifurcata 4.13.1<br />
— bilineata 4.5.6<br />
— bilseli 4.5, 4.5.7<br />
— bilturio 10.2.2<br />
— bimucronata 10.2.2<br />
— biwae 4.5.8<br />
— bosniaca 4.20.2<br />
— botia 10.2.2<br />
— boutanensis 10.26.1<br />
— bulgarica 4.20.4<br />
— calderoni 4.5.9<br />
— carnaticus 4.10.16<br />
— caspia 4.20.5<br />
— caucasica 4.20.6<br />
— cha-ny 2.4.11<br />
— chlorosoma 10.2.9<br />
— choii 4.5.10, 4.5.27<br />
— choii<br />
— choirorhynchos 4.2.1<br />
— chrysolaimos 10.21.5<br />
— cincticauda 10.34.36<br />
— cinnamomea 4.16, 4.16.25<br />
— cirrhifurcata 4.13.1<br />
— conspersa 4.20.8<br />
— corica 10.21.7<br />
— cucura 4.4.1<br />
— curta 2.5.4<br />
— dalmatina 4.5.11<br />
— danubialis 4.5.15<br />
— dario 2.2, 2.2.3<br />
— decemcirrosus 4.13.1<br />
— delicata 4.15, 4.15.1<br />
— dolichorhynchus 4.5.12<br />
— dorsalis 10.42.24<br />
— elata 4.20.2<br />
— elazigensis 4.5.13<br />
— elegans 10.42.27<br />
— elongata 4.5, 4.5.14, 4.5.48, 4.20.2<br />
— elongatoides 4.5.15<br />
— erythropterus 4.13.1<br />
— evreni 4.5.16<br />
— fahireae 4.5.17<br />
— faridpaki 4.5.18<br />
— fasciata 10.21, 10.21.9<br />
— fossilis 4.13, 4.13.3
Cobitis frenata 10.25.16<br />
— furstenbergii 10.4.2, 10.4.2<br />
— galilaea 10.25, 10.25.17<br />
— geto 2.2.3<br />
— gibbosa 10.2.2<br />
— gladkovi 4.5.19, 4.5.34<br />
— gongota 4.4.1<br />
— grandis 2.7<br />
— granoei 4.5.48<br />
— guntea 4.10.9<br />
— guttata 4.5.3, 4.10.9<br />
— haasi 4.5.40<br />
— haematopterus 4.13.1<br />
— hankugensis 4.5.20<br />
— hasselti 4.10, 4.10.10<br />
— hellenica 4.5.21<br />
— heteroclita 10.4<br />
— he-tsieou 4.13.1<br />
— hohenackeri 4.20.1<br />
— hymenophysa 2.7.4<br />
— illyrica 4.5.22<br />
— insignis 10.25.20<br />
— jadovaensis 4.5.23<br />
— jaklesii 10.21.10<br />
— japonica 4.5.8<br />
— kellei 4.5.24<br />
— keyvani 4.5.25<br />
— koreensis 4.6, 4.6.2<br />
— kuhlii 4.16.16<br />
— kurui 4.5.62<br />
— lachnostoma 4.5.1<br />
— laoensis 4.5.26<br />
— larvata 4.20.8<br />
— laterimaculata 4.15.2<br />
— lebedevi 4.5.27<br />
— leopardus 10.25.31<br />
— levantina 4.5.28<br />
— linea 4.5.29<br />
— longicauda 10.26, 10.26.4<br />
— longicorpus 4.6.3<br />
— longimanus 4.5<br />
— longitaeniatus 4.5.26<br />
— lutheri 4.5.30<br />
— macracanthus 2.3, 2.3.1<br />
— macrochir 4.11, 4.11.1<br />
— macrorhynchos 4.2.1<br />
— macrostigma 4.5.31<br />
— maculata 4.13.1<br />
— malapterura 10.26, 10.26.5<br />
— marmorata 10.42.65<br />
— maroccana 4.5.32<br />
— matsubarae 4.5.33<br />
— maya 4.10.9<br />
— megaspila 4.5.15<br />
— melanoleuca 4.5.19, 4.5.34, 4.5.48<br />
— merga 10.25.28<br />
— meridionalis 4.5.35<br />
— microcephala 4.5.36<br />
— microps 10.42.68<br />
— micropus 4.13.1<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Cobitis misgurnoides 4.12, 4.12.1<br />
— mohoity 4.13.4<br />
— monocera 10.2.2<br />
— montana 4.20.2<br />
— montanus 4.20.2, 10<br />
— mooreh 10.2.4<br />
— multifasciatus 4.7.2<br />
— multimaculata 4.5.37<br />
— mysorensis 4.10.16<br />
— narentana 4.5.38<br />
— nuicocensis 4.12.1<br />
— nurga 10.25.28<br />
— ny-tsieou 4.13.1<br />
— oblonga 4.16.24<br />
— obscura 10.34<br />
— ocellata 10.2.2<br />
— octocirrhus 4.10.10<br />
— oculata 4.4.1<br />
— ohridana 4.5.39<br />
— olivai 4.5.48<br />
— pacifica 4.6.4<br />
— paludica 4.5.40<br />
— paludicola 4.5.40<br />
— pangia 4.16.25<br />
— panthera 10.25.31<br />
— parisiensis 10.4.2<br />
— pavonacea 10.2.5<br />
— pectoralis 4.13.1<br />
— persa 10.25, 10.25.33<br />
— peschevi 4.5.54<br />
— pfeifferi 10.21.25<br />
— phoxocheila 4.10.9<br />
— phrygica 4.5.41<br />
— pictava 10.4.2<br />
— pironae 10.4.2<br />
— poecilopleura 4.5<br />
— pontica 4.5.42<br />
— psammismus 4.13.1<br />
— pumilus 4.6.5<br />
— puncticulata 4.5.43<br />
— punctilineata 4.5.44<br />
— puta 4.5.6<br />
— radnensis 4.20.2<br />
— rara 4.5.45<br />
— rhodopensis 4.5.54<br />
— romanica 4.20.9<br />
— rossomeridionalis 4.5.57<br />
— rotundicaudata 4.8, 4.8.2<br />
— rubidipinnis 4.10.16, 4.13.1, 10.2.7<br />
— rubripinnis 4.10.16, 4.13.1<br />
— ruppelli 10.22<br />
— rupelli 10.22.1<br />
— satunini 4.5.46<br />
— savona 10.34.157<br />
— schlegeli 4.5.8<br />
— semizonata 10.2.7<br />
— septa 4.5.6<br />
— shikokuensis 4.5.47<br />
— sibirica 4.5.34, 4.5.48<br />
— simplicispina 4.5, 4.5.13, 4.5.41, 4.5.49<br />
180<br />
Cobitis sinensis 4.5.50<br />
— spiloptera 10.34.176<br />
— spilura 4.5.55<br />
— splendens 4.5.51<br />
— squataeniatus 4.5.64<br />
— stephanidisi 4.5.52<br />
— stolickai 10.42.100<br />
— striata 4.5.53<br />
— strumicae 4.5.54<br />
— suborbitalis 10.21.9<br />
— taenia 4.5, 4.5.55, 10.4<br />
— taenioides 4.5.55<br />
— takatsuensis 4.5.56<br />
— tanaitica 4.5.57<br />
— taurica 4.5.58<br />
— tenuicauda 10.14.5<br />
— tesselatus 4.20.4<br />
— tetralineata 4.5.59<br />
— thermalis 4.10.16<br />
— tigris 10.25.41<br />
— toni 10.4.15<br />
— trichonica 4.5.60<br />
— turcica 4.5.61<br />
— turio 10.2.2<br />
— uranoscopus 10.42.107<br />
— vallachica 4.20.10<br />
— vardarensis 4.5.62<br />
— variabilis 10.4.2<br />
— variegata 2.4, 2.4.1<br />
— vettonica 4.5.63<br />
— victoriae 4.5.40<br />
— vitoriae 4.5.40<br />
— vittata 10.42.65<br />
— yeni 4.5<br />
— ylengensis 4.5.64<br />
— zanandreai 4.5.65<br />
— zhejiangensis 4.5.66<br />
— zonalternans 10.2, 10.2.9<br />
Cobitophis 4.16<br />
— perakensis 4.16.5<br />
compactus, Erromyzon 8.3.2<br />
compressicauda, Botia 2.4.13<br />
compressirostris, Barbatula 10.4.4<br />
compressirostris, Nemachilus 10.4.4<br />
confluens, Hemimyzon 7.5.1<br />
confuzona, Homaloptera 7.6.2<br />
coniptera, Triplophysa 10.42.19<br />
conipterus, Nemachilus 10.42.19<br />
conirostris, Nemachilus 10.34.38<br />
conirostris, Schistura 10.34.38<br />
conjungens, Nemachilus 10.16.4<br />
conspersa, Cobitis 4.20.8<br />
contractus, Gastromyzon 8.5.6<br />
coreanus, Elxis 10.17.1<br />
corica, Cobitis 10.21.7<br />
corica, Nemacheilus 10.21.7<br />
cornusaccus, Gastromyzon 8.5.7<br />
coromandelensis, Lepidocephalichthys<br />
4.10.6<br />
coruscans, Schistura 10.34.39
costata, Diplophysa 10.17.1<br />
costata, Lefua 10.17.1<br />
crabro, Schistura 10.34.40<br />
cranbrooki, Gastromyzon 8.5.8<br />
crassicauda, Indotriplophysa 10.14.2<br />
crassicauda, Nemachilus 10.14.2<br />
crassicauda, Vanmanenia 8.17.7<br />
crassilabris, Triplophysa 10.42.20<br />
crassiobex, Neogastromyzon 8.10.3<br />
crassipedunculatus, Oreias 10.5.1<br />
crenastus, Gastromyzon 8.5.9<br />
cristata, Metaschistura 10.19.1<br />
cristatus, Nemacheilus 10.19, 10.19.1<br />
crossochilus, Misgurnus 4.13.1<br />
Crossostoma 8, 8.4<br />
— chenyiyui 8.4.1<br />
— davidi 8.4, 8.4.2<br />
— fangi 8.15, 8.15.3<br />
— fascicauda 8.4.3<br />
— fasciolatus 8.4.4<br />
— foochowensis 8.4.2<br />
— galericula 8.4.5<br />
— lacustre 8.4.6<br />
— paucisquama 8.4.7<br />
— stigmata 8.4.8<br />
— tengi 8.4.9<br />
— tinkhami 8.4.10<br />
Crossostominae 8<br />
cruciatus, Micronemacheilus 10.20.1<br />
cruciatus, Nemacheilus 10.20, 10.20.1<br />
crypt<strong>of</strong>asciata, Schistura 10.34.41<br />
Cryptotora 7.4<br />
— thamicola 7.4.1<br />
ctenocephalus, Gastromyzon 8.5.10<br />
cucura, Cobitis 4.4.1<br />
cueljuensis, Nemachilus 10.42.25<br />
cuneicephala, Triplophysa 10.42.21<br />
cuneicephalus, Barbatula 10.42.21<br />
cuneovirgata, Pangio 4.16.9<br />
cuneovirgatus, Acanthophthalmus 4.16.9<br />
curta, Cobitis 2.5.4<br />
curtistigma, Schistura 10.34.13, 10.34.42<br />
curtus, Parabotia 2.5.4<br />
cyclica, Beaufortia 8.2.2, 8.2.4<br />
cyri, Nemachilus 10.25.11<br />
cyri, Oxynoemacheilus 10.25.11<br />
D<br />
dabryanus, Paramisgurnus 4.13.1, 4.18,<br />
4.18.1<br />
dabryi, Claea 10.5.1, 10.42.6<br />
dabryi, Oreias 10.5, 10.5.1<br />
dabryi, Schistura 10.5.1<br />
dalaica, Diplophysa 14.40.22<br />
dalaica, Triplophysa 10.42.22<br />
dalatensis, Schistura 10.34.43<br />
dalmatina, Cobitis 4.5.11<br />
danubialis, Cobitis 4.5.15<br />
danumensis, Gastromyzon 8.5.11<br />
daon, Beaufortia 8.2.3<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
daon, Gastromyzon 8.2.3<br />
daon, Sinogastromyzon 7.13.2<br />
daqiaoensis, Triplophysa 10.42.23<br />
dario, Botia 2.2.3<br />
dario, Cobitis 2.2, 2.2.3<br />
dattai, Noemacheilus 10.34.184<br />
daubentoni, Schistura 10.34.44<br />
davidi, Crossostoma 8.4, 8.4.2<br />
davidi, Formosania 8.4.2<br />
dayi, Botia 2.2.7<br />
dayi, Nemachilus 10.34.47<br />
deansmarti, Schistura 10.34.45<br />
decemcirrosus, Cobitis 4.13.1<br />
defectiva, Schistura 10.34.46<br />
deignani, Noemacheilus 10.34.87<br />
delacouri, Oreoglanis 7.7.4<br />
delicata, Cobitis 4.15, 4.15.1<br />
delicata, Niwaella 4.15.1<br />
delphax, Acanthopsoides 4.1.1<br />
denisoni, Nemacheilus 10.34.47<br />
denisoni, Schistura 10.34.47, 10.34.111<br />
denudata, Glaniopsis 8.6.1<br />
desmotes, Nemacheilus 10.34.48<br />
desmotes, Schistura 10.34.48<br />
deterrai, Nemachilus 10.42.34<br />
Deuterophysa 10.42<br />
devdevi, Nemachilus 10.34.49<br />
devdevi, Schistura 10.34.49<br />
dezeensis, Sinogastromyzon 7.13.3<br />
dgebuadzei, Barbatula 10.4.5<br />
dgebuadzei, Orthrias 10.4.5<br />
diabolica, Barbucca 6.1, 6.1.1<br />
Diacantha 2.2<br />
Diacanthus 2.2<br />
dialuzona, Acantopsis 4.2, 4.2.1<br />
dianchiensis, Sphaerophysa 10.38, 10.38.1<br />
diardi, Sewellia 8.16, 8.16.4<br />
Diardichthys 8.16<br />
dibruensis, Lepidocephalus 4.10.9<br />
dichachrous, Cobitichthys 4.13.1<br />
Didymophysa 10.42<br />
Dienbienia 7.5<br />
— namnuaensis 7.5, 7.5.11<br />
diminuta, Schistura 10.34.50<br />
dingi, Triplophysa 10.42.15<br />
Diplophysa costata 10.17.1<br />
— dalaica 14.40.22<br />
— herzensteini 10.16.2<br />
— intermedia 10.42.46<br />
— kungessana 10.42, 10.42.51<br />
— labiata 10.16, 10.16.4<br />
— microphthalma 10.16.5, 10.42.67<br />
— nasalis 10.16.1<br />
— papillosolabiata 10.42.78<br />
— pedaschenkoi 10.42.82<br />
— stewarti 10.42.99<br />
— strauchii 10.42, 10.42.101<br />
Diplophysoides 10.42<br />
discoloris, Yunnanilus 10.46.7<br />
disparis, Liniparhomaloptera 8.9.1<br />
181<br />
disparis, Parhomaloptera 8.9, 8.9.1<br />
disparizona, Schistura 10.34.51<br />
dixoni, Nemacheilus 10.17.1<br />
djaggasteensis, Nemachilus 10.42.22<br />
doiranica, Sabanejewia 4.20.2<br />
dolichopterus, Lepturichthys 7.10.1<br />
dolichorhynchus, Cobitis 4.5.12<br />
doonensis, Noemacheilus 10.34.52<br />
doonensis, Schistura 10.34.52<br />
dori, Orthrias 10.25.20<br />
doriae, Eucirrhichthys 4.16, 4.16.10<br />
doriae, Pangio 4.16.10<br />
dorsalis, Cobitis 10.42.24<br />
dorsalis, Triplophysa 10.42.24<br />
dorsaloides, Nemacheilus 10.42.106<br />
dorsizona, Schistura 10.34.53<br />
dorsonotata, Triplophysa 10.42.25,<br />
10.42.100<br />
dorsonotatus, Nemachilus 10.42.25<br />
Draconectes 10.6<br />
Draconectes narinosus 10.6.1<br />
drassensis, Noemacheilus 10.42.26<br />
drassensis, Triplophysa 10.42.26<br />
dubia, Schistura 10.34.54<br />
dubius, Parabotia 2.5.5<br />
dunckeri, Nemachilus 10.21.25<br />
Dzihunia 10.7<br />
— amudarjensis 10.7.1<br />
— ilan 10.7.2<br />
— turdakovi 10.7.3<br />
E<br />
ecdyonuroides, Hemimyzon 7.5.2<br />
echigonia, Lefua 10.17.2<br />
eddsi, Balitora 7.1.4<br />
edsinica, Triplophysa 10.40.2<br />
edsinicus, Tarimichthys 10.40.2<br />
elakatis, Yunnanilus 10.46.8<br />
elata, Cobitis 4.20.2<br />
elazigensis, Cobitis 4.5.13<br />
elegans, Cobitis 10.42.27<br />
elegans, Triplophysa 10.42.27<br />
elegantissimus, Nemacheilus 10.21.8<br />
Ellopostoma 5, 5.1<br />
— megalomycter 5.1.1<br />
— mystax 5.1.2<br />
Ellopostomatidae 5<br />
elongata, Balitora 7.5.3<br />
elongata, Beaufortia 8.2.4<br />
elongata, Botia 2.4, 2.4.1, 2.5.5<br />
elongata, Cobitis 4.5, 4.5.14, 4.5.48, 4.20.2<br />
elongata, Leptobotia 2.4.1, 2.4.7<br />
elongata, Pangio 4.16.11<br />
elongata, Physoschistura 10.30.3<br />
elongata, Sewellia 8.16.5<br />
elongata, Travancoria 7.14.1<br />
elongatoides, Cobitis 4.5.15<br />
elongatus, Aborichthys 10.1.1, 10.30.3<br />
elongatus, Gastromyzon 8.2.4<br />
elongatus, Hemimyzon 7.5.3
elongatus, Misgurnus 4.13.1<br />
elongatus, Nemachilus 10.1.1, 10.30.3,<br />
10.42.77<br />
elongatus, Oreonectes 10.24.1<br />
Elxis 10.17<br />
— coreanus 10.17.1<br />
— nikkonis 10.17, 10.17.4<br />
— sabanus 10.39.2<br />
embalohensis, Gastromyzon 8.5.12<br />
enalios, Cobitichthys 4.13.1<br />
Enobarbichthys 4.10<br />
Enobarbus 4.10<br />
Eonemachilus 10.8, 10.46<br />
— longidorsalis 10.8.1<br />
— nigromaculatus 10.8.2<br />
— yangzonghaiensis 10.8.3<br />
eos, Botia 2.8.2<br />
eos, Yasuhikotakia 2.8.2<br />
ephelis, Schistura 10.34.55<br />
ercisianus, Orthrias 10.25.12<br />
ercisianus, Oxynoemacheilus 10.25.12<br />
erdali, Barbatula 10.25.13<br />
erdali, Oxynoemacheilus 10.25.13<br />
eregliensis, Orthrias 10.25.14<br />
eregliensis, Oxynoemacheilus 10.25.14<br />
erhaiensis, Homatula 10.11.4<br />
erhaiensis, Paracobitis 10.11.4<br />
erikssoni, Misgurnus 4.13.4<br />
Erromyzon 8.3<br />
— compactus 8.3.2<br />
— pachychilus 8.3.1<br />
— sinensis 8.3.3<br />
— yangi 8.3.4<br />
erythrinna, Nemacheilus 10.4.2<br />
erythropterus, Cobitis 4.13.1<br />
erythrorhina, Balitora 7.6.5<br />
eubranchus, Hypergastromyzon 8.7.1<br />
Eucirrhichthys 4.16<br />
— doriae 4.16, 4.16.10<br />
euepiptera, Vaillantella 3.1.2<br />
euepipterus, Nemacheilus 3.1, 3.1.2<br />
eugeniae, Indotriplophysa 10.14.3<br />
eugeniae, Triplophysa 10.14.3<br />
euphraticus, Noemacheilus 10.25.4<br />
evezardi, Indoreonectes 10.13.1<br />
evezardi, Nemacheilus 10.13.1<br />
evreni, Cobitis 4.5.16<br />
evreni, Oxynoemacheilus 10.25.15<br />
evreni, Schistura 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
extrorsus, Gastromyzon 8.5.13<br />
F<br />
fahireae, Cobitis 4.5.17<br />
fangi, Botia 2.4.10<br />
fangi, Crossostoma 8.15, 8.15.3<br />
fangi, Pseudogastromyzon 8.15.3<br />
faridpaki, Cobitis 4.5.18<br />
farragus, Gastromyzon 8.5.14<br />
farsicus, Orthrias 10.25.33<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
farwelli, Nemachilus 10.42.28<br />
farwelli, Triplophysa 10.42.28<br />
fasciata, Cobitis 10.21, 10.21.9<br />
fasciata, Homaloptera 7.7.6<br />
fasciata, Parabotia 2.5<br />
fasciata, Schistura 10.34.56<br />
fasciatus, Acanthophthalmus 4.16, 4.16.16<br />
fasciatus, Gastromyzon 8.5.15<br />
fasciatus, Nemacheilus 10.21.9<br />
fasciatus, Noemacheilus 10.21.9<br />
fasciatus, Parabotia 2.5.6<br />
fasciatus, Pseudogastromyzon 8.15.4<br />
fasciatus, Psilorhynchus 8.15.4<br />
fascicauda, Crossostoma 8.4.3<br />
fascicauda, Formosania 8.4.3<br />
fascimaculata, Schistura 10.34.57<br />
fasciolata, Beaufortia 8.2.16<br />
fasciolata, Formosania 8.4.4<br />
fasciolata, Homaloptera 10.34.58<br />
fasciolata, Schistura 10.34.58<br />
fasciolatus, Crossostoma 8.4.4<br />
fedtschenkoae, Nemachilus 10.42.107<br />
filinaris, Pangio 4.16.12<br />
fimbriata, Homaloptera 7.10, 7.10.2<br />
fimbriatus, Lepturichthys 7.10.2<br />
finis, Schistura 10.34.59<br />
flavicauda, Canthophrys 2.2.3<br />
flavicorpus, Triplophysa 10.42.29<br />
flav<strong>of</strong>asciata, Vaillantella 3.1.3<br />
flavolineata, Leptobotia 2.4.2<br />
flavus, Nemachilus 10.25<br />
foochowensis, Crossostoma 8.4.2<br />
forkicaudalis, Yunnanilus 10.46.14<br />
formosa, Botia 2.7.5<br />
formosana, Homaloptera 7.5, 7.5.4<br />
Formosania 8.4<br />
— chenyiyui 8.4.1<br />
— davidi 8.4.2<br />
— fascicauda 8.4.3<br />
— fasciolata 8.4.4<br />
— galericula 8.4.5<br />
— gilberti 8.4.6<br />
— lacustris 8.4.6<br />
— paucisquama 8.4.7<br />
— stigmata 8.4.8<br />
— tengi 8.4.9<br />
— tinkhami 8.4.10<br />
— yaoshanensis 8.17.11<br />
formosanum, Homaloptera 8.4.6<br />
formosanus, Hemimyzon 7.5.4<br />
formosanus, Misgurnus 4.13.1<br />
fossilis, Cobitis 4.13, 4.13.3<br />
fossilis, Misgurnus 4.13.3<br />
fowleri, Barbatula 10.4.8<br />
fowlerianus, Noemacheilus 10.34.162<br />
frenata, Cobitis 10.25.16<br />
frenatus, Oxynoemacheilus 10.25.16<br />
fukien, Misgurnus 4.13.1<br />
furcatus, Lepidocephalichthys 4.10.7<br />
furcatus, Oreias 10.5.1<br />
182<br />
furcocaudalis, Oreonectes 10.24.2<br />
furstenbergii, Cobitis 10.4.2<br />
furva, Triplophysa 10.42.30<br />
fusca, Apua 4.16, 4.16.13<br />
fusca, Pangio 4.16.13<br />
fusinotata, Schistura 10.34.60<br />
fuxianensis, Triplophysa 10.42.31<br />
G<br />
galericula, Crossostoma 8.4.5<br />
galericula, Formosania 8.4.5<br />
galilaea, Cobitis 10.25, 10.25.17<br />
galilaeus, Oxynoemacheilus 10.25.17<br />
gangetica, Schistura 10.34.61<br />
gangeticus, Noemacheilus 10.34.61<br />
ganheensis, Yunnanilus 10.46.9<br />
garoensis, Aborichthys 10.1.2<br />
Gastromyzon 8, 8.5<br />
— aequabilis 8.5.1<br />
— aeroides 8.5.2<br />
— auronigrus 8.5.3<br />
— bario 8.5.4<br />
— borneensis 8.5, 8.5.5<br />
— buas 8.2.1<br />
— contractus 8.5.6<br />
— cornusaccus 8.5.7<br />
— cranbrooki 8.5.8<br />
— crenastus 8.5.9<br />
— ctenocephalus 8.5.10<br />
— danumensis 8.5.11<br />
— daon 8.2.3<br />
— elongatus 8.2.4<br />
— embalohensis 8.5.12<br />
— extrorsus 8.5.13<br />
— farragus 8.5.14<br />
— fasciatus 8.5.15<br />
— ingeri 8.5.16<br />
— introrsus 8.5.17<br />
— katibasensis 8.5.18<br />
— kweichowensis 8.2.7<br />
— lepidogaster 8.5.19<br />
— leveretti 8.2, 8.2.8<br />
— loos 8.2.10<br />
— megalepis 8.5.20<br />
— monticola 8.5.21<br />
— ocellatus 8.5.22<br />
— ornaticauda 8.5.23<br />
— pariclavis 8.5.24<br />
— pauciradiatus 8.10.6<br />
— pingi 8.2.12<br />
— praestans 8.5.25<br />
— psiloetron 8.5.26<br />
— punctulatus 8.5.27<br />
— ridens 8.5.28<br />
— russulus 8.5.29<br />
— scitulus 8.5.30<br />
— spectabilis 8.5.31<br />
— stellatus 8.5.32<br />
— szechuanensis 8.2.14<br />
— umbrus 8.5.33
Gastromyzon venustus 8.5.34<br />
— viriosus 8.5.35<br />
— zebrinus 8.5.36<br />
— zebroidus 8.2.16<br />
Gastromyzontidae 8<br />
geisleri, Schistura 10.34.62<br />
gejiuensis, Noemacheilus 10.42.32<br />
gejiuensis, Triplophysa 10.42.32<br />
genilepis, Paranemachilus 10.27, 10.27.1<br />
germencica, Barbatula 10.25.18<br />
germencicus, Oxynoemacheilus 10.25.18<br />
gerzeensis, Triplophysa 10.42.33<br />
geto, Cobitis 2.2.3<br />
ghazniensis, Noemacheilus 10.26.2<br />
ghazniensis, Paracobitis 10.26.2<br />
gibbosa, Cobitis 10.2.2<br />
gibbusnazus, Nemachilus 10.25.7<br />
gigantea, Botia 2.6.2<br />
gilberti, Formosania 8.4.6<br />
gladkovi, Cobitis 4.5.19, 4.5.34<br />
Glaniopsini 8<br />
Glaniopsis 8, 8.6<br />
— denudata 8.6.1<br />
— gossei 8.6.2<br />
— hanitschi 8.6, 8.6.3<br />
— multiradiata 8.6.4<br />
globiceps, Schistura 10.34.63<br />
goaensis, Acanthophthalmus 4.16.14<br />
goaensis, Pangio 4.16.14<br />
goalparensis, Lepidocephalichthys 4.10.8<br />
Gobius pa-che-tsee-yu 8.17.12<br />
— tsin-ting-yu 7.5.16<br />
Gojeb 10<br />
golubtsovi, Barbatula 10.4.3, 10.4.4, 10.4.6<br />
golubtsovi, Orthrias 10.4.6<br />
gongota, Canthophrys 4.4.1<br />
gongota, Cobitis 4.4.1<br />
gossei, Glaniopsis 8.6.2<br />
gracilentus, Acanthopsoides 4.1.2<br />
gracilentus, Neacanthopsis 4.1, 4.1.2<br />
gracilicauda, Beaufortia 8.2.7<br />
gracilis, Acanthopsoides 4.1, 4.1.3<br />
gracilis, Nemacheilus 10.42.34<br />
gracilis, Triplophysa 10.42.34<br />
grahami, Nemachilus 10.42.35<br />
grahami, Triplophysa 10.42.35<br />
grandis, Botia 2.2.1, 2.7<br />
grandis, Cobitis 2.7<br />
grandis, Schistura 2.7<br />
grangeri, Misgurnus 4.13.1<br />
granoei, Cobitis 4.5.48<br />
griffithii, Nemachilus 10.42.36<br />
griffithii, Triplophysa 10.42.36<br />
griswoldi, Progastromyzon 8.14, 8.14.3<br />
griswoldi, Protomyzon 8.14.3<br />
grummorum, Hedinichthys 10.9.1<br />
guananensis, Oreonectes 10.24.7<br />
guentheri, Lepturichthys 7.10.2<br />
guentheri, Mesonoemacheilus 10.18.1<br />
guentheri, Nemacheilus 10.18.1<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
guilinensis, Leptobotia 2.4.3<br />
guishanensis, Paralepidocephalus 4.17<br />
gundriseri, Triplophysa 10.42.37<br />
guntea, Cobitis 4.10.9<br />
guntea, Lepidocephalichthys 4.10.9<br />
guttata, Cobitis 4.5.3, 4.10.9<br />
guttatus, Acantopsis 4.5.3<br />
gymnetrus, Vanmanenia 8.17.8<br />
gymnogaster, Homaloptera 7.8.2<br />
gymnogaster, Homalopterula 7.8.2<br />
Gyrinocheilidae 1<br />
Gyrinocheilops 1.1<br />
— pennocki 1.1, 1.1.2<br />
Gyrinocheilus 1, 1.1<br />
— aymonieri 1.1.1<br />
— kaznakovi 1.1.1<br />
— monchadskii 1.1.1<br />
— pennocki 1.1.2<br />
— pustulosus 1.1, 1.1.3<br />
H<br />
haarlovi, Nemacheilus 10.28.6<br />
haasi, Cobitis 4.5.40<br />
haematopterus, Cobitis 4.13.1<br />
hagiangensis, Paracobitis 10.34.64<br />
hagiangensis, Schistura 10.34.64<br />
hagiangensis, Sinogastromyzon 7.13.4<br />
hainan, Misgurnus 4.13.1<br />
hainanensis, Vanmanenia 8.17.9<br />
haithanhi, Balitora 7.1.5<br />
hamwii, Nemacheilus 10.25.19<br />
hamwii, Oxynoemacheilus 10.25.19<br />
hangshuiensis, Metahomaloptera 7.11.1<br />
hanitschi, Glaniopsis 8.6, 8.6.3<br />
hankugensis, Cobitis 4.5.20<br />
hansuiensis, Leptobotia 2.4.13<br />
hapalias, Acanthopsoides 4.1.4<br />
harnaiensis, Noemacheilus 10.34.65<br />
harnaiensis, Schistura 10.34.65<br />
hasselti, Cobitis 4.10, 4.10.10<br />
hasselti, Lepidocephalichthys 4.10.10<br />
hazaraensis, Nemacheilus 10.42.36<br />
Hedinichthys 10.9, 10.42<br />
— grummorum 10.9.1<br />
— macropterus 10.9.2<br />
— minutus 10.9.3<br />
— yarkandensis 10.9.2, 10.9.4<br />
Helgia 7.6<br />
— bilineata 7.6<br />
— modesta 7.7.1<br />
hellenica, Cobitis 4.5.21<br />
helodes, Botia 2.7.3<br />
helodes, Syncrossus 2.7.3<br />
Hemimyzon 7, 7.5<br />
— confluens 7.5.1<br />
— ecdyonuroides 7.5.2<br />
— elongatus 7.5.3<br />
— formosanus 7.5.4<br />
— khonensis 7.5.5<br />
— macroptera 7.5.6<br />
183<br />
Hemimyzon macropterus 7.5.6<br />
— megalopseos 7.5.7<br />
— nanensis 7.5.8<br />
— nujiangensis 7.5.9<br />
— papilio 7, 7.5.10<br />
— pengi 7.5.11<br />
— pumilicorpora 7.5.12<br />
— sheni 7.5.13<br />
— songamensis 7.1.5<br />
— taitungensis 7.5.14<br />
— tchangi 7.5.15<br />
— yaotanensis 7.5.16<br />
— zebroidus 8.15, 8.15.4<br />
Heminoemacheilus 10.8, 10.10, 10.36, 10.46<br />
— hyalinus 10.10.1<br />
— zhengbaoshani 10.10, 10.10.2<br />
hengyangensis, Leptobotia 2.4.4<br />
herrei, Mesonoemacheilus 10.18.2<br />
herzensteini, Diplophysa 10.16.2<br />
herzensteini, Labiatophysa 10.16.2<br />
heterocheila, Parabotia 2.5.7<br />
heterocheilus, Parabotia 2.5.7<br />
heteroclita, Cobitis 10.4<br />
heterognathos, Schistura 10.35.2<br />
heterognathos, Sectoria 10.35.2<br />
heterolepis, Homaloptera 7.8.3<br />
heterolepis, Homalopterula 7.8.3<br />
heteroura, Balitora 7.1.5<br />
he-tsieou, Cobitis 4.13.1<br />
heungchow, Misgurnus 4.13.1<br />
hexafurca, Botia 2.6.4<br />
hexaocellum, Sinogastromyzon 7.13.1<br />
hexiensis, Nemachilus 10.42.38<br />
hexiensis, Triplophysa 10.42.38, 10.42.100<br />
heyangensis, Triplophysa 10.42.39<br />
hialmari, Triplophysa 10.42.40<br />
himachalensis, Noemacheilus 10.34.66<br />
himachalensis, Schistura 10.34.66<br />
himalaya, Turcinoemacheilus 10.45.1<br />
hingi, Homaloptera 10.34.67<br />
hingi, Schistura 10.34.67<br />
hipporhynchos, Kottelatlimia 4.9.1<br />
hispanica, Nemacheilus 10.4.11<br />
histrionica, Botia 2.2.4<br />
hoai, Oreias 10.34.68<br />
hoai, Schistura 10.34.68, 10.34.134<br />
h<strong>of</strong>fmanni, Lefua 10.17.3<br />
h<strong>of</strong>fmanni, Sinohomaloptera 7.1.5<br />
hohenackeri, Cobitis 4.20.1<br />
homalocephala, Vanmanenia 8.17.10<br />
Homalophra 7.6<br />
Homaloptera 7, 7.6, 8.17<br />
— abbreviata 7.9.1<br />
— amphisquamata 7.8.1<br />
— batek 7.2.1<br />
— bilineata 7.6.1<br />
— caldwelli 8.17.5<br />
— chekianensis 8.17.14<br />
— confuzona 7.6.2<br />
— fasciata 7.7.6
Homaloptera fasciolata 10.34.58<br />
— fimbriata 7.10, 7.10.2<br />
— formosana 7.5, 7.5.4<br />
— formosanum 8.4.6<br />
— gymnogaster 7.8.2<br />
— heterolepis 7.8.3<br />
— hingi 10.34.67<br />
— indochinensis 7.7.4<br />
— javanica 7.2.10<br />
— johorensis 7.12, 7.12.1<br />
— kwangsiensis 7.1, 7.1.5<br />
— leonardi 7.2.2<br />
— lepidogaster 7.8.2<br />
— lineata 7.7.4<br />
— manipurensis 7.7.3<br />
— maxinae 7.7.4<br />
— menoni 7.6.3<br />
— microstoma 8.12.1<br />
— modiglianii 7.8.4<br />
— monoloba 8.9.2<br />
— montana 7.6.4<br />
— multiloba 8.17.1<br />
— nebulosa 7.7.2<br />
— nigra 7.2.10<br />
— ocellata 7.6, 7.6.5, 7.7.6<br />
— ogilviei 7.6.6<br />
— ophiolepis 7.2.3<br />
— orthogoniata 7.6.7<br />
— parclitella 7.6.8<br />
— pillaii 7.6.9<br />
— polylepis 7.6.5<br />
— rotundicauda 10.24, 10.24.9<br />
— salusur 7.6.5<br />
— santhamparaiensis 7.6.10<br />
— septemmaculata 7.2.4<br />
— sexmaculata 7.2.4<br />
— silasi 7.6.11<br />
— smithi 7.7.4<br />
— stephensoni 7.2.5<br />
— taiwanica 7.5.4<br />
— tatereganii 7.2, 7.2.6<br />
— tetraloba 8.17.3<br />
— thamicola 7.4, 7.4.1<br />
— tweediei 7.7.5<br />
— ulmeri 7.8.6<br />
— valenciennesi 7.7.6<br />
— vanderbilti 7.8.6<br />
— ventrosquamata 8.17.4<br />
— vulgaris 7.2.7<br />
— wassinkii 7.7, 7.7.6<br />
— weberi 7.2.5<br />
— whiteheadi 8.14, 8.14.4<br />
— yuwonoi 7.2.9<br />
— zollingeri 7.2.10<br />
Homalopteraeformes 7<br />
Homalopteridae 7<br />
Homalopteroides 7.7<br />
— modestus 7.7.1<br />
— nebulosus 7.7.2<br />
— rupicola 7.7.3<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Homalopteroides smithi 7.7.4<br />
— tweediei 7.7.5<br />
— wassinkii 7.7.6<br />
Homalopterula 7.8<br />
— amphisquamata 7.8.1<br />
— gymnogaster 7.8.2<br />
— heterolepis 7.8.3<br />
— modiglianii 7.8.4<br />
— ripleyi 7.8, 7.8.5<br />
— vanderbilti 7.8.6<br />
Homalosoma 8.17<br />
— stenosoma 8.17, 8.17.14<br />
Homatula 10.11, 10.26<br />
— acuticephala 10.11.1<br />
— anguillioides 10.11.2<br />
— berezowskii 10.11.3<br />
— erhaiensis 10.11.4<br />
— laxiclathra 10.11.5<br />
— nanpanjiangensis 10.11.7<br />
— oligolepis 10.11.8<br />
— oxygnathus 10.11.7<br />
— potanini 10.11.9<br />
— pycnolepis 10.11.10<br />
— variegata 10.11.11<br />
— wujiangensis 10.11.12<br />
hopeiensis, Leptobotia 2.5.6<br />
horae, Botia 2.8.6<br />
horai, Nemachilus 10.34.69<br />
horai, Schistura 10.34.42, 10.34.69,<br />
10.34.97, 10.34.163<br />
hsiashiensis, Sinogastromyzon 7.13.5<br />
hsutschouensis, Nemacheilus 10.42.41<br />
hsutschouensis, Triplophysa 10.42.41<br />
huangguoshuensis, Beaufortia 8.2.5<br />
huanglongensis, Triplophysa 10.42.42<br />
huanjiangensis, Triplophysa 10.42.43<br />
huapingensis, Nemacheilus 10.34.70<br />
huapingensis, Schistura 10.34.70<br />
huapingensis, Triplophysa 10.42.44<br />
hugowolfeldi, Iksookimia 4.6.1<br />
humilis, Hypergastromyzon 8.7, 8.7.2<br />
humilis, Nemachilus 10.34.58, 10.42.37<br />
huongensis, Schistura 10.34.71<br />
hutchinsoni, Nemachilus 10.14.7<br />
hutjertjuensis, Nemacheilus 10.42, 10.42.45<br />
hutjertjuensis, Triplophysa 10.42.45<br />
hyalinus, Heminoemacheilus 10.10.1<br />
Hymenophysa 2.2, 2.7<br />
— macclellandi 2.7.4<br />
hymenophysa, Cobitis 2.7.4<br />
hymenophysa, Syncrossus 2.7.4<br />
Hymenphysa 2.2, 2.7<br />
hypercorpus, Sinogastromyzon 7.13.6<br />
Hypergastromyzon 8.7<br />
— eubranchus 8.7.1<br />
— humilis 8.7, 8.7.2<br />
Hypogastromyzon 8.7<br />
I<br />
Iberocobitis 4.5<br />
184<br />
Iksookimia 4.6, 4.15<br />
Iksookimia hugowolfeldi 4.6.1<br />
— koreensis 4.6.2<br />
— longicorpus 4.6.3<br />
— pacifica 4.6.4<br />
— pumila 4.6.5<br />
— yongdokensis 4.6.6<br />
Ilamnemacheilus 10.12<br />
— longipinnis 10.12, 10.12.1<br />
ilan, Dzihunia 10.7.2<br />
ilan, Nemachilus 10.7.2<br />
illyrica, Cobitis 4.5.22<br />
imitator, Schistura 10.34.72<br />
implicata, Schistura 10.34.73<br />
incerta, Barbatula 10.34.74<br />
incerta, Schistura 10.34.64, 10.34.74<br />
incipiens, Nemachilus 10.40.3<br />
incipiens, Tarimichthys 10.40.3<br />
incognito, Pangio 4.16.15<br />
indochinensis, Homaloptera 7.7.4<br />
Indoreonectes 10.13<br />
— evezardi 10.13.1<br />
— keralensis 10.13.2<br />
Indotriplophysa 10.14, 10.42<br />
— choprai 10.14.1<br />
— crassicauda 10.14.2<br />
— eugeniae 10.14.3<br />
— leptosoma 10.14.4<br />
— tenuicauda 10.14.5<br />
— tenuis 10.14.6<br />
— yasinensis 10.14.7<br />
Infundibulatus 10.23<br />
ingeri, Gastromyzon 8.5.16<br />
inglisi, Nemachilus 10.34.155<br />
insidiosa, Katibasia 8.8, 8.8.1<br />
insignis, Cobitis 10.25.20<br />
insignis, Oxynoemacheilus 10.25.4,<br />
10.25.20<br />
intermedia, Beaufortia 8.2.6<br />
intermedia, Diplophysa 10.42.46<br />
intermedia, Leptobotia 2.5.6<br />
intermedia, Praeformosania 8.17.12<br />
intermedia, Triplophysa 10.42.46<br />
intermedius, Sinogastromyzon 7.13.20<br />
introrsus, Gastromyzon 8.5.17<br />
iranica, Paracobitis 10.26.3<br />
irregularis, Schistura 10.34.75<br />
irrorata, Lepidocephalichthys 4.10.11<br />
Iskandaria 10.14<br />
— kuschakewitschi 10.15.1<br />
— pardalis 10.15.2<br />
isostigma, Schistura 10.34.76<br />
ispartensis, Seminemacheilus 10.25.27,<br />
10.36.1<br />
israeliticus, Orthrias 10.25.20<br />
J<br />
jadovaensis, Cobitis 4.5.23<br />
jaklesii, Cobitis 10.21.10<br />
jaklesii, Nemacheilus 10.21.10
japonica, Cobitis 4.5.8<br />
jarutanini, Schistura 10.34.77<br />
javanica, Homaloptera 7.2.10<br />
javanicus, Acanthophthalmus 4.16.24<br />
javedi, Botia 2.2.2<br />
Jerdonia 4.10<br />
jianchuanensis, Triplophysa 10.42.47<br />
jiarongensis, Triplophysa 10.42.48<br />
jimbeyi, Nemacheilus 10.42.101<br />
Jinshaia 7, 7.9<br />
— abbreviata 7.9.1<br />
— niulanjiangensis 7.9.2<br />
— sinensis 7.5.16, 7.9.3<br />
jinxiensis, Yunnanilus 10.46.10<br />
jiulongjiangensis, Pseudogastromyzon<br />
8.15.4<br />
johorensis, Homaloptera 7.12, 7.12.1<br />
johorensis, Neohomaloptera 7.12.1<br />
jonesi, Travancoria 7.14, 7.14.2<br />
jonklaasi, Lepidocephalichthys 4.10.12<br />
jordanicus, Noemacheilus 10.25.20<br />
K<br />
kaffa, Afronemacheilus 10.3<br />
kafirnigani, Nemachilus 10.42.49<br />
kafirnigani, Triplophysa 10.42.49<br />
kaimurensis, Nemacheilus 10.21.11<br />
kangjupkhulensis, Nemachilus 10.34.78<br />
kangjupkhulensis, Schistura 10.34.78,<br />
10.34.113<br />
kangjupkhulensis, Schistura<br />
kangrae, Nemachilus 10.34.79<br />
kangrae, Schistura 10.34.79<br />
kapuasensis, Nemacheilus 10.21.12<br />
kashmirensis, Nemachilus 10.14.7<br />
katibasensis, Gastromyzon 8.5.18<br />
Katibasia 8.8<br />
— insidiosa 8.8, 8.8.1<br />
katik, Kottelatlimia 4.9.2, 4.10.15<br />
katik, Lepidocephalichthys 4.9, 4.9.2<br />
kaynaki, Oxynoemacheilus 10.25.21<br />
kaysonei, Schistura 10.34.80<br />
kaznakovi, Gyrinocheilus 1.1.1<br />
kaznakowi, Labiatophysa 10.16.3<br />
kaznakowi, Triplophysa 10.16.3<br />
kellei, Cobitis 4.5.24<br />
kempi, Aborichthys 10.1, 10.1.3<br />
kengtungensis, Nemacheilus 10.34.81<br />
kengtungensis, Schistura 10.34.81<br />
keralensis, Indoreonectes 10.13.2<br />
keralensis, Oreonectes 10.13, 10.13.2<br />
kermanshahensis, Noemacheilus 10.25.22<br />
kermanshahensis, Oxynoemacheilus<br />
10.25.22<br />
kessleri, Nemachilus 10.28.4<br />
kessleri, Paraschistura 10.28.4<br />
kessleri, Schistura 10.28.6<br />
keyvani, Cobitis 4.5.25<br />
khamtanhi, Schistura 10.34.82<br />
khonensis, Hemimyzon 7.5.5<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
khugae, Schistura 10.34.83<br />
kiabii, Oxynoemacheilus 10.25.23<br />
kiangensis, Parabotia 2.5.8<br />
kiangsiensis, Parabotia 2.5.8<br />
Kichulchoia 4.7<br />
— brevifasciata 4.7.1<br />
— multifasciata 4.7.2<br />
kimluani, Parabotia 2.5.5<br />
kirinensis, Barbatula 10.4.8<br />
kloetzliae, Schistura 10.34.84<br />
kodaguensis, Schistura 10.34.85<br />
kohatensis, Schistura 10.34.86<br />
kohchangensis, Nemacheilus 10.34.87<br />
kohchangensis, Schistura 10.34.87<br />
kokshaalensis, Triplophysa 10.14<br />
koladynensis, Schistura 10.34.88<br />
kolhapurensis, Botia 2.2.8<br />
kongphengi, Schistura 10.34.89<br />
kontumensis, Schistura 10.34.90<br />
koreensis, Cobitis 4.6, 4.6.2<br />
koreensis, Iksookimia 4.6.2<br />
Koreocobitis 4.8<br />
— naktongensis 4.8.1<br />
— rotundicaudata 4.8.2<br />
kosswigi, Orthrias 10.25.24<br />
kosswigi, Oxynoemacheilus 10.25.24<br />
kosswigi, Turcinoemacheilus 10.45, 10.45.2<br />
kottelati, Neogastromyzon 8.10.4<br />
Kottelatlimia 4.9<br />
— hipporhynchos 4.9.1<br />
— katik 4.9.2, 4.10.15<br />
— pristes 4.9.3<br />
kranos, Lepidocephalichthys 4.10.13<br />
kubanica, Sabanejewia 4.20.7<br />
kubotai, Botia 2.2.5<br />
kudoii, Leptobotia 2.5.6<br />
kuhlii, Cobitis 4.16.16<br />
kuhlii, Pangio 4.16.16<br />
kuiperi, Nemachilus 10.21.28<br />
kullmanni, Triplophysa 10.42.50<br />
kungessana, Diplophysa 10.42, 10.42.51<br />
kungessana, Triplophysa 10.42.51<br />
kurui, Cobitis 4.5.62<br />
kuschakewitschi, Iskandaria 10.15.1<br />
kuschakewitschi, Nemachilus 10.15, 10.15.1<br />
kwangsiensis, Balitora 7.1.5<br />
kwangsiensis, Botia 2.5.6<br />
kwangsiensis, Homaloptera 7.1, 7.1.5<br />
kweichowensis, Beaufortia 8.2.7<br />
kweichowensis, Gastromyzon 8.2.7<br />
L<br />
labeosus, Nemacheilus 10.23<br />
labeosus, Neonoemacheilus 10.23.2<br />
labeosus, Noemacheilus 10.23.2<br />
labiata, Diplophysa 10.16, 10.16.4<br />
labiata, Labiatophysa 10.16.4<br />
Labiatophysa 10.16, 10.42<br />
— herzensteini 10.16.2<br />
— kaznakowi 10.16.3<br />
185<br />
Labiatophysa labiata 10.16.4<br />
— microphthalma 10.16.5, 10.42.67<br />
— nasalis 10.16.1<br />
Labigastromyzon 8.15<br />
lachnostoma, Acanthopsis 4.5.1<br />
lachnostoma, Cobitis 4.5.1<br />
lactogeneus, Nemacheilus 10.21.14<br />
lacusnigri, Nemachilus 10.42.52<br />
lacusnigri, Triplophysa 10.42.52<br />
lacustre, Crossostoma 8.4.6<br />
lacustris, Formosania 8.4.6<br />
lacustris, Triplophysa 10.42.53<br />
ladacensis, Nemachilus 10.14.5<br />
lancangjiangensis, Balitora 7.1.6<br />
lancangjiangensis, Sinohomaloptera 7.1.6<br />
laoensis, Cobitis 4.5.26<br />
laoensis, Misgurnus 4.5.26<br />
larvata, Cobitis 4.20.8<br />
larvata, Sabanejewia 4.20.8<br />
lateralis, Misgurnus 4.10.9<br />
laterimaculata, Cobitis 4.15.2<br />
laterimaculata, Niwaella 4.15.2<br />
laterimaculata, Schistura 10.34.118<br />
laterimaculata, Triplophysa 10.42.54<br />
laterivittatus, Noemacheilus 10.34.33<br />
laticauda, Balitora 7.1.7<br />
laticeps, Pseudogastromyzon 8.15.5<br />
laticeps, Triplophysa 10.42.55<br />
latidens, Schistura 10.34.91<br />
latifasciata, Schistura 10.34.92<br />
latifasciatus, Noemacheilus 10.34.92<br />
laxiclathra, Homatula 10.11.5<br />
lebedevi, Cobitis 4.5.27<br />
lecontei, Botia 2.8.3<br />
lecontei, Yasuhikotakia 2.8.3<br />
Lefua 10, 10.17<br />
— andrewsi 10.17.1<br />
— costata 10.17.1<br />
— echigonia 10.17.2<br />
— h<strong>of</strong>fmanni 10.17.3<br />
— nikkonis 10.17.4<br />
— pleskei 10.17.5<br />
— sayu 10.17.6<br />
Lefuini 10<br />
lendlii, Nemacheilus 10.36, 10.36.2<br />
lendlii, Seminemacheilus 10.36.2<br />
lenkoranensis, Nemacheilus 10.25.25<br />
lenkoranensis, Oxynoemacheilus 10.25.25<br />
leonardi, Balitoropsis 7.2.2<br />
leonardi, Homaloptera 7.2.2<br />
leontinae, Nemachilus 10.25.26<br />
leontinae, Oxynoemacheilus 10.25.26<br />
leopardus, Cobitis 10.25.31<br />
leopardus, Misgurnus 4.13.1<br />
lepidocaulis, Paraschistura 10.28.5<br />
lepidocaulis, Schistura 10.28.5<br />
Lepidocephalichthys 4.10<br />
— alkaia 4.10.1<br />
— annandalei 4.10.2<br />
— arunachalensis 4.10.3
Lepidocephalichthys barbatuloides<br />
4.10.4<br />
— berdmorei 4.10.5<br />
— coromandelensis 4.10.6<br />
— furcatus 4.10.7<br />
— goalparensis 4.10.8<br />
— guntea 4.10.9<br />
— hasselti 4.10.10<br />
— irrorata 4.10.11<br />
— jonklaasi 4.10.12<br />
— katik 4.9, 4.9.2<br />
— kranos 4.10.13<br />
— lorentzi 4.10.14<br />
— manipurensis 4.10.15<br />
— menoni 4.10.2<br />
— micropogon 4.10.15<br />
— nepalensis 4.10.9<br />
— nudus 4.10.10<br />
— pallens 4.11.1<br />
— pristes 4.9.3<br />
— thermalis 4.10.16<br />
— tomaculum 4.10.17<br />
— zeppelini 4.10.18<br />
Lepidocephalus 4.11<br />
— birmanicus 4.10.5<br />
— cataractus 4.10.5<br />
— caud<strong>of</strong>urcatus 4.10.8<br />
— dibruensis 4.10.9<br />
— macrochir 4.11.1<br />
— spectrum 4.11.2<br />
— taeniatus 4.10.10<br />
— weberi 10.21<br />
lepidogaster, Gastromyzon 8.5.19<br />
lepidogaster, Homaloptera 7.8.2<br />
Lepidoglanidae 8<br />
Lepidoglanis 8, 8.5<br />
— monticola 8.5, 8.5.21<br />
Leptobotia 1, 2.4<br />
— banarescui 2.5.1<br />
— elongata 2.4.1, 2.4.7<br />
— flavolineata 2.4.2<br />
— guilinensis 2.4.3<br />
— hansuiensis 2.4.13<br />
— hengyangensis 2.4.4<br />
— hopeiensis 2.5.6<br />
— intermedia 2.5.6<br />
— kudoii 2.5.6<br />
— mantschurica 2.5.11<br />
— orientalis 2.4.6<br />
— microphthalma 2.4.5<br />
— pellegrini 2.4.7<br />
— posterodorsalis 2.4.8<br />
— punctata 2.4.9<br />
— purpurea 2.4.11<br />
— rubrilabris 2.4.10<br />
— taeniops 2.4.11<br />
— tchangi 2.4.12<br />
— tientainensis 2.4.13<br />
Leptobotiini 1<br />
leptocephala, Ussuria 4.13, 4.13.4<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
leptosoma, Indotriplophysa 10.14.4<br />
leptosoma, Nemachilus 10.14.4<br />
Lepturichthys 7, 7.10<br />
— dolichopterus 7.10.1<br />
— fimbriatus 7.10.2<br />
— guentheri 7.10.2<br />
— nicholsi 7.10.2<br />
leukensis, Schistura 10.34.93<br />
levantina, Cobitis 4.5.28<br />
leveretti, Beaufortia 8.2.8<br />
leveretti, Gastromyzon 8.2, 8.2.8<br />
lhasae, Nemachilus 10.42.98<br />
lianjiangensis, Pseudogastromyzon 8.15.6<br />
liboensis, Nemacheilus 10.42.59<br />
lidi, Pangio 4.16.17<br />
lijiangensis, Parabotia 2.5.9<br />
lindbergi, Noemacheilus 10.28.6<br />
lindbergi, Paraschistura 10.28.6<br />
linea, Acanthopsis 4.5.29<br />
linea, Cobitis 4.5.29<br />
lineata, Homaloptera 7.7.4<br />
lineata, Vanmanenia 8.17.11<br />
lineatus, Oreias 10.34.105<br />
lineolata, Balitora 8.16, 8.16.6<br />
lineolata, Sewellia 8.16.6<br />
lingyunensis, Schistura 10.42.56<br />
lingyunensis, Triplophysa 10.42.56<br />
Liniparhomaloptera 8.9<br />
— disparis 8.9.1<br />
— monoloba 8.9.2<br />
— obtusirostris 8.9.3<br />
— qiongzhongensis 8.9.4<br />
liui, Beaufortia 8.2.9<br />
lividus, Nemachilus 4.13.1<br />
lixianensis, Triplophysa 10.42.57<br />
lixianjiangensis, Sinogastromyzon 7.13.7<br />
lohachata, Botia 2.2.6<br />
longa, Schistura 10.34.94<br />
longchiensis, Triplophysa 10.42.1<br />
longianalis, Nemachilus 10.42.99<br />
longianguis, Triplophysa 10.42.58<br />
longibarba, Niwaella 4.15.3<br />
longibarbata, Balitora 7.1.9<br />
longibarbata, Paracobitis 10.42.59<br />
longibarbata, Triplophysa 10.42.59<br />
longibarbatus, Sinohomaloptera 7.1.9<br />
longibarbatus, Yunnanilus 10.46.11<br />
longibarbus, Nemachilus 10.9.4<br />
longibulla, Yunnanilus 10.46.12<br />
longicauda, Cobitis 10.26, 10.26.4<br />
longicauda, Metahomaloptera 7.11.2<br />
longicauda, Paracobitis 10.26.4<br />
longicorpus, Cobitis 4.6.3<br />
longicorpus, Iksookimia 4.6.3<br />
longidorsalis, Botia 2.84<br />
longidorsalis, Eonemachilus 10.8.1<br />
longidorsalis, Homatula 10.11.6<br />
longidorsalis, Paracobitis 10.11.6<br />
longidorsalis, Yasuhikotakia 2.8.4<br />
longidorsalis, Yunnanilus 10.8.1<br />
186<br />
longimanus, Cobitis 4.5<br />
longimanus, Pangio 4.16.18<br />
longipectoralis, Nemacheilus 10.21.13<br />
longipectoralis, Protonemacheilus 10.31,<br />
10.31.1<br />
longipectoralis, Triplophysa 10.42.60<br />
longipinnis, Acanthocobitis 10.2, 10.2.1,<br />
10.2.5, 10.21.14<br />
longipinnis, Acanthophthalmus 4.10.5<br />
longipinnis, Ilamnemacheilus 10.12, 10.12.1<br />
longipinnis, Nemacheilus 10.21.14<br />
Longischistura 10.34<br />
longistriatus, Nemacheilus 10.21.15<br />
longitaeniatus, Cobitis 4.5.26<br />
longiventralis, Botia 2.6.1<br />
longiventralis, Sinibotia 2.6.1<br />
longus, Nemachilus 10.34.94<br />
loos, Beaufortia 8.2.10<br />
loos, Gastromyzon 8.2.10<br />
lorentzi, Acanthophthalmus 4.10.14<br />
lorentzi, Lepidocephalichthys 4.10.14<br />
lucasbahi, Botia 2.7.5<br />
lucasbahi, Syncrossus 2.7.5<br />
lucidorsum, Pteronemacheilus 10.32,<br />
10.32.1<br />
lumbriciformis, Pangio 4.16.19<br />
lunanensis, Noemacheilus 10<br />
ludongensis, Balitora 7.1.8<br />
lungkowensis, Paraprotomyzon 8.11.2<br />
luochengensis, Oreonectes 10.24.8<br />
lutheri, Cobitis 4.5.30<br />
M<br />
maassi, Vaillantella 3.1.3<br />
macclellandi, Hymenophysa 2.7.4<br />
macedonicus, Orthrias 10.25.8<br />
machensis, Noemacheilus 10.34.95<br />
machensis, Schistura 10.34.95<br />
mackenziei, Nemachilus 10.2.2<br />
macmahoni, Nemacheilus 10.26, 10.26.6<br />
macracanthus, Chromobotia 2.3.1<br />
macracanthus, Cobitis 2.3, 2.3.1<br />
macrocephala, Triplophysa 10.42.61<br />
macrocephalus, Schistura 10.34.83,<br />
10.34.96<br />
macrochir, Cobitis 4.11, 4.11.1<br />
macrochir, Lepidocephalus 4.11.1<br />
macrogaster, Yunnanilus 10.46.13<br />
macroiscalus, Yunnanilus 10.46<br />
macrolepis, Oreonectes 10.24.3<br />
macrolepis, Schistura 10.34.13, 10.34.97<br />
macrolepis, Yunnanilus 10.46.20<br />
macrolineata, Botia 2.2.3<br />
macromaculata, Triplophysa 10.42.62<br />
macrophthalma, Triplophysa 10.42.63<br />
macroptera, Hemimyzon 7.5.6<br />
macropterus, Hemimyzon 7.5.6<br />
macropterus, Hedinichthys 10.9.2<br />
macropterus, Nemachilus 10.9.2<br />
macrorhynchos, Cobitis 4.2.1
macrositanus, Yunnanilus 10.46.14<br />
macrostigma, Cobitis 4.5.31<br />
macrostoma, Sinogastromyzon 7.13.8<br />
macrotaenia, Nemacheilus 10.34.98<br />
macrotaenia, Schistura 10.34.98<br />
maculata, Balitora 7.1.2<br />
maculata, Cobitis 4.13.1<br />
maculata, Schistura 2.2.1<br />
maculatum, Pseudogastromyzon 8.15.8<br />
maculatus, Misgurnus 4.13.1<br />
maculatus, Platacanthus 4.10, 4.10.16<br />
maculiceps, Nemacheilus 10.34.99<br />
maculiceps, Schistura 10.34.99<br />
maculosa, Botia 2.5.10<br />
maculosus, Parabotia 2.5.10<br />
Madrasia 4.10<br />
maepaiensis, Schistura 10.34.100<br />
magnifluvis, Schistura 10.34.101<br />
mahnerti, Schistura 10.34.102<br />
malaisei, Schistura 10.34.103<br />
malapterura, Cobitis 10.26, 10.26.5<br />
malapterura, Paracobitis 10.26.5<br />
malayana, Pangio 4.16.20<br />
malayanus, Acanthophthalmus 4.16.20<br />
malloryi, Triplophysa 10.42<br />
mandalayensis, Acanthocobitis 10.2.3<br />
mandalayensis, Nemacheilus 10.2.3<br />
manipurensis, Homaloptera 7.7.3<br />
manipurensis, Lepidocephalichthys 4.10.15<br />
manipurensis, Nemachilus 10.34.104<br />
manipurensis, Schistura 10.34.104<br />
mantschurica, Leptobotia 2.5.11<br />
mantschuricus, Parabotia 2.5.11<br />
maolanensis, Paracobitis 10.42.59<br />
maon, Sinogastromyzon 7.13.9<br />
marang, Nemacheilus 10.21.16<br />
mariae, Acanthopthalmus 4.16.21<br />
mariarum, Pangio 4.16.21<br />
markakulensis, Barbatula 10.4.4, 10.4.7,<br />
10.4.15<br />
markakulensis, Nemacheilus 10.4.7<br />
markehenensis, Nemachilus 10.42.64<br />
markehenensis, Triplophysa 10.42.64<br />
marmorata, Cobitis 10.42.65<br />
marmorata, Sewellia 8.16.7<br />
marmorata, Triplophysa 10.42.65<br />
maroccana, Cobitis 4.5.32<br />
masyae, Nemacheilus 10.21, 10.21.17<br />
matsubarae, Cobitis 4.5.33<br />
maxinae, Homaloptera 7.7.4<br />
maya, Cobitis 4.10.9<br />
maydelli, Neoeucirrhichthys 4.14, 4.14.1<br />
media, Sewellia 8.16.8<br />
mediterraneus, Barbatula 10.25.27, 10.36.1<br />
mediterraneus, Oxynoemacheilus 10.25.27<br />
megalepis, Gastromyzon 8.5.20<br />
megalomycter, Aperioptus 5.1.1<br />
megalomycter, Ellopostoma 5.1.1<br />
megalopseos, Hemimyzon 7.5.7<br />
megaspila, Cobitis 4.5.15<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
megastoma, Sectoria 10.35.2<br />
meihuashanensis, Pseudogastromyzon<br />
8.15.7<br />
melanoleuca, Cobitis 4.5.19, 4.5.34, 4.5.48<br />
melanosoma, Balitora 7.1.3<br />
melarancia, Schistura 10.34.105<br />
menanensis, Noemacheilus 10.34.106<br />
menanensis, Schistura 10.34.106<br />
mengdingensis, Neonoemacheilus 10.23.3<br />
menoni, Homaloptera 7.6.3<br />
menoni, Lepidocephalichthys 4.10.2<br />
menoni, Mesonoemacheilus 10.18.3<br />
menoni, Noemacheilus 10.18.3<br />
merga, Cobitis 10.25.28<br />
merga, Oxynoemacheilus 10.25.28<br />
meridionalis, Balitora 7.1.10<br />
meridionalis, Cobitis 4.5.35<br />
meridionalis, Nemachilus 10.32.2<br />
meridionalis, Pteronemacheilus 10.32.2<br />
Mesomisgurnus 4.13<br />
Mesonoemacheilus 10.18<br />
— guentheri 10.18.1<br />
— herrei 10.18.2<br />
— menoni 10.18.3<br />
— pambarensis 10.18.4<br />
— petrubanarescui 10.18.5<br />
— pulchellus 10.18.6<br />
— remadevii 10.18.7<br />
— reticul<strong>of</strong>asciatus 10.34.151<br />
— triangularis 10.18.8<br />
Metahomaloptera 7, 7.11<br />
— hangshuiensis 7.11.1<br />
— longicauda 7.11.2<br />
— omeiensis 7.11, 7.11.3<br />
Metaschistura 10.19<br />
— cristata 10.19.1<br />
microcephala, Cobitis 4.5.36<br />
Microcobitis 4.12<br />
— misgurnoides 4.12.1<br />
microlabra, Paraschistura 10.28.7<br />
microlabra, Schistura 10.28.7<br />
microlepis, Vanmanenia 8.17.7<br />
Micronemacheilus 10.20<br />
— bacmeensis 10.41.2<br />
— cruciatus 10.20.1<br />
— zispi 10.41.2<br />
microphthalma, Diplophysa 10.16.5,<br />
10.42.67<br />
microphthalma, Labiatophysa 10.16.5,<br />
10.42.67<br />
microphthalma, Leptobotia 2.4.5<br />
microphthalma, Triplophysa 10.42.67<br />
microphthalmus, Oreonectes 10.24.4<br />
microphthalmus, Schistura 10.42.67<br />
microphysus, Nemacheilus 10.42.66<br />
microphysus, Triplophysa 10.42.66<br />
micropogon, Acanthopis 4.10.15<br />
micropogon, Lepidocephalichthys 4.10.15<br />
microps, Cobitis 10.42.68<br />
microps, Triplophysa 10.42.11, 10.42.68<br />
187<br />
micropus, Cobitis 4.13.1<br />
microstoma, Homaloptera 8.12.1<br />
microstoma, Parhomaloptera 8.12.1<br />
minuta, Schistura 10.34.107<br />
minutum, Sinogastromyzon 7.13.10<br />
minutus, Hedinichthys 10.9.3<br />
minutus, Nemachilus 10.9.3<br />
minutus, Sinogastromyzon 7.13.10<br />
minxianensis, Nemachilus 10.42.13<br />
Misgurninae 4<br />
misgurnoides, Cobitis 4.12, 4.12.1<br />
misgurnoides, Microcobitis 4.12.1<br />
Misgurnus 4, 4.13<br />
— anguillicaudatus 4.13.1, 4.18.1<br />
— arenae 4.5.2<br />
— buphoensis 4.13.**<br />
— cestoideus 4.13.4<br />
— crossochilus 4.13.1<br />
— elongatus 4.13.1<br />
— erikssoni 4.13.4<br />
— formosanus 4.13.1<br />
— fossilis 4.13.3<br />
— fukien 4.13.1<br />
— grangeri 4.13.1<br />
— hainan 4.13.1<br />
— heungchow 4.13.1<br />
— laoensis 4.5.26<br />
— lateralis 4.10.9<br />
— leopardus 4.13.1<br />
— maculatus 4.13.1<br />
— mizolepis 4.18.1<br />
— mohoity 4.13.4<br />
— multimaculatus 4.13.5<br />
— nikolskyi 4.13.6<br />
— oligolepos 4.18.1<br />
— olygoskolopos 4.18.1<br />
— punctatus 4.13.1<br />
— spilurus 4.13.4<br />
— tonkinensis 4.13.7<br />
— tungting 4.13.1<br />
— unicolor 4.13.1<br />
— yunnan 4.13.1<br />
mizolepis, Misgurnus 4.18.1<br />
mobbsi, Schistura 10.34.108<br />
modesta, Botia 2.8, 2.8.5<br />
modesta, Helgia 7.7.1<br />
modesta, Yasuhikotakia 2.8.5<br />
modestus, Homalopteroides 7.7.1<br />
Modigliania 10.21<br />
— papillosa 10.21, 10.21.23<br />
modiglianii, Homaloptera 7.8.4<br />
modiglianii, Homalopterula 7.8.4<br />
moeiensis, Schistura 10.34.109<br />
mohoity, Cobitis 4.13.4<br />
mohoity, Misgurnus 4.13.4<br />
molobrion, Acanthopsoides 4.1.5<br />
monchadskii, Gyrinocheilus 1.1.1<br />
monilis, Nemacheilus 10.21.18<br />
monocera, Cobitis 10.2.2<br />
mon<strong>of</strong>asciodorsalata, Vanmanenia 8.17.13
monoloba, Homaloptera 8.9.2<br />
monoloba, Liniparhomaloptera 8.9.2<br />
monolobata, Parasewellia 8.16.9<br />
monolobata, Sewellia 8.16.9<br />
montana, Cobitis 4.20.2<br />
montana, Homaloptera 7.6.4<br />
montana, Schistura 10, 10.34, 10.34.79,<br />
10.34.110<br />
montanus, Cobitis 4.20.2, 10<br />
monticola, Gastromyzon 8.5.21<br />
monticola, Lepidoglanis 8.5, 8.5.21<br />
mooreh, Acanthocobitis 10.2.4<br />
mooreh, Cobitis 10.2.4<br />
moquensis, Triplophysa 10.42.69<br />
morehensis, Neonoemacheilus 10.23.4<br />
morleti, Botia 2.8.6<br />
morleti, Yasuhikotakia 2.8.6<br />
mugah, Nemacheilus 10.34.193<br />
mukambbikaensis, Noemacheilus 10.34.111<br />
mukambbikaensis, Schistura 10.34.47,<br />
10.34.111<br />
multifasciata, Botia 2.5.6<br />
multifasciata, Kichulchoia 4.7.2<br />
multifasciata, Schistura 10.34.112,<br />
10.34.155<br />
multifasciatus, Cobitis 4.7.2<br />
multifasciatus, Nemachilus 10.34.112<br />
multifasciatus, Paraprotomyzon 8.11, 8.11.3<br />
multiloba, Homaloptera 8.17.1<br />
multiloba, Vanmanenia 8.17.1<br />
multimaculata, Cobitis 4.5.37<br />
multimaculatus, Misgurnus 4.13.5<br />
multiocellata, Beaufortia 8.2.16<br />
multiocellum, Sinogastromyzon 7.13.11<br />
multiradiata, Glaniopsis 8.6.4<br />
multistigmatus, Acantopsis 4.2.2<br />
muraeniformis, Acanthophthalmus 4.16.22<br />
muraeniformis, Pangio 4.16.22<br />
myersi, Acanthophthalmus 4.16.23<br />
myersi, Pangio 4.16.23<br />
myersi, Pseudogastromyzon 8.15.8<br />
myrmekia, Nemacheilus 10.34.48<br />
mysorensis, Balitora 7.1.11<br />
mysorensis, Cobitis 4.10.16<br />
mystax, Ellopostoma 5.1.2<br />
N<br />
Naemacheilus 10.21<br />
nagaensis, Noemacheilus 10.34.113<br />
nagaensis, Schistura 10.34.113<br />
nagodiensis, Schistura 10.34.114<br />
nahangensis, Vanmanenia 8.17.2<br />
naktongensis, Koreocobitis 4.8.1<br />
nalbanti, Noemacheilus 10.28.12<br />
namboensis, Schistura 10.34.115<br />
namiri, Nemacheilus 10.25.29<br />
namiri, Oxynoemacheilus 10.25.29<br />
namnaensis, Sinogastromyzon 7.13.12<br />
namnuaensis, Dienbienia 7.5., 7.5.11<br />
namromensis, Acanthopsoides 4.1.2<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
nandanensis, Triplophysa 10.42.70<br />
nandingensis, Noemacheilus 10.34.116<br />
nandingensis, Schistura 10.34.116<br />
nanensis, Hemimyzon 7.5.8<br />
Nanoschistura 10.25<br />
nanpanjiangensis, Homatula 10.11.7<br />
nanpanjiangensis, Oreias 10.42.71<br />
nanpanjiangensis, Paracobitis 10.11.7<br />
nanpanjiangensis, Sinogastromyzon 7.13.13<br />
nanpanjiangensis, Triplophysa 10.42.71<br />
nanpanjiangensis, Yunnanilus 10.46.15<br />
nantaiensis, Sinogastromyzon 7.13.14<br />
nantingensis, Balitora 7.1.12<br />
narentana, Cobitis 4.5.38<br />
narinosus, Draconectes 10.6.1<br />
nasalis, Diplophysa 10.16.1<br />
nasalis, Labiatophysa 10.16.1<br />
naseeri, Noemacheilus 10.28.8<br />
naseeri, Paraschistura 10.28.8<br />
nasifilis, Nemachilus 10.34.117<br />
nasifilis, Schistura 10.34.117<br />
nasobarbatula, Triplophysa 10.42.72<br />
Naunacheilus 10.21<br />
naziri, Noemacheilus 10.42.36<br />
Neacanthopsis 4.1<br />
— gracilentus 4.1, 4.1.2<br />
nebulosa, Botia 2.2.7<br />
nebulosa, Homaloptera 7.7.2<br />
nebulosus, Homalopteroides 7.7.2<br />
Nemacheilidae 10<br />
Nemacheilos 10.21<br />
Nemacheilus 10, 10.21<br />
— abyssinicus 10.3, 10.3.1<br />
— acuticephalus 10.34.3<br />
— afghana 10.42.65<br />
— afrenatus 10.25.16<br />
— ajmonis 10.14.7<br />
— akhtari 10.42.1<br />
— alasanicus 10.25.25<br />
— alepidotus 10.28.1<br />
— alticeps 10.33, 10.33.1<br />
— altipedunculatus 10.34.8<br />
— amudarjensis 10.7, 10.7.1<br />
— anambarensis 10.34.10<br />
— angeli 10.4.8, 10.42.6<br />
— anglicana 10.4.2<br />
— angorae 10.25.2<br />
— anguilla 10.21.1<br />
— arenicolus 10.21.2<br />
— arunachalensis 4.10.3, 10.34.184<br />
— assamensis 10.23.1<br />
— atriceps 10.35, 10.35.1<br />
— aureus 10.2.4<br />
— badamensis 10.15.1<br />
— baenzigeri 10.44, 10.44.1<br />
— balteatus 10.34.19<br />
— baluchiorum 10.34.20<br />
— bampurensis 10.28.2<br />
— banar 10.21.3<br />
— barapaniensis 10.30.3<br />
188<br />
Nemacheilus beavani 10.34.22<br />
— bellibarus 10.42.11<br />
— berezowskii 10.11.3<br />
— bergi 10.25.25<br />
— bergianus 10.25.6<br />
— bertini 10.42.86<br />
— bhimachari 10.34.24<br />
— binotatus 10.21.4<br />
— bipartitus 4.13, 4.13.4<br />
— blackiana 10.4.2<br />
— bleekeri 10.42.12<br />
— blythii 10<br />
— bombifrons 10.40<br />
— brachypterus 10.42.13<br />
— brahui 10.42.14<br />
— brandtii 10.25.7<br />
— brevibarbus 10.9.4, 10.42.15<br />
— brevicauda 10.42.16<br />
— breviceps<br />
— brevis 10.29, 10.29.2<br />
— brunneanus 10.30, 10.30.1<br />
— bucculentus 10.34.27<br />
— bureschi 10.25.8<br />
— callichromus 10.34.28<br />
— carassus 10.42.86<br />
— carletoni 10.34.31<br />
— caucasicus 10.4.2<br />
— chapaensis 10.34.34<br />
— cheni 2.4<br />
— chindwinicus 10.34.35<br />
— chondrostoma 10.42.18<br />
— choprai 10.14.1<br />
— choresmi 10.7.1<br />
— chryseus 10.34.47<br />
— chrysolaimos 10.21.5<br />
— cleopatra 10.21.6<br />
— cobdonensis 10.4.3<br />
— compressirostris 10.4.4<br />
— conipterus 10.42.19<br />
— conirostris 10.34.38<br />
— conjungens 10.16.4<br />
— corica 10.21.7<br />
— crassicauda 10.14.2<br />
— cristatus 10.19, 10.19.1<br />
— cruciatus 10.20, 10.20.1<br />
— cueljuensis 10.42.25<br />
— cyri 10.25.11<br />
— dattai 10.34.184<br />
— dayi 10.34.47<br />
— deignani 10.34.87<br />
— denisoni 10.34.47<br />
— desmotes 10.34.48<br />
— deterrai 10.42.34<br />
— devdevi 10.34.49<br />
— dixoni 10.17.1<br />
— djaggasteensis 10.42.22<br />
— doonensis 10.34.52<br />
— dorsaloides 10.42.106<br />
— dorsonotatus 10.42.25<br />
— drassensis 10.42.26
Nemacheilus dunckeri 10.21.25<br />
— elegantissimus 10.21.8<br />
— elongatus 10.1.1, 10.30.3, 10.42.77<br />
— erythrinna 10.4.2<br />
— euepipterus 3.1, 3.1.2<br />
— euphraticus 10.25.4<br />
— evezardi 10.13.1<br />
— farwelli 10.42.28<br />
— fasciatus 10.21.9<br />
— fedtschenkoae 10.42.107<br />
— flavus 10.25<br />
— fowlerianus 10.34.162<br />
— gangeticus 10.34.61<br />
— gejiuensis 10.42.32<br />
— ghazniensis 10.26.2<br />
— gibbusnazus 10.25.7<br />
— gracilis 10.42.34<br />
— grahami 10.42.35<br />
— griffithii 10.42.36<br />
— guentheri 10.18.1<br />
— haarlovi 10.28.6<br />
— hamwii 10.25.19<br />
— harnaiensis 10.34.65<br />
— hazaraensis 10.42.36<br />
— hexiensis 10.42.38<br />
— himachalensis 10.34.66<br />
— hispanica 10.4.11<br />
— horai 10.34.69<br />
— hsutschouensis 10.42.41<br />
— huapingensis 10.34.70<br />
— humilis 10.34.58, 10.42.37<br />
— hutchinsoni 10.14.7<br />
— hutjertjuensis 10.42, 10.42.45<br />
— ilan 10.7.2<br />
— incipiens 10.40.3<br />
— inglisi 10.34.155<br />
— jaklesii 10.21.10<br />
— jimbeyi 10.42.101<br />
— jordanicus 10.25.20<br />
— kafirnigani 10.42.49<br />
— kaimurensis 10.21.11<br />
— kangjupkhulensis 10.34.78<br />
— kangrae 10.34.79<br />
— kapuasensis 10.21.12<br />
— kashmirensis 10.14.7<br />
— kengtungensis 10.34.81<br />
— kermanshahensis 10.25.22<br />
— kessleri 10.28.4<br />
— kohchangensis 10.34.87<br />
— kuiperi 10.21.28<br />
— kuschakewitschi 10.15, 10.15.1<br />
— labeosus 10.23, 10.23.2<br />
— lactogeneus 10.21.14<br />
— lacusnigri 10.42.52<br />
— ladacensis 10.14.5<br />
— laterivittatus 10.34.33<br />
— latifasciatus 10.34.92<br />
— lendlii 10.36, 10.36.2<br />
— lenkoranensis 10.25.25<br />
— leontinae 10.25.26<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
Nemacheilus leptosoma 10.14.4<br />
— lhasae 10.42.98<br />
— liboensis 10.42.59<br />
— lindbergi 10.28.6<br />
— lividus 4.13.1<br />
— longianalis 10.42.99<br />
— longibarbus 10.9.4<br />
— longipectoralis 10.21.13<br />
— longipinnis 10.21.14<br />
— longistriatus 10.21.15<br />
— longus 10.34.94<br />
— lunanensis 10<br />
— machensis 10.34.95<br />
— mackenziei 10.2.2<br />
— macmahoni 10.26, 10.26.6<br />
— macropterus 10.9.2<br />
— macrotaenia 10.34.98<br />
— maculiceps 10.34.99<br />
— mandalayensis 10.2.3<br />
— manipurensis 10.34.104<br />
— marang 10.21.16<br />
— markakulensis 10.4.7<br />
— markehenensis 10.42.64<br />
— masyae 10.21, 10.21.17<br />
— menanensis 10.34.106<br />
— menoni 10.18.3<br />
— meridionalis 10.32.2<br />
— microphysus 10.42.66<br />
— minutus 10.9.3<br />
— minxianensis 10.42.13<br />
— monilis 10.21.18<br />
— mugah 10.34.193<br />
— mukambbikaensis 10.34.111<br />
— multifasciatus 10.34.112<br />
— myrmekia 10.34.48<br />
— nagaensis 10.34.113<br />
— nalbanti 10.28.12<br />
— namiri 10.25.29<br />
— nandingensis 10.34.116<br />
— naseeri 10.28.8<br />
— nasifilis 10.34.117<br />
— naziri 10.42.36<br />
— nicholsi 10.34.118<br />
— nigromaculatus 10.8, 10.8.3<br />
— nilgiriensis 10.34.119<br />
— nordkansuensis 10.9.2<br />
— notostigma 10.34.121<br />
— nudus 10.4.8, 10.42.6<br />
— obesus 10.39, 10.39.1<br />
— obscurus 10.34.18<br />
— oedipus 10.34.14<br />
— olivaceus 10.21.19<br />
— orientalis 10.42.77<br />
— ornatissimus 10.21.8<br />
— ornatus 10.21.20<br />
— oxiana 10.25<br />
— oxygnathus 10.11.7<br />
— pakistanicus 10.28.10<br />
— pallidus 10.21.21<br />
— paludani 10.28.12<br />
189<br />
Nemacheilus pambaensis 10.34.47<br />
— pambarensis 10.18.4<br />
— panguri 10.42.99<br />
— papillos 10.21.22<br />
— papillosus 10.21.23<br />
— pappenheimi 10.42.79<br />
— paradoxus 10.42.80<br />
— pardalis 10.15.2<br />
— paucifasciatus 10.34.129<br />
— paucimaculatus 10.21.24<br />
— pechiliensis 10.4.8<br />
— peguensis 10.23, 10.23.5<br />
— pellegrini 10.34.176<br />
— periyarensis 10.18.8<br />
— petrubanarescui 10.18.5<br />
— pfeifferae 10.21.25<br />
— phuketensis 10.2.9<br />
— plagiognathus 10.42.86<br />
— platiceps 10.21.26<br />
— pleurotaenia 10.46, 10.46.22<br />
— poculi 10.34.134<br />
— polytaenia 10.34.135<br />
— poonaensis 10.21.1<br />
— potanini 10.11, 10.11.9<br />
— prashadi 10.34.137<br />
— prashari 10.28.11<br />
— productus 10.14.4<br />
— pseudoscleropterus 10.42.84<br />
— pulchellus 10.18.6<br />
— pulcher 10.41.1<br />
— pulsiz 10.25.12<br />
— punjabensis 10.28.12<br />
— putaoensis 10.34.169<br />
— quignardi 10.4.11<br />
— rajasthanicus 10.34.147<br />
— raoe 10.30.5<br />
— rarus 10.34.148<br />
— reidi 10.34.149<br />
— rendahli 10.34.47<br />
— retropinnis 10.42.86<br />
— reuniens 10.42.106<br />
— rhadinaeus 10.26.6<br />
— rivulicola 10.30.6<br />
— robustus 10.42.87<br />
— rotundiventris 10.33.2<br />
— ruzskyi 10.42.101<br />
— salari 10.42.19<br />
— salmonides 10.29.1<br />
— saravacensis 10.21.27<br />
— sargadensis 10.28, 10.28.13<br />
— savona 10.34.47<br />
— schultzi 10.34.159<br />
— scleropterus 10.42.90<br />
— selangoricus 10.21.28<br />
— semiarmatus 10.34.160<br />
— serpentarius 7.6.1<br />
— sewerzowi 10.42.92<br />
— sexcauda 10.34.162<br />
— seyhanensis 10.25.37<br />
— shanensis 10.30.7
Nemacheilus shebbearei 10.34.165<br />
— shehensis 10.42.94<br />
— shiyangensis 10.42.96<br />
— shuangjiangensis 10.34.166<br />
— sibiricus 10.4.14, 10.4.15<br />
— sijuensis 10.34.168<br />
— sikkimensis 10<br />
— sikmaiensis 10.34.169<br />
— siluroides 10.42.97<br />
— singhi 10.34.171<br />
— sinuatus 10.2.4<br />
— smithi 10.26.7<br />
— spilotus 10.34.177<br />
— spiniferus 10.21.29<br />
— starostini 10.43, 10.43.1<br />
— stenurus 10.42.98<br />
— stigm<strong>of</strong>asciatus 10.21.30<br />
— striatus 10.34, 10.34.178<br />
— sturanyi 10.4.13<br />
— taeniata 10.41.2<br />
— taeniatus 10.41<br />
— tambaraparniensis 10.18.8<br />
— tanggulaensis 10.42.102<br />
— tangtianhensis 10.42.99<br />
— tarimensis 10.9.4<br />
— taurica 10.4.2<br />
— tebo 10.21.31<br />
— tenuis 10.14, 10.14.6<br />
— terekensis 10.25.28<br />
— thai 10.34.118<br />
— tibetanus 10.42.104<br />
— tomianus 10.4.14<br />
— tortonesei 10.25.20<br />
— transiens 10.42.101<br />
— translineatus 10.21.28<br />
— triangularis 10.18, 10.18.8<br />
— troglocataractus 10.21.32<br />
— tschaiyssuensis 10.25.4<br />
— tuberigum 10.21.33<br />
— turcmenicus 10.28.13<br />
— turcomanus 10.28.14<br />
— turfanensis 10.42.118<br />
— ulacholicus 10.42.106<br />
— urophthalmus 10.2.8<br />
— valerii 10.42.24<br />
— vardarensis 10.4.2<br />
— variegatus 10.11.11<br />
— vinciguerrae 10.34.188<br />
— waltoni 10.34.189<br />
— wuweiensis 10.42.110<br />
— xanthi 2.5.6<br />
— xiangxiensis 10.42.112<br />
— xingshanensis 10.42.114<br />
— yarkandensis 10.9, 10.9.4<br />
— yasinensis 10.14.7<br />
— yingjiangensis 10.34.192<br />
— zaidamensis 10.33.3<br />
— zaisanicus 10.42.101<br />
— zamegacephalus 10.33.4<br />
— zeravschani 10.25<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Nemachilichthys 10.22<br />
Nemachilichthys ruppelli 10.22.1<br />
— shimogensis 10.22.1<br />
Nemachilus 10.21 (see species under<br />
Nemacheilus)<br />
Nematocheilos 10.21<br />
Neoeucirrhichthys 4.14<br />
— maydelli 4.14, 4.14.1<br />
Neogastromyzon 8.10<br />
— brunei 8.10.1<br />
— chini 8.10.2<br />
— crassiobex 8.10.3<br />
— kottelati 8.10.4<br />
— nieuwenhuisii 8.10, 8.10.5<br />
— pauciradiatus 8.10.6<br />
Neohomaloptera 7.12<br />
— johorensis 7.12.1<br />
Neonoemacheilus 10.23<br />
— assamensis 10.23.1<br />
— labeosus 10.23.2<br />
— mengdingensis 10.23.3<br />
— morehensis 10.23.4<br />
— peguensis 10.23.5<br />
nepalensis, Lepidocephalichthys 4.10.9<br />
nicholsi, Lepturichthys 7.10.2<br />
nicholsi, Nemacheilus 10.34.118<br />
nicholsi, Schistura 10.34.118<br />
nielseni, Paraschistura 10.28.9<br />
nielseni, Schistura 10.28.9<br />
nieuwenhuisii, Neogastromyzon 8.10, 8.10.5<br />
niger, Yunnanilus 10.46.16<br />
nigrolineata, Botia 2.1<br />
nigra, Homaloptera 7.2.10<br />
nigrocorpa, Balitora 7.1.5<br />
nigrolineata, Ambastaia 2.1.1<br />
nigrolineata, Botia 2.1.1<br />
nigromaculatus, Eonemachilus 10.8.2<br />
nigromaculatus, Nemacheilus 10.8, 10.8.2<br />
nikkonis, Elxis 10.17, 10.17.4<br />
nikkonis, Lefua 10.17.4<br />
nikolskyi, Misgurnus 4.13.6<br />
nilgiriensis, Noemacheilus 10.34.119<br />
nilgiriensis, Schistura 10.34.119<br />
ninglangensis, Triplophysa 10.42.73<br />
niulanensis, Beaufortia 8.2.11<br />
niulanensis, Yunnanilus 10.46.17<br />
niulanjiangensis, Claea 10.5.2<br />
niulanjiangensis, Jinshaia 7.9.2<br />
niulanjiangensis, Paraprotomyzon 8.11.4<br />
niulanjiangensis, Schistura 10.5.2<br />
Niwaella 4.15<br />
— brevifasciata 4.7, 4.7.1<br />
— delicata 4.15.1<br />
— laterimaculata 4.15.2<br />
— longibarba 4.15.3<br />
— xinjiangensis 4.15.4<br />
Noemacheilus 10.21 (see species under<br />
Nemacheilus)<br />
nomi, Schistura 10.34.120<br />
nordkansuensis, Nemachilus 10.9.2<br />
190<br />
normani, Annamia 8.1.1<br />
normani, Parhomaloptera 8.1, 8.1.1<br />
notostigma, Nemacheilus 10.34.121<br />
notostigma, Schistura 10.34.121<br />
novemradiata, Schistura 10.34.122<br />
nuda, Barbatula 10.4.8, 10.4.15, 10.42.86<br />
nudidorsum, Schistura 10.34.123<br />
nudus, Lepidocephalichthys 4.10.10<br />
nudus, Nemacheilus 10.4.8, 10.42.6<br />
nuicocensis, Cobitis 4.12.1<br />
nujiangensa, Triplophysa 10.42.74<br />
nujiangensis, Balitora 7.5.9<br />
nujiangensis, Hemimyzon 7.5.9<br />
Nun 10.25<br />
nurga, Cobitis 10.25.28<br />
ny-tsieou, Cobitis 4.13.1<br />
O<br />
obeini, Schistura 10.34.124<br />
obesus, Nemacheilus 10.39, 10.39.1<br />
obesus, Sundoreonectes 10.39.1<br />
oblonga, Cobitis 4.16.24<br />
oblonga, Pangio 4.16.24<br />
obscura, Cobites 10.34.157, 10.34.189<br />
obscura, Cobitis 10.34<br />
obscura, Parhomaloptera 8.12, 8.12.1<br />
obscura, Triplophysa 10.42.75<br />
obscurus, Noemacheilus 10.34.18,<br />
10.34.189<br />
obtusirostra, Triplophysa 10.42.76,<br />
10.42.100<br />
obtusirostris, Liniparhomaloptera 8.9.3<br />
obtusirostris, Yunnanilus 10.46.18<br />
ocellata, Balitora 7.7.6<br />
ocellata, Cobitis 10.2.2<br />
ocellata, Homaloptera 7.6, 7.6.5, 7.7.6<br />
ocellata, Schistura 10<br />
ocellatus, Gastromyzon 8.5.22<br />
octoactinotos, Acantopsis 4.2.2<br />
octocirrhus, Cobitis 4.10.10<br />
Octonema 10.17, 10.24<br />
— pleskei 10.17, 10.17.5<br />
octozona, Serpenticobitis 9.1, 9.1.2<br />
oculata, Cobitis 4.4.1<br />
oedipus, Nemacheilus 10.34.125<br />
oedipus, Schistura 10.34.125<br />
ogilviei, Homaloptera 7.6.6<br />
ohridana, Cobitis 4.5.39<br />
oligolepis, Homatula 10.11.8<br />
oligolepis, Paracobitis 10.11.8<br />
oligolepos, Misgurnus 4.18.1<br />
olivaceus, Cantophrys 4.10.9<br />
olivaceus, Nemacheilus 10.21.19<br />
olivai, Cobitis 4.5.48<br />
olygoskolopos, Misgurnus 4.18.1<br />
omeiensis, Metahomaloptera 7.11, 7.11.3<br />
ophiolepis, Balitoropsis 7.2.3<br />
ophiolepis, Homaloptera 7.2.3<br />
oreas, Barbatula 10.4.9, 10.4.15<br />
oreas, Orthrias 10.4, 10.4.9
Oreias 10.5<br />
Oreias crassipedunculatus 10.5.1<br />
— dabryi 10.5, 10.5.1<br />
— furcatus 10.5.1<br />
— hoai 10.34.68<br />
— lineatus 10.34.105<br />
— nanpanjiangensis 10.42.71<br />
— punctatus 10.34.14<br />
— sonlaensis 10.34.1<br />
— trilineatus 10.34.2<br />
Oreoglanis delacouri 7.7.4<br />
Oreoleuciscus potanini 10.4.4<br />
Oreonectes 10.24<br />
— anophthalmus 10.24.6<br />
— elongatus 10.24.1<br />
— furcocaudalis 10.24.2<br />
— guananensis 10.24.7<br />
— keralensis 10.13, 10.13.2<br />
— luochengensis 10.24.8<br />
— macrolepis 10.24.3<br />
— microphthalmus 10.24.4<br />
— platycephalus 10.24, 10.24.9<br />
— polystigmus 10.24.10<br />
— retrodorsalis 10.24.11<br />
— sayu 10.17.6<br />
— translucens 10.24.5<br />
— yenlingi 10.24.9<br />
orientalis, Leptobotia 2.4.6<br />
orientalis, Nemachilus 10.42.77<br />
orientalis, Triplophysa 10.42.77<br />
ornaticauda, Gastromyzon 8.5.23<br />
ornatissimus, Nemacheilus 10.21.8<br />
ornatus, Nemacheilus 10.21.20<br />
orthocauda, Barbatula 10.34.126<br />
orthocauda, Schistura 10.34.126<br />
orthogoniata, Homaloptera 7.6.7<br />
Orthrias 10.4<br />
— araxensis 10.25.3<br />
— banarescui 10.25.5<br />
— dgebuadzei 10.4.5<br />
— dori 10.25.20<br />
— ercisianus 10.25.12<br />
— eregliensis 10.25.14<br />
— farsicus 10.25.33<br />
— golubtsovi 10.4.6<br />
— israeliticus 10.25.20<br />
— kosswigi 10.25.24<br />
— macedonicus 10.25.8<br />
— oreas 10.4, 10.4.9<br />
— pantheroides 10.25.20<br />
— pindus 10.25.35<br />
— potaninorum 10.4.10<br />
— samantica 10.25.36<br />
— sawadai 10.4.12<br />
— simavica 10.25.39<br />
— zetensis 10.4.16<br />
oxiana, Nemachilus 10.25<br />
oxianus, Oxynoemacheilus 10.25.30<br />
oxygnathus, Homatula 10.11.7<br />
oxygnathus, Nemachilus 10.11.7<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
Oxynoemacheilus 10.25<br />
Oxynoemacheilus anatolicus 10.25.1<br />
— angorae 10.25.2<br />
— araxensis 10.25.3<br />
— argyrogramma 10.25.4<br />
— banarescui 10.25.5<br />
— bergianus 10.25.6<br />
— brandtii 10.25.4, 10.25.7<br />
— bureschi 10.25.8<br />
— ceyhanensis 10.25.9<br />
— cinicus 10.25.10<br />
— cyri 10.25.11<br />
— ercisianus 10.25.12<br />
— erdali 10.25.13<br />
— eregliensis 10.25.14<br />
— evreni 10.25.15<br />
— frenatus 10.25.16<br />
— galilaeus 10.25.17<br />
— germencicus 10.25.18<br />
— hamwii 10.25.19<br />
— insignis 10.25.4, 10.25.20<br />
— kaynaki 10.25.21<br />
— kermanshahensis 10.25.22<br />
— kiabii 10.25.23<br />
— kosswigi 10.25.24<br />
— lenkoranensis 10.25.25<br />
— leontinae 10.25.26<br />
— mediterraneus 10.25.27<br />
— merga 10.25.28<br />
— namiri 10.25.29<br />
— oxianus 10.25.30<br />
— panthera 10.25.26, 10.25.31<br />
— paucilepis 10.25.32<br />
— persa 10.25.33<br />
— phoxinoides 10.25.34<br />
— pindus 10.25.35<br />
— samanticus 10.25.36<br />
— seyhanensis 10.25.37<br />
— seyhanicola 10.25.38<br />
— simavicus 10.25.39<br />
— theophilii 10.25.40<br />
— tigris 10.25.41<br />
— tongiorgii 10.25.42<br />
P<br />
pa-che-tsee-yu, Gobius 8.17.12<br />
pachycephalus, Yunnanilus 10.46.19<br />
pachychilus, Erromyzon 8.3.1<br />
pachychilus, Protomyzon 8.3.1<br />
pacifica, Cobitis 4.6.4<br />
pacifica, Iksookimia 4.6.4<br />
pahangensis, Acanthophthalmus 4.1.1<br />
pakistanica, Paraschistura 10.28.10<br />
pakistanicus, Noemacheilus 10.28.10<br />
pallens, Lepidocephalichthys 4.11.1<br />
pallidus, Nemacheilus 10.21.21<br />
paludani, Noemacheilus 10.28.12<br />
paludica, Acanthopsis 4.5<br />
paludica, Cobitis 4.5.40<br />
paludicola, Cobitis 4.5.40<br />
191<br />
paludosus, Yunnanilus 10.46.20<br />
pambaensis, Noemacheilus 10.34.47<br />
pambarensis, Mesonoemacheilus 10.18.4<br />
pangia, Cobitis 4.16.25<br />
pangia, Pangio 4.16.25<br />
Pangio 4.16<br />
— agma 4.16.1<br />
— alcoides 4.16.2<br />
— alternans 4.16.3<br />
— ammophila 4.16.4<br />
— anguillaris 4.16.5<br />
— apoda 4.16.6<br />
— atactos 4.16.7<br />
— bashai 4.16<br />
— bitaimac 4.16.8<br />
— cuneovirgata 4.16.9<br />
— doriae 4.16.10<br />
— elongata 4.16.11<br />
— filinaris 4.16.12<br />
— fusca 4.16.13<br />
— goaensis 4.16.14<br />
— incognito 4.16.15<br />
— kuhlii 4.16.16<br />
— lidi 4.16.17<br />
— longimanus 4.16.18<br />
— lumbriciformis 4.16.19<br />
— malayana 4.16.20<br />
— mariarum 4.16.21<br />
— muraeniformis 4.16.22<br />
— myersi 4.16.23<br />
— oblonga 4.16.24<br />
— pangia 4.16.25<br />
— piperata 4.16.26<br />
— pulla 4.16.27<br />
— robiginosa 4.16.28<br />
— semicincta 4.16.29<br />
— shelfordii 4.16.30<br />
— signicauda 4.16.31<br />
— superba 4.16.32<br />
panguri, Nemachilus 10.42.99<br />
panthera, Cobitis 10.25.31<br />
panthera, Oxynoemacheilus 10.25.26,<br />
10.25.31<br />
pantheroides, Orthrias 10.25.20<br />
papilio, Hemimyzon 7, 7.5.10<br />
papillos, Nemacheilus 10.21.22<br />
papillosa, Modigliania 10.21, 10.21.23<br />
papillosolabiata, Diplophysa 10.42.78<br />
papillosolabiata, Triplophysa 10.42.78<br />
papillosus, Nemacheilus 10.21.23<br />
pappenheimi, Nemacheilus 10.42.79<br />
pappenheimi, Triplophysa 10.42.79<br />
papulifera, Schistura 10.34.127<br />
Parabotia 2.5<br />
— banarescui 2.5.1<br />
— bimaculatus 2.5.2<br />
— brevirostris 2.5.3<br />
— curtus 2.5.4<br />
— dubius 2.5.5<br />
— fasciata 2.5
Parabotia fasciatus 2.5.6<br />
— heterocheila 2.5.7<br />
— heterocheilus 2.5.7<br />
— kiangensis 2.5.8<br />
— kiangsiensis 2.5.8<br />
— kimluani 2.5.5<br />
— lijiangensis 2.5.9<br />
— maculosus 2.5.10<br />
— mantschuricus 2.5.11<br />
— parvus 2.5.12<br />
— rubrilabris 2.5<br />
— taeniops 2.4.11<br />
— vancuongi 2.5.5<br />
Paracanthocobitis 10.2<br />
Paracobitis 10.11, 10.26<br />
— acuticephala 10.11.1<br />
— anguillioides 10.11.2<br />
— boutanensis 10.26.1<br />
— erhaiensis 10.11.4<br />
— ghazniensis 10.26.2<br />
— hagiangensis 10.34.64<br />
— iranica 10.26.3<br />
— longibarbata 10.42.59<br />
— longicauda 10.26.4<br />
— longidorsalis 10.11.6<br />
— malapterura 10.26.5<br />
— maolanensis 10.42.59<br />
— nanpanjiangensis 10.11.7<br />
— oligolepis 10.11.8<br />
— phongthoensis 10.34.133<br />
— posterodorsalus 10.42.59<br />
— rhadinaea 10.26.6<br />
— smithi 10.26.7<br />
— tigris 10.25.4, 10.25.32<br />
— vignai 10.26.8<br />
— wujiangensis 10.11.12<br />
paradoxa, Triplophysa 10.42.80<br />
paradoxus, Nemacheilus 10.42.80<br />
Paralepidocephalus 4.17<br />
— guishanensis 4.17<br />
— yui 4.17, 4.17.1<br />
Paramisgurnus 4.18<br />
— dabryanus 4.13.1, 4.18, 4.18.1<br />
Paranemachilus 10.27<br />
— genilepis 10.27, 10.27.1<br />
Paraprotomyzon 8.11<br />
— bamaensis 8.11.1<br />
— lungkowensis 8.11.2<br />
— multifasciatus 8.11, 8.11.3<br />
— niulanjiangensis 8.11.4<br />
— yunnanensis 8.2.15<br />
Paraschistura 10.28<br />
— alepidota 10.28.1<br />
— bampurenris 10.28.2, 10.34.20<br />
— chrysicristinae 10.28.3<br />
— kessleri 10.28.4<br />
— lepidocaulis 10.28.5<br />
— lindbergi 10.28.6<br />
— microlabra 10.28.7<br />
— naseeri 10.28.8<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Paraschistura nielseni 10.28.9<br />
— pakistanica 10.28.10<br />
— prashari 10.28.11<br />
— punjabensis 10.28.12<br />
— sargadensis 10.28.13<br />
— turcomana 10.28.14<br />
Parasewellia 8.16<br />
— monolobata 8.16.9<br />
— polylobata 8.16.9<br />
— tetralobata 8.16, 8.16.1<br />
parclitella, Homaloptera 7.6.8<br />
pardalis, Iskandaria 10.15.2<br />
pardalis, Nemachilus 10.15.2<br />
Parhomaloptera 8, 8.12<br />
— disparis 8.9, 8.9.1<br />
— microstoma 8.12.1<br />
— normani 8.1, 8.1.1<br />
— obscura 8.12, 8.12.1<br />
Parhomalopterini 8<br />
pariclavis, Gastromyzon 8.5.24<br />
parisiensis, Cobitis 10.4.2<br />
parva, Triplophysa 10.42.81<br />
parvus, Parabotia 2.5.12<br />
parvus, Yunnanilus 10.46.21<br />
patella, Sewellia 8.16.10<br />
paucicincta, Schistura 10.34.128<br />
paucifasciata, Schistura 10.34.129<br />
paucifasciatus, Nemachilus 10.34.129<br />
paucilepis, Barbatula 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
paucilepis, Oxynoemacheilus 10.25.32<br />
paucimaculatus, Nemacheilus 10.21.24<br />
pauciradiatus, Gastromyzon 8.10.6<br />
pauciradiatus, Neogastromyzon 8.10.6<br />
paucisquama, Crossostoma 8.4.7<br />
paucisquama, Formosania 8.4.7<br />
pavonacea, Acanthocobitis 10.2.5<br />
pavonacea, Cobitis 10.2.5<br />
pavonina, Balitora 7.6.5<br />
pechiliensis, Nemachilus 10.4.8<br />
pectoralis, Cobitis 4.13.1<br />
pedaschenkoi, Diplophysa 10.42.82<br />
pedaschenkoi, Triplophysa 10.42.82<br />
peguensis, Nemacheilus 10.23, 10.23.5<br />
peguensis, Neonoemacheilus 10.23.5<br />
pellegrini, Leptobotia 2.4.7<br />
pellegrini, Nemacheilus 10.34.176<br />
pengi, Balitora 7.5.11<br />
pengi, Hemimyzon 7.5.11<br />
pennocki, Gyrinocheilops 1.1, 1.1.2<br />
pennocki, Gyrinocheilus 1.1.2<br />
perakensis, Cobitophis 4.16.5<br />
peristictus, Pseudogastromyzon 8.15.9<br />
periyarensis, Nemacheilus 10.18.8<br />
persa, Cobitis 10.25, 10.25.33<br />
persa, Oxynoemacheilus 10.25.33<br />
personata, Schistura 10.34.130<br />
pertica, Schistura 10.34.131<br />
pervagata, Schistura 10.34.132<br />
peschevi, Cobitis 4.5.54<br />
192<br />
Petromizon variegatus 4.13.3<br />
petrubanarescui, Mesonoemacheilus 10.18.5<br />
petrubanarescui, Noemacheilus 10.18.5<br />
Petruichthys 10.29, 10.46<br />
— brevis 10.29.2<br />
— salmonides 10.29.1<br />
pfeifferae, Nemacheilus 10.21.25<br />
pfeifferi, Cobitis 10.21.25<br />
phongthoensis, Paracobitis 10.34.133<br />
phongthoensis, Schistura 10.34.133<br />
phoxinoides, Barbatula 10.25.34<br />
phoxinoides, Oxynoemacheilus 10.25.34<br />
phoxocheila, Cobitis 4.10.9<br />
phrygica, Cobitis 4.5.41<br />
phuketensis, Noemacheilus 10.2.9<br />
Physoschistura 10.30, 10.31<br />
— brunneana 10.30.1<br />
— chindwinensis 10.30.2<br />
— elongata 10.30.3<br />
— pseudobrunneana 10.30.4<br />
— raoi 10.30.5<br />
— rivulicola 10.30.6<br />
— shanensis 10.30.7<br />
— tuivaiensis 10.30.8<br />
— yunnaniloides 10.30.9<br />
pictava, Cobitis 10.4.2<br />
pictilis, Acanthocobitis 10.2.6<br />
pictorius, Aperioptus 4.2, 4.2.1<br />
pillaii, Homaloptera 7.6.9<br />
pindus, Barbatula 10.25.35<br />
pindus, Orthrias 10.25.35<br />
pindus, Oxynoemacheilus 10.25.35<br />
pingchowensis, Praeformosania 8.17,<br />
8.17.12<br />
pingchowensis, Vanmanenia 8.17.12<br />
pingi, Beaufortia 8.2.12<br />
pingi, Gastromyzon 8.2.12<br />
piperata, Pangio 4.16.26<br />
pironae, Cobitis 10.4.2<br />
plagiognathus, Nemachilus 10.42.86<br />
Platacanthus 4.10<br />
— agrensis 4.10, 4.10.16<br />
— maculatus 4.10, 4.10.16<br />
platiceps, Nemacheilus 10.21.26<br />
Platycara anisura 7.1.2<br />
— australis 7.3, 7.3.1<br />
platycephalus, Oreonectes 10.24, 10.24.9<br />
Platyptera sinensis 7<br />
Plesiomyzon 8.13<br />
— baotingensis 8.13, 8.13.1<br />
pleskei, Lefua 10.17.5<br />
pleskei, Octonema 10.17, 10.17.5<br />
pleurotaenia, Nemacheilus 10.46, 10.46.22<br />
pleurotaenia, Yunnanilus 10.46.22<br />
poculi, Noemacheilus 10.34.134<br />
poculi, Schistura 10.34.68, 10.34.134<br />
poecilopleura, Cobitis 4.5<br />
Pogononemacheilus 10.21<br />
polliciformis, Beaufortia 8.2.2<br />
polyfasciata, Triplophysa 10.42.83
polylepis, Beaufortia 8.2.13<br />
polylepis, Homaloptera 7.6.5<br />
polylepis, Protocobitis 4.19, 4.19.1<br />
polylepis, Vanmanenia 8.17.12<br />
polylobata, Parasewellia 8.16.9<br />
polynema, Cobitichthys 4.13.1<br />
polystigmus, Oreonectes 10.24.10<br />
polytaenia, Nemachilus 10.34.135<br />
polytaenia, Schistura 10.34.135<br />
pontica, Cobitis 4.5.42<br />
poonaensis, Nemachilus 10.21.1<br />
porthos, Schistura 10.34.136<br />
posterodorsalis, Leptobotia 2.4.8<br />
posterodorsalus, Paracobitis 10.42.59<br />
posteroventralis, Barbatula 10.42.22<br />
potanini, Homatula 10.11.9<br />
potanini, Nemacheilus 10.11, 10.11.9<br />
potanini, Oreoleuciscus 10.4.4<br />
potaninorum, Barbatula 10.4.10<br />
potaninorum, Orthrias 10.4.10<br />
Praeformosania 8.17<br />
— intermedia 8.17.12<br />
— pingchowensis 8.17, 8.17.12<br />
praestans, Gastromyzon 8.5.25<br />
prashadi, Nemachilus 10.34.137<br />
prashadi, Schistura 10.34.137<br />
prashari, Nemachilus 10.28.11<br />
prashari, Paraschistura 10.28.11<br />
pratti, Botia 2.4.10<br />
pridii, Schistura 10.34.138<br />
pristes, Kottelatlimia 4.9.3<br />
pristes, Lepidocephalichthys 4.9.3<br />
procera, Schistura 10.34.139<br />
productus, Nemachilus 10.14.4<br />
Progastromyzon 8.14<br />
— griswoldi 8.14, 8.14.3<br />
prolixifasciata, Schistura 10.34.140<br />
Prostheacanthus 4.2<br />
— spectabilis 4.2, 4.2.2<br />
Protenemacheilus 10.31<br />
Protocobitis 4.19<br />
— polylepis 4.19, 4.19.1<br />
— typhlops 4.19, 4.19.2<br />
Protomyzon 8.14<br />
— aphelocheilus 8.14.1<br />
— borneensis 8.14.2<br />
— griswoldi 8.14.3<br />
— pachychilus 8.3.1<br />
— sinensis 8.3, 8.3.3<br />
— whiteheadi 8.14.4<br />
Protonemacheilus 10.31<br />
— longipectoralis 10.31, 10.31.1<br />
psammismus, Cobitis 4.13.1<br />
pseudobrunneana, Physoschistura 10.30.4<br />
Pseudodon 10.26<br />
pseud<strong>of</strong>asciolata, Schistura 10.34.141<br />
Pseudogastromyzon 8, 8.15<br />
— changtingensis 8.15.1<br />
— cheni 8.15.2<br />
— fangi 8.15.3<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
Pseudogastromyzon fasciatus 8.15.4<br />
— jiulongjiangensis 8.15.4<br />
— laticeps 8.15.5<br />
— lianjiangensis 8.15.6<br />
— maculatum 8.15.8<br />
— meihuashanensis 8.15.7<br />
— myersi 8.15.8<br />
— peristictus 8.15.9<br />
— tungpeiensis 8.15.1<br />
Pseudogastromyzoni 8<br />
Pseudohomaloptera 7.2<br />
pseudoscleroptera, Triplophysa 10.42.84<br />
pseudoscleropterus, Nemachilus 10.42.84<br />
psiloetron, Gastromyzon 8.5.26<br />
Psilorhynchus aymonieri 1.1.1<br />
— fasciatus 8.15.4<br />
— sinensis 7.5.16, 7.9, 7.9.3<br />
psittacula, Schistura 10.34.142<br />
pterolineata, Sewellia 8.16.11<br />
Pteronemacheilus 10.31, 10.32<br />
— lucidorsum 10.32, 10.32.1<br />
— meridionalis 10.32.2<br />
pulchellus, Mesonoemacheilus 10.18.6<br />
pulchellus, Nemacheilus 10.18.6<br />
pulcher, Nemacheilus 10.41.1<br />
pulcher, Traccatichthys 10.41.1<br />
pulcherrimus, Yunnanilus 10.46.23<br />
pulchra, Botia 2.6.2<br />
pulchra, Sinibotia 2.6.2<br />
pulchripinnis, Botia 2.8.3<br />
puliensis, Sinogastromyzon 7.13.15<br />
pulla, Pangio 4.16.27<br />
pulsiz, Nemacheilus 10.25.12<br />
pumatensis, Schistura 10.34.143<br />
pumila, Iksookimia 4.6.5<br />
pumilicorpora, Hemimyzon 7.5.12<br />
pumilus, Cobitis 4.6.5<br />
punctata, Leptobotia 2.4.9<br />
punctata, Schistura 10.21.7<br />
punctatus, Misgurnus 4.13.1<br />
punctatus, Oreias 10.34.14<br />
puncticulata, Cobitis 4.5.43<br />
punctifasciata, Schistura 10.34.144<br />
punctilineata, Cobitis 4.5.44<br />
punctulatus, Gastromyzon 8.5.27<br />
punjabensis, Nemachilus 10.28.12<br />
punjabensis, Paraschistura 10.28.12<br />
purpurea, Leptobotia 2.4.11<br />
pustulosus, Gyrinocheilus 1.1, 1.1.3<br />
puta, Cobitis 4.5.6<br />
putaoensis, Nemacheilus 10.34.169<br />
pycnolepis, Homatula 10.11.10<br />
Q<br />
Qinghaichthys 10.33, 10.42<br />
— alticeps 10.33.1<br />
— rotundiventris 10.33.2<br />
— zaidamensis 10.33.3<br />
— zamegacephalus 10.33.4<br />
qiongzhongensis, Liniparhomaloptera 8.9.4<br />
193<br />
qiubeiensis, Triplophysa 10.42.85<br />
quaesita, Schistura 10.34.145<br />
quasimodo, Schistura 10.34.146<br />
quignardi, Barbatula 10.4.11<br />
quignardi, Noemacheilus 10.4.11<br />
R<br />
radnensis, Cobitis 4.20.2<br />
rajasthanica, Schistura 10.34.147<br />
rajasthanicus, Noemacheilus 10.34.147<br />
raoe, Nemachilus 10.30.5<br />
raoi, Physoschistura 10.30.5<br />
rara, Cobitis 4.5.45<br />
rara, Schistura 10.34.148<br />
rarus, Noemacheilus 10.34.148<br />
reevesae, Botia 2.6.3<br />
reevesae, Sinibotia 2.6.3<br />
reidi, Noemacheilus 10.34.149<br />
reidi, Schistura 10.34.149<br />
remadevii, Mesonoemacheilus 10.18.7<br />
rendahli, Noemacheilus 10.34.47<br />
reticulata, Schistura 10.34.150<br />
reticul<strong>of</strong>asciata, Schistura 10.34.151<br />
retrodorsalis, Oreonectes 10.24.11<br />
retropinnis, Nemachilus 10.42.86<br />
retropinnis, Triplophysa 10.42.86,<br />
10.42.100<br />
reuniens, Nemachilus 10.42.106<br />
reversa, Botia 2.7.6<br />
reversus, Syncrossus 2.7.6<br />
rhadinaea, Paracobitis 10.26.6<br />
rhadinaeus, Nemachilus 10.26.6<br />
rhodopensis, Cobitis 4.5.54<br />
ridens, Gastromyzon 8.5.28<br />
rikiki, Schistura 10.34.152<br />
ripleyi, Homalopterula 7.8, 7.8.5<br />
rivulicola, Nemachilus 10.30.6<br />
rivulicola, Physoschistura 10.30.6<br />
robertsi, Acanthopsoides 4.1.6<br />
robertsi, Schistura 10.34.153<br />
robiginosa, Pangio 4.16.28<br />
robiginosus, Acanthophthalmus 4.16.28<br />
robusta, Botia 2.6.4<br />
robusta, Sinibotia 2.6.4<br />
robusta, Triplophysa 10.42.87<br />
robustus, Nemachilus 10.42.87<br />
romanica, Cobitis 4.20.9<br />
romanica, Sabanejewia 4.20.9<br />
rosa, Triplophysa 10.42.88<br />
rosammai, Aborichthys 10.34.154<br />
rosammai, Schistura 10.34.154<br />
rossomeridionalis, Cobitis 4.5.57<br />
rossoperegrinatorum, Triplophysa 10.42.38<br />
rostrata, Botia 2.2.7<br />
rotundicauda, Homaloptera 10.24, 10.24.9<br />
rotundicaudata, Cobitis 4.8, 4.8.2<br />
rotundicaudata, Koreocobitis 4.8.2<br />
rotundiventris, Nemacheilus 10.33.2<br />
rotundiventris, Qinghaichthys 10.33.2<br />
rubidipinnis, Acanthocobitis 10.2.3, 10.2.7
ubidipinnis, Cobitis 10.2.7<br />
rubiginosus, Canthophrys 4.16.25<br />
rubrilabris, Leptobotia 2.4.10<br />
rubrilabris, Parabotia 2.5<br />
rubripinnis, Botia 2.8.5<br />
rubripinnis, Cobitis 4.10.16, 4.13.1<br />
ruppelli, Cobitis 10.22<br />
ruppelli, Nemachilichthys 10.22.1<br />
rugocauda, Sinogastromyzon 7.13.16<br />
rupecula, Schistura 10.34, 10.34.155<br />
rupelli, Cobitis 10.22.1<br />
rupicola, Chopraia 7.7, 7.7.3<br />
rupicola, Homalopteroides 7.7.3<br />
rupicola, Schistura 10.34.155<br />
russa, Schistura 10.34.156<br />
russulus, Gastromyzon 8.5.29<br />
ruzskyi, Nemachilus 10.42.101<br />
S<br />
Sabanejewia 4.20<br />
— aurata 4.20.1<br />
— balcanica 4.20.2<br />
— baltica 4.20.3<br />
— bulgarica 4.20.4<br />
— caspia 4.20.5<br />
— caucasica 4.20.6<br />
— doiranica 4.20.2<br />
— kubanica 4.20.7<br />
— larvata 4.20.8<br />
— romanica 4.20.9<br />
— thrakica 4.20.2<br />
— vallachica 4.20.10<br />
sabanus, Elxis 10.39.2<br />
sabanus, Sundoreonectes 10.39.2<br />
salari, Nemachilus 10.42.19<br />
salmonides, Nemachilus 10.29.1<br />
salmonides, Petruichthys 10.29.1<br />
salusur, Homaloptera 7.6.5<br />
samantica, Orthrias 10.25.36<br />
samanticus, Oxynoemacheilus 10.25.36<br />
sandakanensis, Acanthophthalmus 4.21,<br />
4.21.1<br />
sandakanensis, Theriodes 4.21.1<br />
sanhoensis, Sinogastromyzon 7.13.20<br />
santhamparaiensis, Homaloptera 7.6.10<br />
saravacensis, Nemacheilus 10.21.27<br />
sargadensis, Nemachilus 10.28, 10.28.13<br />
sargadensis, Paraschistura 10.28.13<br />
satunini, Cobitis 4.5.46<br />
savona, Cobitis 10.34.157<br />
savona, Nemachilus 10.34.47<br />
savona, Schistura 10.34.157<br />
sawadai, Barbatula 10.4.12<br />
sawadai, Orthrias 10.4.12<br />
sayu, Lefua 10.17.6<br />
sayu, Oreonectes 10.17.6<br />
scapanognatha, Triplophysa 10.42.89<br />
scaturigina, Schistura 10.34.158,<br />
10.34.165, 10.34.193<br />
Schistura 10.34<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Schistura aculeata 4.10.9<br />
— acuticephala 10.34.3<br />
— afasciata 10.34.4, 10.34.13<br />
— aizawlensis 10.34.5<br />
— alta 10.34.6<br />
— alticrista 10.34.7<br />
— altipedunculata 10.34.8<br />
— amplizona 10.34.9<br />
— anambarensis 10.34.10<br />
— antennata 10.34.11<br />
— aramis 10.34.12<br />
— arifi 10.34.4, 10.34.13, 10.34.42,<br />
10.34.97<br />
— athos 10.34.14<br />
— atra 10.34.15<br />
— aurantiaca 10.34.16<br />
— bachmaensis 10.34.17<br />
— bairdi 10.34.18<br />
— balteata 10.34.19<br />
— baluchiorum 10.34.20<br />
— bannaensis 10.34.21<br />
— beavani 10.34.22<br />
— bella 10.34.23<br />
— bhimachari 10.34.24<br />
— bolavenensis 10.34.25<br />
— breviceps 10.34.26<br />
— bucculenta 10.34.27<br />
— callichroma 10.34.28<br />
— callidora 10.34.29<br />
— carbonaria 10.34.30<br />
— carletoni 10.34.31<br />
— cataracta 10.34.32<br />
— caud<strong>of</strong>urca 10.34.33<br />
— ceyhanensis 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
— chapaensis 10.34.34<br />
— chindwinica 10.34.35<br />
— chrysicristinae 10.28.3<br />
— cincticauda 10.34.36<br />
— clatrata 10.34.37<br />
— conirostris 10.34.38<br />
— coruscans 10.34.39<br />
— crabro 10.34.40<br />
— crypt<strong>of</strong>asciata 10.34.41<br />
— curtistigma 10.34.13, 10.34.42<br />
— dabryi 10.5.1<br />
— dalatensis 10.34.43<br />
— daubentoni 10.34.44<br />
— deansmarti 10.34.45<br />
— defectiva 10.34.46<br />
— denisoni 10.34.47, 10.34.111<br />
— desmotes 10.34.48<br />
— devdevi 10.34.49<br />
— diminuta 10.34.50<br />
— disparizona 10.34.51<br />
— doonensis 10.34.52<br />
— dorsizona 10.34.53<br />
— dubia 10.34.54<br />
— ephelis 10.34.55<br />
194<br />
Schistura evreni 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
— fasciata 10.34.56<br />
— fascimaculata 10.34.57<br />
— fasciolata 10.34.58<br />
— finis 10.34.59<br />
— fusinotata 10.34.60<br />
— gangetica 10.34.61<br />
— geisleri 10.34.62<br />
— globiceps 10.34.63<br />
— grandis 2.7<br />
— hagiangensis 10.34.64<br />
— harnaiensis 10.34.65<br />
— heterognathos 10.35.2<br />
— himachalensis 10.34.66<br />
— hingi 10.34.67<br />
— hoai 10.34.68<br />
— horai 10.34.42, 10.34.69, 10.34.97,<br />
10.34.163<br />
— huapingensis 10.34.70<br />
— huongensis 10.34.71<br />
— imitator 10.34.72<br />
— implicata 10.34.73<br />
— incerta 10.34.64, 10.34.74<br />
— irregularis 10.34.75<br />
— isostigma 10.34.76<br />
— jarutanini 10.34.77<br />
— kangjupkhulensis 10.34.78, 10.34.113<br />
— kangrae 10.34.79<br />
— kaysonei 10.34.80<br />
— kengtungensis 10.34.81<br />
— kessleri 10.28.6<br />
— khamtanhi 10.34.82<br />
— khugae 10.34.83<br />
— kloetzliae 10.34.84<br />
— kodaguensis 10.34.85<br />
— kohatensis 10.34.86<br />
— kohchangensis 10.34.87<br />
— koladynensis 10.34.88<br />
— kongphengi 10.34.89<br />
— kontumensis 10.34.90<br />
— laterimaculata 10.34.118<br />
— latidens 10.34.91<br />
— latifasciata 10.34.92<br />
— lepidocaulis 10.28.5<br />
— leukensis 10.34.93<br />
— lingyunensis 10.42.56<br />
— longa 10.34.94<br />
— machensis 10.34.95<br />
— macrocephalus 10.34.96, 10.34.83<br />
— macrolepis 10.34.13, 10.34.97<br />
— macrotaenia 10.34.98<br />
— maculata 2.2.1<br />
— maculiceps 10.34.99<br />
— maepaiensis 10.34.100<br />
— magnifluvis 10.34.101<br />
— mahnerti 10.34.102<br />
— malaisei 10.34.103<br />
— manipurensis 10.34.104<br />
— melarancia 10.34.105
Schistura menanensis 10.34.106<br />
— microlabra 10.28.7<br />
— microphthalmus 10.42.67<br />
— minuta 10.34.107<br />
— mobbsi 10.34.108<br />
— moeiensis 10.34.109<br />
— montana 10, 10.34, 10.34.79, 10.34.110<br />
— mukambbikaensis 10.34.47, 10.34.111<br />
— multifasciata 10.34.112, 10.34.155<br />
— nagaensis 10.34.113<br />
— nagodiensis 10.34.114<br />
— namboensis 10.34.115<br />
— nandingensis 10.34.116<br />
— nasifilis 10.34.117<br />
— nicholsi 10.34.118<br />
— nielseni 10.28.9<br />
— nilgiriensis 10.34.119<br />
— niulanjiangensis 10.5.2<br />
— nomi 10.34.120<br />
— notostigma 10.34.121<br />
— novemradiata 10.34.122<br />
— nudidorsum 10.34.123<br />
— obeini 10.34.124<br />
— ocellata 10<br />
— oedipus 10.34.125<br />
— orthocauda 10.34.126<br />
— papulifera 10.34.127<br />
— paucicincta 10.34.128<br />
— paucifasciata 10.34.129<br />
— personata 10.34.130<br />
— pertica 10.34.131<br />
— pervagata 10.34.132<br />
— phongthoensis 10.34.133<br />
— poculi 10.34.68, 10.34.134<br />
— polytaenia 10.34.135<br />
— porthos 10.34.136<br />
— prashadi 10.34.137<br />
— pridii 10.34.138<br />
— procera 10.34.139<br />
— prolixifasciata 10.34.140<br />
— pseud<strong>of</strong>asciolata 10.34.141<br />
— psittacula 10.34.142<br />
— pumatensis 10.34.143<br />
— punctata 10.21.7<br />
— punctifasciata 10.34.144<br />
— quaesita 10.34.145<br />
— quasimodo 10.34.146<br />
— rajasthanica 10.34.147<br />
— rara 10.34.148<br />
— reidi 10.34.149<br />
— reticulata 10.34.150<br />
— reticul<strong>of</strong>asciata 10.34.151<br />
— rikiki 10.34.152<br />
— robertsi 10.34.153<br />
— rosammai 10.34.154<br />
— rupecula 10.34, 10.34.155<br />
— rupicola 10.34.155<br />
— russa 10.34.156<br />
— savona 10.34.157<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
Schistura scaturigina 10.34.158,<br />
10.34.165, 10.34.193<br />
— schultzi 10.34.159<br />
— semiarmata 10.34.160<br />
— sertata 10.34.161<br />
— sexcauda 10.34.162<br />
— seyhanicola 10.25.15, 10.25.32,<br />
10.25.38, 10.25.9<br />
— shadiwalensis 10.34.163<br />
— sharavathiensis 10.34.164<br />
— shebbearei 10.34.158, 10.34.165<br />
— shuangjiangensis 10.34.166<br />
— sigillata 10.34.167<br />
— sijuensis 10.34.168<br />
— sikmaiensis 10.34.169<br />
— similis 10.34.170<br />
— singhi 10.34.171<br />
— sokolovi 10.34.172<br />
— sombooni 10.34.173<br />
— sonlaensis 10.34.1<br />
— spekuli 10.34.174<br />
— spiesi 10.34.175<br />
— spiloptera 10.34.176<br />
— spilota 10.34.177<br />
— striata 10.34.24, 10.34.178<br />
— suber 10.34.179<br />
— subfusca 10.34.158<br />
— susannae 10.34.180<br />
— tenura 10.34.181<br />
— thanho 10.34.182<br />
— tigrina 10.34.183<br />
— tirapensis 10.34.184<br />
— tizardi 10.34.185<br />
— trilineata 10.34.2<br />
— tubulinaris 10.34.186<br />
— udomritthiruji 10.34.187<br />
— vinciguerrae 10.34.188<br />
— waltoni 10.34.189<br />
— xhatensis 10.34.190<br />
— yenjitee 10.34.157<br />
— yersini 10.34.191<br />
— yingjiangensis 10.34.192<br />
— zonata 10.34.158, 10.34.193<br />
schlegeli, Cobitis 4.5.8<br />
schultzi, Noemacheilus 10.34.159<br />
schultzi, Schistura 10.34.159<br />
scitulus, Gastromyzon 8.5.30<br />
scleroptera, Triplophysa 10.42.90<br />
scleropterus, Nemachilus 10.42.90<br />
Sectoria 10.35<br />
— atriceps 10.35.1<br />
— heterognathos 10.35.2<br />
— megastoma 10.35.2<br />
selangoricus, Nemacheilus 10.21.28<br />
sellaefer, Barbatula 10.42.91<br />
sellaefer, Triplophysa 10.42.91<br />
semiarmata, Schistura 10.34.160<br />
semiarmatus, Nemacheilus 10.34.160<br />
semicincta, Pangio 4.16.29<br />
semicinctus, Acanthophthalmus 4.16.29<br />
195<br />
Seminemacheilus 10.36<br />
— ispartensis 10.25.27, 10.36.1<br />
— lendlii 10.36.2<br />
— tongiorgii 10.25.42<br />
semizonata, Cobitis 10.2.7<br />
septa, Cobitis 4.5.6<br />
septemmaculata, Homaloptera 7.2.4<br />
serpentarius, Nemacheilus 7.6.1<br />
Serpenticobitidae 9<br />
Serpenticobitis 9, 9.1<br />
— cingulata 9.1.1<br />
— octozona 9.1, 9.1.2<br />
— zonata 9.1.3<br />
serrilineata, Vanmanenia 8.17.13<br />
sertata, Schistura 10.34.161<br />
Sewellia 8.16<br />
— albisuera 8.16.1<br />
— analis 8.16.2<br />
— brevis 8.16.8<br />
— breviventralis 8.16.3<br />
— diardi 8.16, 8.16.4<br />
— elongata 8.16.5<br />
— lineolata 8.16.6<br />
— marmorata 8.16.7<br />
— media 8.16.8<br />
— monolobata 8.16.9<br />
— patella 8.16.10<br />
— pterolineata 8.16.11<br />
— songboensis 8.16.6<br />
— speciosa 8.16.12<br />
— trakhucensis 8.16.13<br />
sewerzowi, Nemachilus 10.42.92<br />
sewerzowi, Triplophysa 10.42.92<br />
sexcauda, Nemacheilus 10.34.162<br />
sexcauda, Schistura 10.34.162<br />
sexmaculata, Balitoropsis 7.2.4<br />
sexmaculata, Homaloptera 7.2.4<br />
seyhanensis, Noemacheilus 10.25.37<br />
seyhanensis, Oxynoemacheilus 10.25.37<br />
seyhanicola, Oxynoemacheilus 10.25.38<br />
seyhanicola, Schistura 10.25.9, 10.25.15,<br />
10.25.32, 10.25.38<br />
shaanxiensis, Triplophysa 10.42.93<br />
shadiwalensis, Schistura 10.34.163<br />
shanensis, Nemachilus 10.30.7<br />
shanensis, Physoschistura 10.30.7<br />
shansi, Barbatula 10.4.8<br />
sharavathiensis, Schistura 10.34.164<br />
shebbearei, Nemachilus 10.34.165<br />
shebbearei, Schistura 10.34.158, 10.34.165<br />
shehensis, Noemacheilus 10.42.94<br />
shehensis, Triplophysa 10.42.26, 10.42.94<br />
shelfordii, Acanthophthalmus 4.16.30<br />
shelfordii, Pangio 4.16.30<br />
sheni, Hemimyzon 7.5.13<br />
shikokuensis, Cobitis 4.5.47<br />
shilinensis, Triplophysa 10.42.95<br />
shimogensis, Balitora 7.1.7<br />
shimogensis, Nemachilichthys 10.22.1<br />
shiyangensis, Nemachilus 10.42.96
shiyangensis, Triplophysa 10.42.96<br />
shuangjiangensis, Noemacheilus 10.34.166<br />
shuangjiangensis, Schistura 10.34.166<br />
sibirica, Cobitis 4.5.34, 4.5.48<br />
sibiricus, Nemacheilus 10.4.14, 10.4.15<br />
sichangensis, Sinogastromyzon 7.13.17<br />
sichuanensis, Yunnanilus 10.46.24<br />
sidthimunki, Ambastaia 2.1.2<br />
sidthimunki, Botia 2.1.2<br />
sigillata, Schistura 10.34.167<br />
signicauda, Pangio 4.16.31<br />
sijuensis, Noemacheilus 10.34.168<br />
sijuensis, Schistura 10.34.168<br />
sikkimensis, Noemacheilus 10<br />
sikmaiensis, Nemachilus 10.34.169<br />
sikmaiensis, Schistura 10.34.169<br />
silasi, Homaloptera 7.6.11<br />
siluroides, Nemachilus 10.42.97<br />
siluroides, Triplophysa 10.42.97<br />
simavica, Orthrias 10.25.39<br />
simavicus, Oxynoemacheilus 10.25.39<br />
similis, Schistura 10.34.170<br />
simplicispina, Cobitis 4.5, 4.5.13, 4.5.41,<br />
4.5.49<br />
sinensis, Cobitis 4.5.50<br />
sinensis, Erromyzon 8.3.3<br />
sinensis, Jinshaia 7.5.16, 7.9.3<br />
sinensis, Platyptera 7<br />
sinensis, Protomyzon 8.3, 8.3.3<br />
sinensis, Psilorhynchus 7.5.16, 7.9, 7.9.3<br />
singhi, Noemacheilus 10.34.171<br />
singhi, Schistura 10.34.171<br />
Sinibotia 2.6<br />
Sinibotia longiventralis 2.6.1<br />
— pulchra 2.6.2<br />
— reevesae 2.6.3<br />
— robusta 2.6.4<br />
— superciliaris 2.6.5<br />
— zebra 2.6.6<br />
Sinogastromyzon 7, 7.13<br />
— chapaensis 7.13.1<br />
— daon 7.13.2<br />
— dezeensis 7.13.3<br />
— hagiangensis 7.13.4<br />
— hexaocellum 7.13.1<br />
— hsiashiensis 7.13.5<br />
— hypercorpus 7.13.6<br />
— intermedius 7.13.20<br />
— lixianjiangensis 7.13.7<br />
— macrostoma 7.13.8<br />
— maon 7.13.9<br />
— minutum 7.13.10<br />
— minutus 7.13.10<br />
— multiocellum 7.13.11<br />
— namnaensis 7.13.12<br />
— nanpanjiangensis 7.13.13<br />
— nantaiensis 7.13.14<br />
— puliensis 7.13.15<br />
— rugocauda 7.13.16<br />
— sanhoensis 7.13.20<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
Sinogastromyzon sichangensis 7.13.17<br />
— szechuanensis 7.13.18<br />
— tonkinensis 7.13.19<br />
— wui 7.13, 7.13.20<br />
Sinohomaloptera 7, 7.1<br />
— acuticauda 7.5.16<br />
— h<strong>of</strong>fmanni 7.1.5<br />
— lancangjiangensis 7.1.6<br />
— longibarbatus 7.1.9<br />
— yaotanensis 7.5.16<br />
Sinohomalopterini 7<br />
sinuatus, Nemacheilus 10.2.4<br />
smithi, Homaloptera 7.7.4<br />
smithi, Homalopteroides 7.7.4<br />
smithi, Noemacheilus 10.26.7<br />
smithi, Paracobitis 10.26.7<br />
sokolovi, Schistura 10.34.172<br />
sombooni, Schistura 10.34.173<br />
Somileptes 4.4<br />
— bispinosa 4.4, 4.4.1<br />
Somileptus 4.4<br />
songamensis, Hemimyzon 7.1.5<br />
songboensis, Sewellia 8.16.6<br />
sonlaensis, Oreias 10.34.1<br />
sonlaensis, Schistura 10.34.1<br />
spanisbripes, Yunnanilus 10.46.25<br />
speciosa, Sewellia 8.16.12<br />
spectabilis, Acantopsis 4.2.3<br />
spectabilis, Gastromyzon 8.5.31<br />
spectabilis, Prostheacanthus 4.2, 4.2.2<br />
spectrum, Lepidocephalus 4.11.2<br />
spekuli, Schistura 10.34.174<br />
Speonectes 10.37<br />
— tiomanensis 10.37.1<br />
Sphaerophysa 10.38<br />
— dianchiensis 10.38, 10.38.1<br />
spiesi, Schistura 10.34.175<br />
spiloptera, Cobitis 10.34.176<br />
spiloptera, Schistura 10.34.176<br />
spilota, Schistura 10.34.177<br />
spilotus, Nemacheilus 10.34.177<br />
spilura, Cobitis 4.5.55<br />
spilurus, Misgurnus 4.13.4<br />
spiniferus, Nemacheilus 10.21.29<br />
splendens, Cobitis 4.5.51<br />
splendida, Botia 2.8.7<br />
splendida, Yasuhikotakia 2.8.7<br />
squataeniatus, Cobitis 4.5.64<br />
starostini, Noemacheilus 10.43, 10.43.1<br />
starostini, Troglocobitis 10.43.1<br />
stellatus, Gastromyzon 8.5.32<br />
stenosoma, Homalosoma 8.17, 8.17.14<br />
stenosoma, Vanmanenia 8.17.14<br />
stenura, Triplophysa 10.42.98<br />
stenurus, Nemachilus 10.42.98<br />
stephanidisi, Cobitis 4.5.52<br />
stephensoni, Balitoropsis 7.2.5<br />
stephensoni, Homaloptera 7.2.5<br />
stewarti, Diplophysa 10.42.99<br />
stewarti, Triplophysa 10.42.99<br />
196<br />
stigmata, Crossostoma 8.4.8<br />
stigmata, Formosania 8.4.8<br />
stigm<strong>of</strong>asciatus, Nemacheilus 10.21.30<br />
stolickai, Cobitis 10.42.100<br />
stolickai, Triplophysa 10.42.1, 10.4.8,<br />
10.42.27, 10.42.100<br />
strauchii, Diplophysa 10.42, 10.42.101<br />
strauchii, Triplophysa 10.42.101<br />
striata, Botia 2.2.8<br />
striata, Cobitis 4.5.53<br />
striata, Schistura 10.34.24, 10.34.178<br />
striata, Vanmanenia 8.17.15<br />
striatus, Nemacheilus 10.34, 10.34.178<br />
strumicae, Cobitis 4.5.54<br />
sturanyi, Barbatula 10.4.13<br />
sturanyi, Nemachilus 10.4.13<br />
suber, Schistura 10.34.179<br />
subfusca, Schistura 10.34.158<br />
suborbitalis, Cobitis 10.21.9<br />
sumatranus, Acanthopthalmus 4.16.29<br />
Sundoreonectes 10.39<br />
— obesus 10.39.1<br />
— sabanus 10.39.2<br />
— tiomanensis 10.37, 10.37.1<br />
superba, Pangio 4.16.32<br />
superbus, Acantophthalmus 4.16.32<br />
superciliaris, Botia 2.6, 2.6.5<br />
superciliaris, Sinibotia 2.6.5<br />
susannae, Schistura 10.34.180<br />
Syncrossus 2.7<br />
— beauforti 2.7.1<br />
— berdmorei 2.7, 2.7.2<br />
— helodes 2.7.3<br />
— hymenophysa 2.7.4<br />
— lucasbahi 2.7.5<br />
— reversus 2.7.6<br />
szechuanensis, Beaufortia 8.2.14<br />
szechuanensis, Gastromyzon 8.2.14<br />
szechuanensis, Sinogastromyzon 7.13.18<br />
T<br />
taenia, Cobitis 4.5, 4.5.55, 10.4<br />
taeniata, Nemacheilus 10.41.2<br />
taeniatus, Lepidocephalus 4.10.10<br />
taeniatus, Nemacheilus 10.41<br />
taeniatus, Traccatichthys 10.41.2<br />
taenioides, Cobitis 4.5.55<br />
taeniops, Leptobotia 2.4.11<br />
taeniops, Parabotia 2.4.11<br />
taitungensis, Hemimyzon 7.5.14<br />
taiwanica, Homaloptera 7.5.4<br />
takatsuensis, Cobitis 4.5.56<br />
tambaraparniensis, Nemacheilus 10.18.8<br />
tanaitica, Cobitis 4.5.57<br />
tanggulaensis, Nemachilus 10.42.102<br />
tanggulaensis, Triplophysa 10.42.102<br />
tangtianhensis, Nemachilus 10.42.99<br />
tarimensis, Nemachilus 10.9.4<br />
Tarimichthys 10.40, 10.42<br />
— bombifrons 10.40.1
Tarimichthys edsinicus 10.40.2<br />
— incipiens 10.40.3<br />
tatereganii, Balitoropsis 7.2.6<br />
tatereganii, Homaloptera 7.2, 7.2.6<br />
Tauphysa 10.42<br />
taurica, Cobitis 4.5.58<br />
taurica, Nemachilus 10.4.2<br />
tchangi, Balitora 7.5.15<br />
tchangi, Hemimyzon 7.5.15<br />
tchangi, Leptobotia 2.4.12<br />
tebo, Nemacheilus 10.21.31<br />
tengi, Crossostoma 8.4.9<br />
tengi, Formosania 8.4.9<br />
tenuicauda, Cobitis 10.14.5<br />
tenuicauda, Indotriplophysa 10.14.5<br />
tenuis, Indotriplophysa 10.14.6<br />
tenuis, Nemacheilus 10.14, 10.14.6<br />
tenura, Schistura 10.34.181<br />
terekensis, Nemacheilus 10.25.28<br />
tesselatus, Cobitis 4.20.4<br />
tetralineata, Cobitis 4.5.59<br />
tetraloba, Homaloptera 8.17.3<br />
tetraloba, Vanmanenia 8.17.3<br />
tetralobata, Parasewellia 8.16, 8.16.1<br />
thai, Nemacheilus 10.34.118<br />
thamicola, Cryptotora 7.4.1<br />
thamicola, Homaloptera 7.4, 7.4.1<br />
thanho, Schistura 10.34.182<br />
theophilii, Oxynoemacheilus 10.25.40<br />
Theriodes 4.21<br />
— sandakanensis 4.21.1<br />
thermalis, Cobitis 4.10.16<br />
thermalis, Lepidocephalichthys 4.10.16<br />
thiemmedhi, Acantopsis 4.2.4<br />
thrakica, Sabanejewia 4.20.2<br />
thuathienensis, Annamia 8.1.1<br />
tianeensis, Triplophysa 10.42.103<br />
tibetana, Triplophysa 10.42.104<br />
tibetanus, Nemachilus 10.42.104<br />
tientainensis, Botia 2.4.13<br />
tientainensis, Leptobotia 2.4.13<br />
tigeriveinus, Yunnanilus 10.46.22<br />
tigrina, Schistura 10.34.183<br />
tigris, Barbatula 10.4.14<br />
tigris, Cobitis 10.25.41<br />
tigris, Oxynoemacheilus 10.25.41<br />
tigris, Paracobitis 10.25.4, 10.25.32<br />
tikaderi, Aborichthys 10.1.4<br />
tinkhami, Crossostoma 8.4.10<br />
tinkhami, Formosania 8.4.10<br />
tiomanensis, Speonectes 10.37.1<br />
tiomanensis, Sundoreonectes 10.37, 10.37.1<br />
tirapensis, Schistura 10.34.184<br />
tizardi, Schistura 10.34.185<br />
tomaculum, Lepidocephalichthys 4.10.17<br />
tomiana, Barbatula 10.4.4, 10.4.14, 10.4.15<br />
tomianus, Barbatula 10.4.15<br />
tomianus, Nemacheilus 10.4.14<br />
tongiorgii, Oxynoemacheilus 10.25.42<br />
tongiorgii, Seminemacheilus 10.25.42<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
toni, Barbatula 10.4.3, 10.4.4, 10.4.8,<br />
10.4.14, 10.4.15<br />
toni, Cobitis 10.4.15<br />
tonkinensis, Misgurnus 4.13.7<br />
tonkinensis, Sinogastromyzon 7.13.19<br />
tortonesei, Nemacheilus 10.25.20<br />
Traccatichthys 10.41<br />
— pulcher 10.41.1<br />
— taeniatus 10.41.2<br />
trakhucensis, Sewellia 8.16.13<br />
transiens, Nemachilus 10.42.101<br />
translineatus, Nemacheilus 10.21.28<br />
translucens, Oreonectes 10.24.5<br />
Travancoria 7.14<br />
— elongata 7.14.1<br />
— jonesi 7.14, 7.14.2<br />
trewavasae, Triplophysa 10.42.34<br />
triangularis, Mesonoemacheilus 10.18.8<br />
triangularis, Nemacheilus 10.18, 10.18.8<br />
trichonica, Cobitis 4.5.60<br />
trilineata, Schistura 10.34.2<br />
trilineatus, Oreias 10.34.2<br />
trifasciodorsalata, Vanmanenia 8.17.13<br />
triocellata, Beaufortia 8.2.16<br />
Triplophysa 10, 10.42<br />
— akhtari 10.42.1<br />
— alexandrae 10.42.2<br />
— aliensis 10.42.3<br />
— altipinnis 10.42.4<br />
— aluensis 10.42.5<br />
— angeli 10.42.6<br />
— anterodorsalis 10.42.7<br />
— aquaecaeruleae 10.42.8, 10.42.40<br />
— arnoldii 10.42.9<br />
— bashanensis 10.42.10<br />
— bellibarus 10.42.11<br />
— bleekeri 10.42.12<br />
— brachyptera 10.42.13<br />
— brahui 10.42.14<br />
— brevibarba 10.42.15<br />
— brevibarbus 10.42.15<br />
— brevicauda 10.42.16<br />
— cakaensis 10.42.17, 10.42.100<br />
— chandagaitensis 10.42.37<br />
— chondrostoma 10.42.18<br />
— coniptera 10.42.19<br />
— crassilabris 10.42.20<br />
— cuneicephala 10.42.21<br />
— dalaica 10.42.22<br />
— daqiaoensis 10.42.23<br />
— dingi 10.42.15<br />
— dorsalis 10.42.24<br />
— dorsonotata 10.42.25, 10.42.100<br />
— drassensis 10.42.26<br />
— edsinica 10.40.2<br />
— elegans 10.42.27<br />
— eugeniae 10.14.3<br />
— farwelli 10.42.28<br />
— flavicorpus 10.42.29<br />
— furva 10.42.30<br />
197<br />
Triplophysa fuxianensis 10.42.31<br />
— gejiuensis 10.42.32<br />
— gerzeensis 10.42.33<br />
— gracilis 10.42.34<br />
— grahami 10.42.35<br />
— griffithii 10.42.36<br />
— gundriseri 10.42.37<br />
— hexiensis 10.42.38, 10.42.100<br />
— heyangensis 10.42.39<br />
— hialmari 10.42.40<br />
— hsutschouensis 10.42.41<br />
— huanglongensis 10.42.42<br />
— huanjiangensis 10.42.43<br />
— huapingensis 10.42.44<br />
— hutjertjuensis 10.42.45<br />
— intermedia 10.42.46<br />
— jianchuanensis 10.42.47<br />
— jiarongensis 10.42.48<br />
— kafirnigani 10.42.49<br />
— kaznakowi 10.16.3<br />
— kokshaalensis 10.14<br />
— kullmanni 10.42.50<br />
— kungessana 10.42.51<br />
— lacusnigri 10.42.52<br />
— lacustris 10.42.53<br />
— laterimaculata 10.42.54<br />
— laticeps 10.42.55<br />
— lingyunensis 10.42.56<br />
— lixianensis 10.42.57<br />
— longchiensis 10.42.1<br />
— longianguis 10.42.58<br />
— longibarbata 10.42.59<br />
— longipectoralis 10.42.60<br />
— macrocephala 10.42.61<br />
— macromaculata 10.42.62<br />
— macrophthalma 10.42.63<br />
— malloryi 10.42<br />
— markehenensis 10.42.64<br />
— marmorata 10.42.65<br />
— microphthalma 10.42.67<br />
— microphysus 10.42.66<br />
— microps 10.42.11, 10.42.68<br />
— moquensis 10.42.69<br />
— nandanensis 10.42.70<br />
— nanpanjiangensis 10.42.71<br />
— nasobarbatula 10.42.72<br />
— ninglangensis 10.42.73<br />
— nujiangensa 10.42.74<br />
— obscura 10.42.75<br />
— obtusirostra 10.42.76, 10.42.100<br />
— orientalis 10.42.77<br />
— papillosolabiata 10.42.78<br />
— pappenheimi 10.42.79<br />
— paradoxa 10.42.80<br />
— parva 10.42.81<br />
— pedaschenkoi 10.42.82<br />
— polyfasciata 10.42.83<br />
— pseudoscleroptera 10.42.84<br />
— qiubeiensis 10.42.85<br />
— retropinnis 10.42.86, 10.42.100
Triplophysa robusta 10.42.87<br />
— rosa 10.42.88<br />
— rossoperegrinatorum 10.42.38<br />
— scapanognatha 10.42.89<br />
— scleroptera 10.42.90<br />
— sellaefer 10.42.91<br />
— sewerzowi 10.42.92<br />
— shaanxiensis 10.42.93<br />
— shehensis 10.42.26, 10.42.94<br />
— shilinensis 10.42.95<br />
— shiyangensis 10.42.96<br />
— siluroides 10.42.97<br />
— stenura 10.42.98<br />
— stewarti 10.42.99<br />
— stolickai 10.4.8, 10.42.1, 10.42.27,<br />
10.42.100<br />
— strauchii 10.42.101<br />
— tanggulaensis 10.42.102<br />
— tianeensis 10.42.103<br />
— tibetana 10.42.104<br />
— trewavasae 10.42.34<br />
— turpanensis 10.42.105<br />
— ulacholica 10.42.106<br />
— uranoscopus 10.42.100, 10.42.107<br />
— venusta 10.42.108<br />
— waisihani 10.42.109<br />
— wuweiensis 10.42.110<br />
— xiangshuingensis 10.42.111<br />
— xiangxiensis 10.42.112<br />
— xichangensis 10.42.113<br />
— xingshanensis 10.42.114<br />
— xiqiensis 10.42.115<br />
— yaopeizhii 10.42.116<br />
— yarkandensis 10.42.15<br />
— yunnanensis 10.42.117<br />
— zhaoi 10.42.118<br />
— zhenfengensis 10.42.119<br />
Triplophysini 10<br />
troglocataractus, Nemacheilus 10.21.32<br />
Troglocobitis 10.43<br />
Troglocobitis starostini 10.43.1<br />
tschaiyssuensis, Noemacheilus 10.25.4<br />
tsin-ting-yu, Gobius 7.5.16<br />
tuberigum, Nemacheilus 10.21.33<br />
Tuberoschistura 10.44<br />
— baenzigeri 10.44.1<br />
— cambodgiensis 10.44.2<br />
tubulinaris, Schistura 10.34.186<br />
tuivaiensis, Physoschistura 10.30.8<br />
tungpeiensis, Pseudogastromyzon 8.15.1<br />
tungting, Misgurnus 4.13.1<br />
turcica, Cobitis 4.5.61<br />
Turcinoemacheilus 10.45<br />
— himalaya 10.45.1<br />
— kosswigi 10.45, 10.45.2<br />
turcmenicus, Nemachilus 10.28.13<br />
turcomana, Paraschistura 10.28.14<br />
turcomanus, Nemachilus 10.28.14<br />
turdakovi, Dzihunia 10.7.3<br />
turfanensis, Nemachilus 10.42.118<br />
Kottelat: <strong>Conspectus</strong> <strong>cobitidum</strong><br />
turio, Cobitis 10.2.2<br />
turpanensis, Triplophysa 10.42.105<br />
tweediei, Homaloptera 7.7.5<br />
tweediei, Homalopteroides 7.7.5<br />
typhlops, Protocobitis 4.19, 4.19.2<br />
U<br />
udomritthiruji, Botia 2.2.9<br />
udomritthiruji, Schistura 10.34.187<br />
ulacholica, Triplophysa 10.42.106<br />
ulacholicus, Nemachilus 10.42.106<br />
ulmeri, Homaloptera 7.8.6<br />
umbrus, Gastromyzon 8.5.33<br />
unicolor, Misgurnus 4.13.1<br />
uniformis, Barbatula 10.34.74<br />
unispina, Canthophrys 10.2.2<br />
unistriatus, Acanthophthalmus 4.10.11<br />
uranoscopus, Cobitis 10.42.107<br />
uranoscopus, Triplophysa 10.42.100,<br />
10.42.107<br />
urophthalma, Acanthocobitis 10.2.8<br />
urophthalmus, Nemachilus 10.2.8<br />
Ussuria 4.13<br />
— leptocephala 4.13, 4.13.4<br />
V<br />
Vaillantella 3, 3.1<br />
— cinnamomea 3.1.1<br />
— euepiptera 3.1.2<br />
— flav<strong>of</strong>asciata 3.1.3<br />
— maassi 3.1.3<br />
Vaillantellidae 3<br />
valenciennesi, Homaloptera 7.7.6<br />
valerii, Nemacheilus 10.42.24<br />
vallachica, Cobitis 4.20.10<br />
vallachica, Sabanejewia 4.20.10<br />
vancuongi, Parabotia 2.5.5<br />
vanderbilti, Homaloptera 7.8.6<br />
vanderbilti, Homalopterula 7.8.6<br />
vanlani, Balitora 7.1.5<br />
vanlongi, Balitora 7.1.6<br />
Vanmanenia 8.17<br />
— caldwelli 8.17.5<br />
— caobangensis 8.17.6<br />
— crassicauda 8.17.7<br />
— gymnetrus 8.17.8<br />
— hainanensis 8.17.9<br />
— homalocephala 8.17.10<br />
— lineata 8.17.11<br />
— microlepis 8.17.7<br />
— mon<strong>of</strong>asciodorsalata 8.17.13<br />
— multiloba 8.17.1<br />
— nahangensis 8.17.2<br />
— pingchowensis 8.17.12<br />
— polylepis 8.17.12<br />
— serrilineata 8.17.13<br />
— stenosoma 8.17.14<br />
— striata 8.17.15<br />
— tetraloba 8.17.3<br />
— trifasciodorsalata 8.17.13<br />
198<br />
Vanmanenia ventrosquamata 8.17.4<br />
— xinyiensis 8.17.16<br />
vardarensis, Cobitis 4.5.62<br />
vardarensis, Nemachilus 10.4.2<br />
variabilis, Cobitis 10.4.2<br />
variegata, Botia 2.4<br />
variegata, Cobitis 2.4, 2.4.1<br />
variegata, Homatula 10.11.11<br />
variegatus, Nemacheilus 10.11.11<br />
variegatus, Petromizon 4.13.3<br />
ventrosquamata, Homaloptera 8.17.4<br />
ventrosquamata, Vanmanenia 8.17.4<br />
venusta, Triplophysa 10.42.108<br />
venustus, Gastromyzon 8.5.34<br />
vermicularis, Acanthophthalmus 4.16.5<br />
vettonica, Cobitis 4.5.63<br />
victoriae, Cobitis 4.5.40<br />
vignai, Paracobitis 10.26.8<br />
viriosus, Gastromyzon 8.5.35<br />
vitoriae, Cobitis 4.5.40<br />
vittata, Cobitis 10.42.65<br />
vittatus, Canthophrys 4.10.9<br />
vulgaris, Balitoropsis 7.2.7<br />
vulgaris, Homaloptera 7.2.7<br />
W<br />
waisihani, Triplophysa 10.42.109<br />
waltoni, Nemacheilus 10.34.189<br />
waltoni, Schistura 10.34.189<br />
wassinkii, Homaloptera 7.7, 7.7.6<br />
wassinkii, Homalopteroides 7.7.6<br />
weberi, Homaloptera 7.2.5<br />
weberi, Lepidocephalus 10.21<br />
whiteheadi, Homaloptera 8.14, 8.14.4<br />
whiteheadi, Protomyzon 8.14.4<br />
wui, Botia 2.5.6<br />
wui, Sinogastromyzon 7.13, 7.13.20<br />
wujiangensis, Homatula 10.11.12<br />
wujiangensis, Paracobitis 10.11.12<br />
wuweiensis, Nemachilus 10.42.110<br />
wuweiensis, Triplophysa 10.42.110<br />
X<br />
xanthi, Nemachilus 2.5.6<br />
xhatensis, Schistura 10.34.190<br />
xiangshuingensis, Triplophysa 10.42.111<br />
xiangxiensis, Noemacheilus 10.42.112<br />
xiangxiensis, Triplophysa 10.42.112<br />
xichangensis, Triplophysa 10.42.113<br />
xingshanensis, Nemachilus 10.42.114<br />
xingshanensis, Triplophysa 10.42.114<br />
xinjiangensis, Niwaella 4.15.4<br />
xinyiensis, Vanmanenia 8.17.16<br />
xiqiensis, Triplophysa 10.42.115<br />
Y<br />
yangi, Erromyzon 8.3.4<br />
yangzonghaiensis, Eonemachilus 10.8.3<br />
yangzonghaiensis, Yunnanilus 10.8.3<br />
yaopeizhii, Triplophysa 10.42.116
yaoshanensis, Formosania 8.17.11<br />
yaotanensis, Hemimyzon 7.5.16<br />
yaotanensis, Sinohomaloptera 7.5.16<br />
yarkandensis, Hedinichthys 10.9.2, 10.9.4<br />
yarkandensis, Nemacheilus 10.9, 10.9.4<br />
yarkandensis, Triplophysa 10.42.15<br />
yasinensis, Indotriplophysa 10.14.7<br />
yasinensis, Nemachilus 10.14.7<br />
Yasuhikotakia 2.8<br />
— caudipunctata 2.8.1<br />
— eos 2.8.2<br />
— lecontei 2.8.3<br />
— longidorsalis 2.8.4<br />
— modesta 2.8.5<br />
— morleti 2.8.6<br />
— splendida 2.8.7<br />
yeni, Cobitis 4.5<br />
yenjitee, Schistura 10.34.157<br />
yenlingi, Oreonectes 10.24.9<br />
yersini, Schistura 10.34.191<br />
yingjiangensis, Nemachilus 10.34.192<br />
yingjiangensis, Schistura 10.34.192<br />
ylengensis, Cobitis 4.5.64<br />
yongdokensis, Iksookimia 4.6.6<br />
yui, Paralepidocephalus 4.17, 4.17.1<br />
yunnan, Misgurnus 4.13.1<br />
yunnanensis, Balitoropsis 7.2.8<br />
yunnanensis, Beaufortia 8.2.15<br />
yunnanensis, Botia 2.7.5<br />
yunnanensis, Paraprotomyzon 8.2.15<br />
yunnanensis, Triplophysa 10.42.117<br />
Yunnanilini 10<br />
THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />
yunnaniloides, Physoschistura 10.30.9<br />
Yunnanilus 10, 10.46<br />
Yunnanilus altus 10.46.1<br />
— analis 10.46.2<br />
— bajiangensis 10.46.3<br />
— beipanjiangensis 10.46.4<br />
— brevis 10.46<br />
— caohaiensis 10.46.5<br />
— chui 10.46.6<br />
— discoloris 10.46.7<br />
— elakatis 10.46.8<br />
— forkicaudalis 10.46.14<br />
— ganheensis 10.46.9<br />
— jinxiensis 10.46.10<br />
— longibarbatus 10.46.11<br />
— longibulla 10.46.12<br />
— longidorsalis 10.8.2<br />
— macrogaster 10.46.13<br />
— macroiscalus 10.46<br />
— macrolepis 10.46.20<br />
— macrositanus 10.46.14<br />
— nanpanjiangensis 10.46.15<br />
— niger 10.46.16<br />
— niulanensis, 10.46.17<br />
— obtusirostris 10.46.18<br />
— pachycephalus 10.46.19<br />
— paludosus 10.46.20<br />
— parvus 10.46.21<br />
— pleurotaenia 10.46.22<br />
— pulcherrimus 10.46.23<br />
— sichuanensis 10.46.24<br />
— spanisbripes 10.46.25<br />
— tigeriveinus 10.46.22<br />
— yangzonghaiensis 10.8.4<br />
— zebrinus 10.46.23<br />
yuwonoi, Balitoropsis 7.2.9<br />
yuwonoi, Homaloptera 7.2.9<br />
199<br />
Z<br />
zaidamensis, Nemachilus 10.33.3<br />
zaidamensis, Qinghaichthys 10.33.3<br />
zaisanicus, Nemachilus 10.42.101<br />
zamegacephalus, Nemachilus 10.33.4<br />
zamegacephalus, Qinghaichthys 10.33.4<br />
zanandreai, Cobitis 4.5.65<br />
zebra, Botia 2.6.6<br />
zebra, Canthophrys 2.2, 2.2.3<br />
zebra, Sinibotia 2.6.6<br />
zebrinus, Gastromyzon 8.5.36<br />
zebrinus, Yunnanilus 10.46.23<br />
zebroida, Beaufortia 8.2.16<br />
zebroidus, Gastromyzon 8.2.16<br />
zebroidus, Hemimyzon 8.15, 8.15.4<br />
zeppelini, Lepidocephalichthys 4.10.18<br />
zeravschani, Nemacheilus 10.25<br />
zetensis, Barbatula 10.4.16<br />
zetensis, Orthrias 10.4.16<br />
zhaoi, Triplophysa 10.42.118<br />
zhejiangensis, Cobitis 4.5.66<br />
zhenfengensis, Triplophysa 10.42.119<br />
zhengbaoshani, Heminoemacheilus<br />
10.10, 10.10.2<br />
zispi, Micronemacheilus 10.41.2<br />
zollingeri, Balitoropsis 7.2.10<br />
zollingeri, Homaloptera 7.2.10<br />
zonalternans, Acanthocobitis 10.2.9<br />
zonalternans, Cobitis 10.2, 10.2.9<br />
zonata, Schistura 10.34.158, 10.34.193<br />
zonata, Serpenticobitis 9.1.3